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La dispersion des poissons des récifs coralliens dépend, pour une large part, de l’existence de larves pélagiques qui peuvent être transportées au sein des courants marins loin de la population source. De l’efficacité de ce transport dépendra l’existence de populations formant un continuum, plus ou moins connectées, tout le long des zones de récifs. Toutefois, ce phénomène de dispersion est complexe et il implique aussi d’autres facteurs, physiques et biologiques, qui influent sur la dispersion de ces organismes.L’objectif de cette thèse a été d’étudier les effets des caractéristiques géographiques de l’archipel cubain et de la biologie des espèces sur la structure génétique et la connectivité des populations de poissons de récifs en utilisant comme modèle trois espèces présentant des traits d’histoire de vie différents : Stegastes partitus, Haemulon flavolineatum et Acanthurus tractus.Nous avons échantillonné des individus adultes dans cinq localités situées autour de l´archipel cubain pendant la période 2005-2010. Le polymorphisme de marqueurs microsatellites a été étudié chez S. partitus et H. flavolineatum, celui de la région non codante de l’ADNmt chez les trois espèces et celui du gène cytb chez A. tractus.L’analyse de la diversité génétique des locus microsatellites révèle des taux d’hétérozygotie observés (Ho) et attendus (He) élevés à chaque locus et pour les deux espèces, mais elles présentent différents patrons d’organisation de la diversité génétique. Chez S. partitus, la comparaison des localités prises deux à deux nous montre une différentiation exprimée par des valeurs de FST très faibles mais significatives, alors que chez H. flavolineatum aucune structuration géographique n’est observée. Ces résultats sont confirmés au niveau de l’ADN mitochondrial, lequel indique aussi une absence de différentiation d’A. tractus pour l’ensemble des localités étudiées. Les différences de structuration génétique des espèces étudiées pourraient être le résultat des effets des courants marins autour de l’ile qui agissent de façon différente sur la dispersion des espèces en fonction de leur comportement reproducteur (e.g. sites de fraie, œufs pélagiques ou benthiques et capacité des larves de rester prêt des récifs natals ou d’être transportés au loin).Pour les marqueurs mitochondriaux, nous avons obtenu des valeurs élevées de diversité haplotypique pour les trois espèces, mais la diversité nucléotidique est très différente selon les espèces, faible chez S. partitus, grande chez H. flavolineatum, et très grande chez A. tractus. La distribution du nombre de différences entre les séquences suggère que ces populations sont passées par des périodes d’expansion démographique. Ceci est confirmé par les estimations des paramètres de tests de neutralité. Les paramètres caractérisant la croissance des populations qui ont été obtenus pour les trois espèces ne sont pas semblables, ce qui suggère que le temps et l’ampleur des événements d’expansion ont été différents pour les trois espèces.L’analyse du gène cytb nous a permis d’identifier des haplotypes chez A. tractus qui n’avaient été observés préalablement que chez A. bahianus. L’aire de répartition de cette dernière espèce a été récemment restreinte à l’Atlantique Sud sur la base de la divergence génétique avec A. tractus et de la séparation géographique qui est maintenu par la barrière que constitue l’embouchure des rivières Orénoque et Amazone. La présence d’haplotypes du sud à Cuba suggère que des individus du sud ont été capables de traverser cette barrière. / Dispersal of marine fishes mainly depends on passive transport of planktonic larvae driven by ocean currents. It allows populations to be connected through thousands of kilometers but also make them sensitive to factors limiting larval dispersion. In this regard it is known that there are several physical and biological factors that determine the distribution of marine species larvae and therefore population connectivity.The present work aims to study the effects of geographic traits of Cuban archipelago and biological characteristics of reef fishes inhabiting it on the genetic structure of their populations. Three different species with distinct biological characteristics and broadly distributed within the Caribbean were selected: Stegastes partitus, Haemulon flavolineatum and Acanthurus tractus. Individuals were sampled from five localities distributed around Cuba during the time period from 2005 to 2010. We analyzed mitochondrial DNA (mtDNA) polymorphism: a fragment of the non coding region (NCR) for the three species and cytb for A. tractus. The polymorphism of nuclear microsatellite loci was studied for S. partitus and H. flavolineatum.Genetic diversity, assessed by means of Ho and He have high values when analysing microsatellite loci. These molecular markers revealed the presence of contrasting geographic structure patterns for the two species. Low but significant pairwise FST estimates were obtained for S. partitus while H. flavolineatum was genetically homogenous. These results were supported by mtDNA analyses. Likewise, A. tractus showed no evidence of significant genetic differentiation at the level of the NCR and