Interactions between non-native yabbies (Cherax albidus) and indigenous marron (Cherax tenuimanus) in the south-west of Western Australia are not well understood. While there is abundant evidence to suggest that invasive freshwater crayfish are detrimental to native species, the nature and degree of impact on marron populations by exotic yabbies remains unclear. Researchers have hypothesized that invasive species make faster and more appropriate use of information about their environment than native species. This greater behavioural plasticity can result in displacement of indigenous species, successful colonisation by invaders, and subsequent disturbance to natural ecosystems and representative biodiversity. / The research presented in this thesis examines the behavioural responses of an indigenous crayfish (C. tenuimanus) and an invasive crayfish (C. albidus) to waterborne odours derived from food, alarm sources and finfish predators. This study was undertaken to assist in the understanding of predatory and competitive interactions between indigenous and non-indigenous crayfish and fish predators, with particular relevance to Western Australia. Predation and competition are major forces influencing community structure in ecosystems; therefore knowledge of competitive and predatory interactions will be of benefit when considering future translocation policies. / Behavioural trials were conducted in two culture systems (54 L aquaria and a 70,000 L mesocosm), where marron and yabbies were exposed to a range of water-borne odours from finfish predators (silver perch and Murray cod), with and without competition from conspecific and heterospecific crayfish. A number of variables likely to influence crayfish behaviour were investigated: strength of chemical odour; crayfish size, gender, diurnal and nocturnal activity patterns; predator size; prior-residence; suitable habitat/shelter; and feed availability. / A key innovation in this research was the high replication in the aquarium-based observation trials using a Latin Cube design, which resulted in greater statistical strength and lower variability. More importantly, this research deviated from the tradition of exclusively using the ‘individual crayfish’ approach for odour-detection experiments and tested these results in a 70,000 L communal observation tank. This was an important development in crayfish behavioural experimentation, particularly as several key findings from the individual crayfish approach were confirmed in a multi-species environment. / Results from this study supported the hypothesis that invasive crayfish species make more appropriate use of a wider range of information about their environment than native crayfish species. Yabbies were found to possess behavioural characteristics not present in marron, such as clearer behavioural modifications to food and heterospecific odour, and cautionary behaviour in the presence of odour from a finfish predator. During simulated daylight conditions, marron displayed behaviours conducive to predation that were not present in yabbies, including less time spent in shelter and more time spent in locomotory activity. However, during specialised night-time observational studies developed during this research, these differences were not evident. This would not seem to be an unusual result, given that crayfish naturally forage at night and become more active; however, it may have important implications for future behavioural studies of crayfish, indicating a bias associated with day-time approaches. Crayfish size also played a role in behavioural modifications to water-borne odours. Larger marron displayed clearer changes in behaviour and were more responsive to heterospecific alarm odour than juveniles. Furthermore, juveniles of both species were more active than adults and sub-adults. / The expansion of the yabby population into Western Australian habitats occupied by marron has been facilitated through translocation for aquaculture, and biological characteristics of the species, some of which are typical of other invasive crayfish species including: tolerance of a variety of conditions; rapid growth; early sexual maturity; burrowing to escape drought and predation; capable of multiple spawns in a growth season; and aggressiveness. Another characteristic of invasive crayfish species also shared by yabbies, as supported by the results of this study, is high behavioural plasticity. / Although marron do not share the same level of behavioural plasticity found in yabbies, their larger body size increases their success in competitive interactions. The comparatively smaller body size of yabbies may be the major factor limiting their population expansion in the presence of marron, especially in water-bodies where shelter is a limited resource. / Marron are an important endemic species in Western Australia, but their conservation is threatened by competition and predation from exotic species. The research presented in this thesis indicates that invasive yabbies are more receptive to chemical stimuli and better equipped to respond to predation risk than marron. This information will be of benefit when considering future translocation policy in Western Australia and highlights the need for a cautious approach to species introductions.