cytb gene. Differences in population genetic structure of surveyed species could be the result of extant marine currents acting distinctly on species dispersion according to their reproductive behaviour (i.e. spawning sites, benthonic or pelagic eggs and capacity of larvae to staying close to the natal reef or being driven away from it). Mitochondrial markers showed high levels of haplotypic diversity for the three species and contrasted nucleotide diversity, low for S. partitus, intermediate for H. flavolineatum and high for A. tractus. According to these observations and results of neutrality tests and mismatch distribution analyses, it is suggested that recent population expansion occurred in these species. However differences in population parameter estimations suggest that the time and the rate of population expansion were different for the different species.The analysis of A. tractus cytb haplotype relationships grouped some Cuban individuals with two haplotypes previously described as A. bahianus. This latter species has been recently proposed as restrict to South Atlantic and separated from A. tractus by the Amazon-Orinoco outflow barrier. The presence of mtDNA haplotypes from the southern region in Cuba suggests that A. bahianus may be able to overcome this barrier.

O recrutamento é extremamente importante no ambiente recifal, sendo o principal responsável pelo reabastecimento de populações adultas de peixes. Esse fenômeno é altamente complexo, não sendo claro se é influenciado apenas por processos estocásticos ou também por processos determinísticos. No presente estudo avaliamos a dinâmica temporal do recrutamento de diversas espécies de peixes recifais, identificando sítios de berçário (i.e. recrutamento estável e alto) e a influência de variáveis ambientais. Para tal, utilizamos dados de um monitoramento de médio prazo (i.e. 2001 a 2014) realizado no Banco dos Abrolhos (BA-Brasil). Foram amostrados mais de 45 sítios, sendo levantados dados sobre a comunidade de peixes, comunidade bentônica e outras variáveis ambientais. A partir desses dados, avaliamos a variação do recrutamento por sítio em dois períodos distintos (i.e. 2001-2008/2006-2014) e a influência de variáveis ambientais no recrutamento, através da técnica Boosted Regression Trees. Constatamos que diversas espécies de peixe apresentam-se com recrutamento estável em distintos sítios de amostragem. Também observamos um efeito positivo da densidade de peixes recifais coespecíficos adultos e da cobertura relativa de algas frondosas no recrutamento de diversas espécies analisadas. No geral, observamos que há certa espécie especificidade no processo de recrutamento, porém, em escalas espaciais maiores, os padrões podem estar ligados a características mais gerais, relacionadas a um grupo taxonômico mais elevado. Em relação aos sítios de berçário, um se destacou, sendo berçário de 5 diferentes espécies, incluindo Scarus trispinosus, uma das espécies prioritárias para conservação na região de Abrolhos. Assim, recomendamos a criação de uma área marinha de proteção integral que englobe o sítio em questão. Além disso, as descobertas deste trabalho nos permitem reforçar a teoria de que o recrutamento de peixes recifais pode ser influenciado por fenômenos determinísticos e não varia simplesmente de maneira estocástica. / Recruitment is extremely important in the reef environment, because it is the main source of population replenishment. Reef fish recruitment is a highly complex process and it is not clear whether it is influenced only by stochastic processes or also by deterministic processes. Herein, we aimed to investigate temporal dynamics of reef fish recruitment, identify nursery sites (i.e. predictably high recruitment sites) and evaluate the influence of environmental variables on recruitment. We used data from a medium-term time series (i.e. 2001-2014) of scientific surveys in Abrolhos Bank (BA-Brazil). We sampled more than 45 sites, for several consecutive years and recorded data about fish community, benthic community and other environmental variables. We assessed the variation of recruitment on each site, during two distinct periods (i.e. 2001-2008 / 2006-2014), and used the Boosted Regression Trees technique to evaluate the influence of environmental variables in recruitment. We found that several reef fish species present a low variable recruitment at different sampling sites. BRT showed a positive effect of the coverage of flesh algae and abundance of conspecific in the abundance of recruits (i.e. young-of-year) of many species. Overall, we notice that the recruitment traits seems to be species specific, but we also found indications that in larger spatial scales, recruitment spatial and temporal patterns may be related to general characteristics among species of the higher taxa. Nursery sites varied among species and one site was a nursery to 5 different reef fish species, including Scarus trispinosus, a species that require priority conservation in the Abrolhos Bank. Therefore, we recommend the creation of a new no-take marine protected area that encompasses this site. Our results also indicated that reef fish recruitment may be influenced by deterministic processes and do not vary only stochastically.

Dimethylsulphoniopropionate (DMSP) and dimethylsulphoxide (DMSO) (collectively DMSP/O) are produced by marine algae, including symbiotic algae within corals. These sulphur compounds are important not only in sulphur cycle dynamics but also in potentially mediating atmospheric conditions, alleviating the effects of climate change and contributing to reef health. Most research has focused on the production of DMSP and its major degradation product, the climatically active gas, dimethylsulphide (DMS) by Acropora corals in the Great Barrier Reef. However, mechanisms for the production and release of DMSP/O by different reef taxa is poorly understood. Recently the importance of mesophotic reefs as refugia for shallow water corals has been postulated, however their role in the marine sulphur cycle is unknown. This research aimed to improve our understanding of the contemporary and climate change induced seawater and tissue production of DMSP/O in a range of reef environments and taxa. This was achieved through a combination of laboratory and field - based studies, using modern and established techniques. An effect of both elevated temperature and OA on increased tissue and seawater concentrations of DMSP/O production is reported in field and laboratory studies. Contrasting effects of benthic cover on tissue DMSP/O distributions and seawater DMSP are also noted. The importance of the physical and hydrodynamic environment on biogeochemical connectivity both within a reef and between neighbouring reefs is also focussed on. Crucially, however, the novel tissue and seawater data from mesophotic sites suggests that deeper reefs could affect the biogeochemistry of their shallow water counterparts. The key finding from this work is that climate change will result in increased seawater DMSP concentrations via two mechanisms; through the increase of cellular production of DMSP/O in all reef taxa, and by increasing the biomass of prolific DMSP producers as reefs transition to a fleshy/macroalgal assemblage. Whilst this could potentially mediate the effects of climate change, it will probably also worsen overall reef health, lead to a restructuring of reef communities from the microbial level upwards and will have possibly permanent and deleterious effects on overall ecosystem function.

珊瑚礁位處熱帶海洋的赤道附近。台灣島被北回歸線分為南與北，為理想的珊瑚礁棲息地。然而，台灣珊瑚礁的保育地位和現代威脅，特別缺乏國際出版的科學研究和數據；加上社會缺乏保護珊瑚礁和沿海資源的必要認識，讓「台灣是海鮮國家，而不是海洋國家」的錯誤認知持續阻礙了保育工作。因此，本文通過文獻回顧和訪問利益相關者，來理解包括政府機構，學者和在地社區成員間的珊瑚礁論述。結果發現，政府機構和在地社區之間的深度不信任使得海洋保護工作複雜化，加上滿足短期利益往往犧牲環境整體資源。然而，通過實施沿海綜合管理計劃(ICZMP)可能為解決方案，在所有利益攸關的共議合作中，確保保育海岸和健康珊瑚礁的諸多可持續利益。 / The majority of the coral reefs are located in the tropical oceans between the Tropics of Cancer and Capricorn, with a particularly high concentration near the equator. The Tropic of Cancer runs through the center of Taiwan, dividing the island into tropical and subtropical climate zones in the South and North, respectively, and creating an ideal habitat for the coral reef ecosystem. Yet, there is an insufficient amount of scientific research and data regarding the status of and threats to Taiwan’s coral reefs, especially in English-language publications. There furthermore exists a lack of awareness of the need to protect the reefs and coastal resources, with the widespread mentality that ‘Taiwan is a seafood country, not an ocean country’ hindering conservation efforts. Strife at the political level, in part a result of Taiwan’s unique historical events, translate into legislative and economic priority given elsewhere. This paper thus aims to realign this information asymmetry through intensive literature reviews and interviews with key stakeholders, including governmental institutions, scholars, and members of indigenous communities. Results show that deep mistrust and miscommunication between governmental institutions and the indigenous communities complicate marine conservation efforts. Satisfying short-term dominant interests frequently comes at the expense of environmental progress, a phenomenon that does not seem to substantially change with the election cycles. Nevertheless, progress has been made through such measures as the implementation of the Integrated Coastal Zone Management Plan. However, persistent issues such as improper enforcement of laws, prioritization of immediate economic gains over long-term ecological concerns, opposition from interest groups, and a lack of incentives and awareness at all levels to protect coastal regions continue to impede effective coral reef and coastal management. Overall, despite the monumental challenges set before marine conservation, the solution lies within a unified collaboration of all the stakeholders determined to protect and cultivate the many benefits of a sustainably managed coast and healthy coral reefs.