The condition at settlement of the western rock lobster pueruli presumably reflects both their energetic condition in reaching the near shore and subsequent recruitment into adult populations. In recruiting to the near shore the pueruli swim across the continental shelf where oceanographic conditions are complex and likely influence the success of recruitment. The results from the biochemical studies are interpreted in light of the oceanography off the coast of Western Australia. I investigated the nutritional condition of larval phyllosomata, post-larval puerulus and first instar post-pueruli juveniles of the spiny lobster, Panulirus cygnus, to determine energy use during the non-feeding transitional puerulus stage. Biochemical analyses of lipid, fatty acid (FA) and protein revealed that lipid, in particular phospholipids, is primarily used for energy during the nonfeeding puerulus stage. Monounsaturated FA showed the greatest decline with development, whereas the polyunsaturated FA showed a high degree of sparing, suggesting these FA are not used as a substrate for energy production. The knowledge gained on the biochemistry of energy use in P. cygnus was then used to investigate the spatial and temporal variability in the nutritional condition, in particular lipid condition, of puerulus collected at three near shore locations (Alkimos, Jurien Bay and Dongara) along the Western Australia coast, and one offshore location (Houtman Abrolhos Islands). The one offshore location was chosen as I hypothesised that arriving pueruli are likely to be in a better state of nutrition than those arriving at more coastal locations where the potential journey from offshore larval feeding grounds to the near shore is considerably greater. This element of my research showed lipid levels to be inversely related, generally, to shelf width but were variable, suggesting pueruli may travel complex trajectories to reach nearshore settlement. The lipid and FA composition of pueruli was also consistent with spatial and seasonal variation in Leeuwin Current and coastal productivity regimes.

Mining induced seismic activity and rockbursting are critical concerns for many underground operations. Seismic activity may arise from the crushing of highly stressed volumes of rock around mine openings or from shear motion on planes of weakness. Shear-slip on major planes of weakness such as faults, shear zones and weak contacts has long been recognized as a dominant mode of failure in underground mines. In certain circumstances, it can generate large seismic events and induce substantial damage to mine openings. The Big Bell Gold mine began experiencing major seismic activity and resultant damage in 1999. Several seismic events were recorded around the second graphitic shear between April 2000 and February 2002. It is likely that the seismic activity occurred as a result of the low strength of the shear structure combined with the high level of mining induced stresses. The stability of the second graphitic shear was examined in order to gain a better understanding of the causes and mechanisms of the seismic activity recorded in the vicinity of the shear structure as mining advanced. The data were derived from the observation of the structure exposures, numerical modelling and seismic monitoring. The numerical modelling predictions and the interpreted seismic monitoring data were subsequently compared in order to identify potential relationships between the two. This thesis proposes the Incremental Work Density (IWD) as a measure to evaluate the relative likelihood of shear-slip induced seismic activity upon major planes of weakness. IWD is readily evaluated using numerical modelling and is calculated as the product of the average driving shear stress and change in inelastic shear deformation during a given mining increment or step. IWD is expected to correlate with shear-slip induced seismic activity in both space and time. In this thesis, IWD was applied to the case study of the second graphitic shear at the Big Bell mine. Exposures of the second graphitic shear yielded information about the physical characteristics of the structure and location within the mine. Numerical modelling was used to examine the influence of mining induced stresses on the overall behaviour of the shear structure. A multi-step model of the mine was created using the three- dimensional boundary element code of Map3D. The shear structure was physically incorporated into the model in order to simulate inelastic shear deformation. An elasto-plastic Mohr-Coulomb material model was used to describe the structure behaviour. The structure plane was divided into several elements in order to allow for the comparison of the numerical modelling predictions and the interpreted seismic data. Stress components, deformation components and IWD values were calculated for each element of the shear structure and