Les récifs coralliens hébergent la plus grande biodiversité sur Terre et fournissent la nourriture, le revenu économique, la protection côtière et les valeurs culturelles pour des millions de personnes. L'accessibilité des écosystèmes via les réseaux routiers est un déterminant majeur de leurs conditions, les écosystèmes les plus accessibles étant les plus menacées. Longtemps, l’estimation de l'accessibilité par l’Homme s’est limitée à l'utilisation de la distance linéaire qui ignore l’irrégularité des côtes maritimes et les réseaux routiers, facteurs influençant le temps nécessaire pour atteindre les lieux de pêche. Cette thèse présente un double enjeu : (i) développer des indices d'accessibilité intégrant l’hétérogénéité du milieu marin pour quantifier les impacts humains sur les récifs ; (ii) évaluer l'importance de ces nouveaux indices, en interaction avec les mesures de protection, pour expliquer les variations de la biomasse de poissons. Dans un premier temps, j’ai caractérisé la proximité des hommes aux récifs, en tenant compte des surfaces rencontrées (routes, chemin, eau) qui influencent le temps de trajet jusqu’aux récifs. Le temps de trajet a permis de calculer un indice de gravité, défini comme la population humaine divisée par le temps de trajet au carré, pour évaluer le niveau de pression humaine de n'importe quel récif au monde. J'ai trouvé que le temps de trajet et la gravité sont des déterminants majeurs de la biomasse de poissons à l’échelle globale. Dans un second temps, trois applications utilisant ces nouveaux indices ont permis de montrer que (i) la gravité permet de révéler l’efficacité des actions de conservation : les réserves placées là où les impacts humains sont modérés à forts fournissent des gains de conservation élevés pour la biomasse de poissons, mais elles sont peu susceptibles de protéger les prédateurs supérieurs, qui sont uniquement présent dans les réserves soumises à de faibles impacts humains ; (ii) en utilisant un cas d'étude réalisé dans le Nord-Ouest de Madagascar, j'ai illustré comment la proximité du marché peut affecter les communautés de pêcheurs et, finalement, engendrer des changements dans l'exploitation des ressources marines ; (iii) j’ai développé une nouvelle approche, le Community-Wide Scan (CWS), pour identifier les espèces de poissons qui contribuent de manière significative à la biomasse et à la couverture corallienne des récifs de l’Indopacifique, celles-ci pouvant constituer des espèces cibles à protéger. Dans le contexte de changements globaux et de perte de biodiversité, la mesure d’accessibilité constitue un outil puissant et permet de relancer le défi de gestion durable et efficace des socio-écosystèmes coralliens. / Coral reefs have the greatest biodiversity of any ecosystem on the planet and support ecosystem goods and services to million people who depend directly on them for food, economic income, coastal protection and cultural values. Ecosystem accessibility, through road networks, is the main driver of their conditions, with the most accessible ecosystems being most at risk of resource depletion. To date, measuring accessibility to humans was strictly limited to examining the linear distance which ignores ragged coastlines and road networks that can affect the time required to reach fishing grounds. This thesis presents a double challenge: (i) developing new metrics of accessibility that account for seascape heterogeneity to better assess human impacts on coral reefs; (ii) evaluating the importance of coral reef accessibility, in interactions with their management, to explain variations of fish biomass. First, I developed novel metrics of reef proximity to human populations and markets based on the friction distance which is related to transport surfaces (paved road, dirt road, water) influencing the effective reach of human settlements. Travel time was used to build the gravity index, defined as human population divided by the squared travel time, to assess the level of human pressure on any reef of the world. I found that both travel time and gravity are strong predictors of fish biomass globally. Second, three applications using these new metrics highlighted that (i) gravity identified critical ecological trade-offs in conservation since reserves placed with moderate-to-high impacts may provide substantial conservation gains for fish biomass while reserve locations with low human impacts were more likely to support higher-order predation, (ii) using a study case in Northwest Madagascar, I illustrated how market proximity can affect fishermen communities and, ultimately, trigger changes in marine resources exploitation, and (iii) I implemented a new Community-Wide Scan (CWS) approach to identify fish species that significantly contribute to the biomass and coral cover on Indo-Pacific reefs and which provided tractable conservation targets. Within the context of global changes and biodiversity loss, the thesis challenges the sustainable and efficient management of coral reef socio-ecological systems with accessibility being the cornerstone.

Coral reef ecosystems are the most diverse on earth, and their subsistence is being threatened by natural and adverse anthropogenic patterns and processes. In an effort to understand and protect these marine environments, several programs have outlined strategies and initiatives. For example, the United States Coral Reef Task Force���s Mapping and Information Working Group has outlined a specific goal to map all coral reefs below 30 m depth by 2009. This study contributes to achieving that goal for three sites around the island of Tutuila, American Samoa, lying in the heart of the South Pacific. American Samoa, a U.S. territory, is home to the Fagatele Bay National Marine Sanctuary, the smallest and most remote in the United States, and to the National Park of American Samoa. Extensive modern scientific surveys were implemented around the territory in 2001 and have since continued and increased. The presence of protected areas and the existence of scientific data collected with state of the art technology have made the site a priority for the Coral Reef Task Force. In this study, methods for classifying surficial seafloor characteristics as bathymetric position index (BPI) zones and structures were developed and applied to the study sites. BPI zones and structures were classified by using algorithms that combine high-resolution (1 m) multibeam bathymetry and its derivatives: bathymetric position index at multiple scales and slope. The development of algorithms and the classification scheme involved the use of historical and current classification studies and three-dimensional visualization. In addition, the BPI zones and structures were compared to limited biological, geological, and physical attributes recorded during accuracy assessment surveys (photos) and towed diver surveys (video). A rugosity (surface ratio) analysis was added to the study to give a picture of the seafloor roughness. The BPI zone and structure classifications overlap and extend existing classifications from Ikonos satellite imagery for water depths shallower than 30 m. Methods, data and classifications developed and applied in this study will be available to the public as a benthic habitat mapping tool (ArcGIS extension), in an online GIS data archive, and on a compact disc attached to this thesis. They contribute to a broader understanding of the marine and coastal environment and will serve as a baseline of information for benthic habitat mapping and future biological, ecological, and geological surveys. The baseline gives a good indication of characteristics that may indicate areas of high biodiversity. The final maps presented here are especially useful to managers, researchers and scientists that seek to establish and monitor a wider and more effective network of marine and coastal protection. / Graduation date: 2005

Title from PDF of title page. / Document formatted into pages; contains 112 pages. / Thesis (M.S.)--University of South Florida, 2003. / Includes bibliographical references. / Text (Electronic thesis) in PDF format. / ABSTRACT: The Florida Keys Coral Reef Monitoring Project (CRMP) began video transect sampling in 1996 and has continuously monitored 107 Florida Keys stations through 2002. The video was downward pointing and produced images from which planar projection data were calculated to determine percent cover of living benthic organisms. An absence of data assessing correlation between octocoral percent cover and octocoral abundance motivated a study to compare octocoral percent cover with abundance data acquired from the same video transects. The methods employed to extract octocoral abundance data from videotape were validated. Temporal changes in octocoral abundance, size and taxonomic group were determined by examination of video transects of 28 randomly selected stations from 1996, 1998, 1999, and 2002. Size classes were defined as [10cm, 10-40cm, ]40cm (short, medium and tall respectively). / ABSTRACT: Taxonomic groups were Gorgonia ventalina and "other octocorals" in three size classes, and Scleraxonia. An in situ study assessed the accuracy of video-derived counts. Average densities of G. ventalina and Scleraxonia were consistently about one colony/m2. Other octocoral as a group averaged 7-9 colonies/m2. When summarized by height, short and tall averaged about 1-2 colonies/m2, while colonies between 10 and 40 cm in height consistently averaged about 6 colonies/m2. Hurricane Georges, in September 1998, impacted the octocoral assemblage. Abundance declined most at stations near the storm center and stations in shallower water. Storm impact was related to octocoral height. Tall octocorals were removed more frequently than medium, short and encrusting forms. A dramatic increase of short individuals in 2002 is indicative of successful post-hurricane recruitment. By 2002, octocoral abundance had recovered to pre-hurricane levels. / ABSTRACT: This study demonstrated that abundance data can reliably be derived from archived video data, reinforcing the value of standardized video data archives. Octocoral abundance and octocoral percent cover are not strongly correlated because tall individuals disproportionately influence percent cover estimates. Nevertheless, trends in octocoral percent cover are reliable indicators of the trends in octocoral abundance. / System requirements: World Wide Web browser and PDF reader. / Mode of access: World Wide Web.

Coral communities have begun to form under small inlet-style bridges along the causeway connecting Cayo Coco to Cayo Guillermo. To understand how these bridge-reefs formed and characterize their attributes relative to surrounding communities, a combination of ecological observations and 13C isotopes were analyzed from the bridges and surrounding reefs, mangrove channels and seagrass patches. Hierarchical Cluster Analysis, Detrended Correspondence Analysis and Canonical Correspondence Analysis were carried out to group which physical attributes contributed to the coral communities in the bridge-reefs and also which habitats they were most similar to, with respect to coral and benthic composition. Bridge reef coral assemblages resembled shallow patch reefs and mangrove channels, while benthic coverage was similar to mangrove channels. Organic matter was from combined seagrass and mangrove sources, with mangrove dominating in some regions more than others. This work demonstrates that habitats within the seascape are inter-related through varying mechanisms and development is multi-dimensional.

This geostrophic current-controlled Zululand/Natal shelf displays a unique assemblage of interesting physical, sedimentological and biological phenomena. The shelf in this area is extremely narrow compared to the global average of 75km, and is characterised by submarine canyons, coral reefs, and steep gradients on the continental slope. A shelf break occurs 2.1km to 4.1km offshore and the shelf can be divided into a northern region and a southern region based on the presence or absence of a defined shelf break. The southern shelf has a poorly-defined shelf break whilst the northern shelf has a well-defined break at -65m. The poor definition of the shelf break on the southern shelf can possibly be attributed to the presence of giant, climbing sand dunes offshore of Jesser Point at depths of -37m to -60m. The northern shelf has a series of coast-parallel oriented patch coral reefs which have colonised carbonate-cemented, coastal-facies sequences. The northern shelf can be divided into three distinct zones: inner-, mid-, and outer-shelf zones. The inner-shelf is defined as the area landward of the general coral reef trend, with depths varying from 0m to -I5m and having an average gradient of 1.1. The mid-shelf is defined by the general coral reef trend, varying from -9m over the shallow central axis of the reefs to -35m along the deep reef-front environments. The outer-shelf is seaward of the coral reefs and occurs at a depth range of -35m to - 65m. Gradients vary from 1° in the south to 2.5° in the northern part of the study area, and are steep compared to world average shelf gradient of 0.116°. Four submarine canyons occur in the study area and are classified as mature- or youthful-phase canyons depending on the degree to which they breach the shelf. The origin of these canyons is not related to the position of modern river mouths but can probably be linked to palaeo-outlets of the Pongola and Mkuze River systems. It is suggested that the canyons