each mining step. The seismic activity recorded in the vicinity of the second graphitic shear was back analysed. The seismic data were also gridded and smoothed. Gridding and smoothing of individual seismic moment and seismic energy values resulted in the definition of indicators of seismic activity for each element and mining step. The numerical model predicted inelastic shear deformation upon the second graphitic shear as mining advanced. The distribution of modelled IWD suggested that shear deformation was most likely seismic upon a zone below the stopes and most likely aseismic upon the upper zone of the shear structure. The distribution of seismic activity recorded in the vicinity of the shear structure verified the above predictions. The seismic events predominantly clustered upon the zone below the stopes. The results indicated that the seismic activity recorded in the vicinity of the second graphitic shear was most likely related to both the change in inelastic shear deformation and the level of driving shear stress during mechanical shearing. Time distribution of the seismic events also indicated that shear deformation and accompanying seismic activity were strongly influenced by mining and were time-dependant. Seismic activity in the vicinity of the second graphitic shear occurred as a result of the overall inelastic shear deformation of the shear structure under mining induced stresses. A satisfactory relationship was found between the spatial distribution of modelled IWD upon the shear structure and the spatial distribution of interpreted seismic activity (measured as either smoothed seismic moment or smoothed seismic energy). Seismic activity predominantly clustered around a zone of higher IWD upon the second graphitic shear as mining advanced. However, no significant statistical relationship was found between the modelled IWD and the interpreted seismic activity. The lack of statistical relationship between the modelled and seismic data may be attributed to several factors including the limitations of the techniques employed (e.g. Map3D modelling, seismic monitoring) and the complexity of the process involved.

This thesis presents the results of historical archaeological research at the Old Farm on Strawberry Hill in Albany, Western Australia. The site is an important colonial farm in Western Australia’s history; the location for the first farm in Western Australia (1827) and linked to many important individuals in the state’s colonial past. The site is owned and managed by the National Trust of Australia (W.A.) and is registered on both the West Australian, Heritage Council Register of Historical Places and the Australian Heritage Commission’s National Estate. Past historical and cultural biases had created an incomplete interpretation of this site that did not represent all social groups, including indentured servants, convict and Aboriginal labourers and women. The research has provided a holistic site interpretation that identified all social groups living and working on this site in the 1800s by analysing historical documents and archaeological excavated materials. The historical documentary record included both personal and official correspondence, diaries and drawings, as well as two valuable farm log books that documented the day to day events on the farm in the early to mid 1800s. The archaeological excavation was restricted to small area excavations in habitation areas still present on the site or in areas identified from 19th century surveyor maps. Both of these data sources were analysed to identify social and economic relationships, such as gender, status, class and ethnicity so that a comparison could be made between historical and archaeological data and a complementary interpretation created. The research was divided into three main periods of site occupation, firstly by convict gardeners during the government farm period from 1827 to 1832. The Spencer family period from 1833 to 1889, which is further defined by two phases, the six years from their arrival until Richard Spencer’s death in 1839 and the dispersal of the family and the property decline until it was sold in 1889. The third period of occupation by the Bird family was not discussed due to the discontinuation of a farming subsistence that distinguished it from a rural rather than an urban property. This study provides the current heritage managers with an updated interpretation of the site’s past and changing social and economic relationships on site and with the early town of Albany. It is hoped that this interpretation will be used to improve the site’s current representation and becomes the basis for a heritage conservation plan which not only recognises the importance of existing site structures, but also sub surface remains. This thesis also identifies a number of avenues for future research that will further enhance the site’s interpretation.