are mass-wasting features which were exploited by palaeo-drainage during regressions. The youthful-phase canyons appear to be mass-wasting features associated with an unstable, rapidly-deposited, progradational late Pliocene sequence and a steep upper continental slope. The mature-phase canyons were probably initiated by mass-wasting but have advanced shoreward, breaching the shelf, due to their link with the palaeo-outlets of the Pongola and Mkuze Rivers during late Pleistocene regressions. Evidence of modem canyon growth has been noted on numerous SCUBA diving surveys carried out on the canyon heads. These take the form of minor wall slumps and small-scale debris flows. The canyons are also supplied with large quantities of sand in the form of large-scale shelf subaqueous dunes generated and transported by the Agulhas Current. As these bedforms meet the canyons the sediment cascades down the canyon thalweg and causes erosion and downcutting of the canyon walls and floor thereby increasing the canyon dimensions. Late Pleistocene beachrock and aeolianite outcrops with or without an Indo-Pacific coral reef veneer are the dominant consolidated lithology on the shelf. These submerged, coast-parallel, carbonate cemented, coastal facies extend semi-continuously from -5m to -95m, and delineate late Pleistocene palaeocoastline events. The rock fabric of these high primary porosity lithologies shows grains floating in a carbonate cement with occasional point-contacts. Grains are mostly quartz (80-90%), minor K-feldspar and plagioclase (5-10%), and various lithic fragments. The rocks contain conspicuous organic grains including foraminifera, bivalve, echinoid, bryozoan, red algal, and occasional sponge spicule fragments; these commonly display replacement fabrics or iron-stained rims. The dominant sedimentary structures found in these sandstone outcrops include high-angle planar cross-bedding and primary depositional dip bedding. Palaeocurrent directions sngest a palaeoenvironment dominated by a combination of longitudinal and transverse dunes with wind directions similar to those observed forming the modem dune systems. Erosional features evident on the submerged beachrocks and aeolianites include gullies trending in two different directions and sea-level planation surfaces with or without the presence of potholes. The unconsolidated sediment on the shelf is either shelf sand, composed mainly of terrigenous quartz grains; or bioclastic sediment which is partially derived from biogenic sources. The quartzose sand from the inner-shelf is generally fine-grained, moderately- to well-sorted, and coarsely- to near symmetrically-skewed. Carbonate content is low, and varies between 4-13%. Quartzose sand from the outer-shelf is fine-grained, moderately- to well-sorted, and coarsely- to very coarsely-skewed. The inner-shelf quartzose sand is better sorted than the outer-shelf sand due to increased reworking of this sediment by the high-energy swell regime. Sediment from the shallower areas of the outer-shelf (< -50m) is better sorted than sediment from depths of greater than -50m. Generally wave-reworking of quartzose shelf sand from the Sodwana Bay shelf results in greater sediment maturity than that observed from geostrophic current effects or a combination of geostrophic and wave-reworking. This sediment was derived by reworking of aeolian and beach sediments, deposited on the shelf during the period leading up to the Last Glacial Maximum (15 000 - 18 000 years B.P.) when sea-level was -130m, during the Holocene (Flandrian) transgression. Bioclastic sediment on the Sodwana Bay shelf is defined as having a CaC03 content of greater than 20% and is a mixture of biogeoically-derived debris and quartzose sand. The distribution of bioclastic sediment in the study area is widespread, with reef-derived and outer-shelf-derived populations being evident. This sediment consists of skeletal detritus originating from the mechanical and biological destruction of carbonate-secreting organisms such as molluscs, foraminifera, alcyonaria, scleractinia, cirripedia, echinodermata, bryozoa, porifera. The reef-derived bioclastic population is confined to depths less than -40m in close proximity to reef areas, whereas the shelf-derived bioclastic population occurs at depths greater than -40m and is derived from carbonate-producing organisms on deep water reefs and soft-substrate environments on the shelf. Large-scale subaqueous dunes form in the unconsolidated sediment on the outer-shelf due to the Agulhas flow acting as a sediment conveyor. These dunes are a common