[Truncated abstract] The use of uncertainty analysis is gaining considerable attention in catchment hydrological modeling. In particular, the choice of an appropriate model structure, the identifiability of parameter values, and the reduction of model predictive uncertainty are deemed as essential elements of hydrological modelling. The chosen model structure must be parsimonious, and the parameters used must either be derivable from field measured data or inferred unambiguously from analysis of catchment response data. In this thesis, a long-term water balance model for the Susannah Brook catchment in Western Australia has been pursued using the ?downward approach?, which is a systematic approach to determine the model with the minimum level of complexity, with parameter values that in theory are derivable from existing physiographic data relating to the catchment. Through the analysis of the rainfall-runoff response at different timescales, and the exploration of the climate, soil and vegetation controls on the water balance response, an initial model structure was formulated, and a priori model parameter values estimated. Further investigation with the use of auxiliary data such as deuterium concentration in the stream and groundwater level data exposed inadequacies in the model structure. Two more model structures were then proposed and investigated through formulating alternative hypotheses regarding the underlying causes of observed variability, including those associated with the absence of a contribution of deep groundwater flow to the streamflow, which was indicated by deuterium concentration and internal dynamics characterized by the observed groundwater levels. ... These differences are due to differences in the time delay between rainfall and recharge between upland and riparian regions. The ages of water recharging the groundwater and discharging from the catchment were estimated by assuming a piston flow mechanism. In the deeper, upland soils, the age of recharging water was considerably larger than the unsaturated zone delay would suggest; a recharge response 16 days after an infiltration event may involve water as much as 160 days old. On the other hand, the delay and the age of recharging water were much lower in the shallow riparian zone. Where the upland zone contributes significantly to discharge, the predicted difference between the rainfall-discharge response time and the average age of discharging water can be significant.

[Truncated abstract] Mine lakes are a water body created after an open-cut mine ceases operating. The lakes develop in the former mine-pit due to the combination of groundwater inflow, surface run-off and, in some cases, due to rapid filling from river diversion. While potentially valuable water resources, these lakes often have poor water quality and managing the water body is an important part of the overall process of mine site rehabilitation. As mine lakes form in man-made pits, they have a bathymetry that is typically quite distinct from natural lakes and this can, in turn, strongly influence the hydrodynamics and hence the water quality of the water body. Despite the potential importance of these water bodies, there have been very few studies on the hydrodynamics of mine lakes. This study describes a field investigation of the hydrodynamics of a former coal mine lake, Lake Kepwari, in south-western Western Australia. In particular, this study examines the hydrodynamic processes in both the surface mixing layers and the internal mixing in the density stratified lake. Wind sheltering in the surface mixing layer occurs due to the presence of the steep walls and lake embankments. A week long field experiment was conducted in December 2003 using a combination of moored thermistor chains with meteorological stations and the deployment of rapid vertical profiling turbulent microstructure instruments and CTD drops from two boats operating on the lake. ... Simulations indicated that inclusion of a site specific sheltering effect, based on the results of the field campaign, significantly improved the models‘ performance in capturing the surface mixed layer deepening associated with episodic strong wind events that occur on the lake. Considerable internal mixing was indicated by the high dissipation rates observed, particularly near the boundaries. Large basin-wide diffusivities were also calculated from the heat budget method over long periods, showed a consistency with time, and were slightly higher in summer than during the Autumn Winter period. Although light, there are persistent winds over the lake and yet little basin-scale internal wave activity or seiching. It is hypothesized that any seiching motion was rapidly damped by strong mixing over the hydraulically rough bathymetry bathymetry created by the remnant benches from the open cut mining operation itself. This boundary mixing, in turn, drives secondary relaxation currents that transport mixed fluid from the boundaries to the interior, resulting in high effective basin-wide diffusivities. A simple boundary mixing model is proposed to describe this process.

[Truncated abstract] Wheat is the main crop in Australia and there are stringent quality requirements. Preharvest sprouting induced by rainfall between maturity and harvest lowers grain quality from premium to feed grades and reduces yield. Wheat production has expanded into the southern Western Australian region where preharvest sprouting occurs in ~1 in 4 seasons and development of more preharvest sprouting tolerant genotypes is required. The main mechanism for improving preharvest sprouting tolerance is grain dormancy. There is genetic variation for dormancy based in the embryo and seed coat but dormancy is complex and is influenced by environmental conditions during grain filling and maturation. Screening and selecting for preharvest sprouting tolerance is problematic and the level of tolerance needed for regions which differ in the level of dormancy they impose, requires clarification. The research presented here aims to answer the underlying question for breeders of how much dormancy is required for preharvest sprouting tolerance in contrasting target environments of the central and coastal wheat belt regions of Western Australia. In the central and coastal wheat belt regions, field trials with modified environments were used to determine the environmental influence on dormancy. Water supply (without directly wetting the grain) and air temperature were modified during grain development in a range of genotypes with different mechanisms of dormancy to determine the influence of environment on dormancy. ... Genotypes with embryo dormancy were consistently the most preharvest sprouting tolerant, even though this dormancy was influenced by the environmental conditions in the different seasons. Pyramiding the embryo component with the specific seed coat component and/or awnless head trait removed some of the environmental variation in preharvest sprouting tolerance, but this was generally considered excessive to the environmental requirements. The methods developed here, of field imposed stresses may provide a valuable tool to further understand the influence of environment on the regulation of dormancy, as different phenotypes can be made with the same genotype. Moisture stress, sudden changes in water supply or high temperatures during the late dough stages influenced dormancy phenotype and should be considered and avoided if possible when selecting locations and running trials for screening for genetic differences in preharvest sprouting tolerance. In the Western Australian context, the embryo component of dormancy appeared to be sufficient and should be adopted as the most important trait for breeding for preharvest sprouting tolerance.

Due to the complex nature of aquatic food webs, the interaction between abiotic and biotic factors that govern ecosystem dynamics is often elusive. Recent advancements in both the collection of reliable field data and the development of ecological models have enabled researchers to gain insights into these more complex interactions. In this study the relationship between physical and ecological processes has been explored by applying a process based coupled physical and ecological model (DYRESM-CAEDYM) to the data sets of three aquatic ecosystems. In the first, the role of zooplankton in the nutrient cycles of Lake Kinneret, Israel was quantified. The model was parameterized and calibrated using an extensive field data set. It was found that the excretion of dissolved nutrients by zooplankton accounted for up to 58% of phytoplankton demand and that this value varied seasonally in response to patterns of stratification and mixing. In the second ecosystem, Mono Lake, USA, results from model simulations were studied to determine the significance of the transport of nutrient rich hypolimnetic water via the benthic boundary layer (BBL) on lake productivity. Model results indicated that although on average the impact of BBL transport on Mono Lake ecology was not large, significant nutrient fluxes were simulated during periods when BBL transport was most active. The timing of these fluxes in the context of seasonal changes were found to be critical to specific aspects of food web dynamics. In the final application, the ecological gradients of the primary salt ponds of Shark Bay, Australia were studied with specific focus on the role of zooplankton as a determinant of ecosystem dynamics. Model results indicated that zooplankton grazing was responsible for reduced water column particulate organic matter and increased light available for the development of microbial mats. However, no direct 8 link between zooplankton grazing and observed changes in planktonic algal species composition or nutrient limitation across the salinity gradient of the ponds was found. Results from this study demonstrate the potential of a lake ecosystem model to extract useful process information to complement field data collection and address questions related to the relationship between physical and ecological processes in aquatic ecosystems.

[Tuncated abstract] Soil is a relatively low cost and robust geochemical sampling medium and is an essential part of most mineral exploration programs. In areas of covered terrain, however, soils are less reliable as a sampling medium because they do not always develop the geochemical signature of the buried mineralisation; possibly a result of limited upward transport of ore related elements into the surficial overburden. As economic demands on the resources industry grow, mineral exploration continues to expand further into areas of covered terrain where the rewards of finding a new deposit relative to the risks of finding it may be comparatively low. Thus, improving the costeffectiveness of a geochemical exploration program requires a sound understanding of the mechanisms by which soil geochemical anomalies form in transported overburden. This thesis examines the deep biotic uplift of ore related elements by deep rooting vegetation as a mechanism for the development of soil geochemical anomalies within transported overburdens, in semi-arid and arid regions. '...' Vegetation and soils were analysed at two Au prospects in Western Australia: Berkley, Coolgardie and Torquata, 210 km south-east of Kambalda, in semi-arid Western Australia to complement both the mass balance and the differential modelling. At Berkley, both the vegetation and soils located directly over the mineralisation showed high concentrations of Au. There may be indirect evidence for the operation of the deep plant uptake flux taking effect from the field evidence at Berkley. Firstly, anomalous concentrations of Au were found in the surface soils, with no detectable Au in the transported overburden. Secondly, the trace element concentrations in vegetation showed correlation to the buried lithology, which to our knowledge has not been reported elsewhere. The results from the samples at Torquata, in contrast, were less conclusive because the Au is almost exclusively associated with a surficial calcrete horizon (at <5 m soil depth). Strong correlations of Ca and Au in leaf samples however, suggest that the vegetation may be involved in the formation of calcrete and the subsequent association of Au with the calcrete. Among the vegetation components, the litter and leaf samples gave the greatest anomaly contrast at both prospects. Finally, three main drivers for the deep biotic uplift of elements were identified based on the results from the mechanistic numerical modelling exercise: i) the deep uptake flux; ii) the maximum plant concentration and; iii) the erosional flux. The relative sizes of these three factors control the rates of formation and decay, and trace element concentrations, of the soil anomaly. The main implication for the use of soils as exploration media in covered terranes is that soil geochemical anomalies may only be transient geological features, forming and dispersing as a result of the relative sizes of the accumulative and loss fluxes. The thesis culminates in the development of the first quantitative, mechanistic model of trace element accumulation in soils by deep biotic uplift.

Geologically-constrained inversion of geophysical data is a powerful method for predicting geology beneath cover. The process seeks 3D physical property models that are consistent with the geology and explain measured geophysical responses. The recovered models can guide mineral explorers to prospective host rocks, structures, alteration and mineralisation. This thesis provides a comprehensive analysis of how the University of British Columbia Geophysical Inversion Facility (UBC–GIF) gravity and magnetic inversions can be applied to subsurface mapping and exploration by demonstrating the necessary approach, data types, and typical results. The non-uniqueness of inversion demands that geological information be included. Commonly available geological data, including structural and physical property measurements, mapping, drilling, and 3D interpretations, can be translated into appropriate inversion constraints using tools developed herein. Surface information provides the greatest improvement in the reliability of recovered models; drilling information enhances resolution at depth. The process used to prepare inversions is as important as the geological constraints themselves. Use of a systematic workflow, as developed in this study, minimises any introduced ambiguity. Key steps include defining the problem, preparing the data, setting inversion parameters and developing geological constraints. Once reliable physical property models are recovered they must be interpreted in a geological context. Where alteration and mineralisation occupy significant volumes, the mineralogy associated with the physical properties can be identified; otherwise a lithological classification of the properties can be applied. This approach is used to develop predictive 3D lithological maps from geologically-constrained gravity and magnetic inversions at several scales in the Agnew-Wiluna greenstone belt in Australia’s Yilgarn Craton. These maps indicate a spatial correlation between thick mafic-ultramafic rock packages and gold deposit locations, suggesting a shared structural control. The maps also identify structural geometries and relationships consistent with the published regional tectonic framework. Geophysical inversion provides a framework into which geological and geophysical data sets can be integrated to produce a holistic prediction of the subsurface. The best possible result is one that cannot be dismissed as inconsistent with some piece of geological knowledge. Such a model can only be recovered by including all available geological knowledge using a consistent workflow process. / Science, Faculty of / Earth, Ocean and Atmospheric Sciences, Department of / Graduate