feature on the Sodwana Bay shelf occurring as two distinct fields at depths of -35m to -70m, the major sediment transport direction being towards the south. The two dune fields, the inner- and outer subaqueous dune fields, are physically divided by Late Pleistocene beachrock and aeolianites ledges. A bedform hierarchy has been recognised. The larger, outer dune field appears to have originated as a system of climbing bedforms with three generations of bedforms being superimposed to form a giant bedform, while the inner dune field has a less complex construction. The largest bedforms are those of the outer dune field off Jesser Point, being up to 12 m high, 4 km long and 1.2 km wide. A major slip face, with a slope of 8° is present. Bedload parting zones exist where the bedform migration direction changes from south to north. Three bedload parting zones occur in the study area at depths of -60m, -47m and -45m; two in the inner dune field and one in the outer dune field. These zones are invariably located at the southern limits of large clockwise eddy systems. Such eddies appear to be the result of topographically induced vorticity changes in the geostrophic flow and/or the response to atmospheric forcing caused by coastal low-pressure system moving up the coastline. It has been demonstrated that the inner subaqueous dune sediment conveyor is not active all the time but only during periods . of increased current strength when the Agulhas Current meanders inshore. The smaller bedforms in the outer dune field undergo continuous transport due to the current velocity on the shelf edge outer dune field being higher than the velocity experienced on the inner dune field. The very large 2·D dune which forms the outer dune field is probably not active at present: this is inferred due to the shallow angle of the mega-crest lee slope (8°). The very large Sodwana Bay subaqueous dune fields may be compared with the very large, reconstructed, subaqueous dunes which occur in Lower Permian sediments of the Vryheid Formation, northern Natal. These Permian dunes are represented, in section, as a fine- to medium-grained distal facies sandstone with giant crossbeds. These large-scale bedforms are unidirectional, but rare directionally-reversed, climbing bedforms do occur, this directional reversal may be related to bedload parting zones. On the evidence presented in this thesis, it is proposed that these Permian subaqueous dunes may be ancient analogues of the modem subaqueous dune field on the Sodwana Bay shelf. Positive-relief hummocks and negative-relief swale structures are fairly common in the fine-grained, quartzose shelf sand at depths of -30m to -60m. These appear to be transitional bedforms related to the reworking by storms of medium 2-D subaqueous dunes. These hummocky structures may be the modem equivalent of hummocky cross-stratification noted in the geological record, and if so, they are probably the first to have ever been observed underwater. The occurrences of ladderback ripples on the Sodwana Bay shelf at depths of -4m to -17m, suggest that subtidal ladderback ripples may be more common than previously thought. Ladderback ripples are common features of tidal flats and beaches where they form by late-stage emergence run-off during the ebb tide. They are generally considered diagnostic of clastic intertidal environments. The mode of formation on the Sodwana Bay shelf is different from the classic late-stage emergence run-off model of intertidal occurrences, being a subtidal setting. Subaqueous observations indicate that ladderback ripples are not environment-specific, and that additional evidence of emergence is therefore necessary to support an intertidal setting in the rock record: ladderback ripples alone are insufficient to prove an intertidal environment. The coral patch reefs of the northern Natal coast are unique, being the most southerly reefs in Africa, and totally unspoilt. The Zululand reefs are formed by a thin veneer of Indo-Pacific type corals which have colonised submerged, late Pleistocene beachrocks and aeolianites. Two-Mile Reef at Sodwana Bay has been used to develop a physiograpbic and biological zoning model for Zululand coral reefs, which has been applied to other reefs in the region. Eight distinct zones can be recognised and differentiated on the basis of physiographic and biological characteristics. The reef fauna is dominated by an abundance of alcyonarian (soft) corals, which constitute 60-70% of the total coral fauna. The Two-Mile Reef zoning model has been successfully applied to larger reefs such as Red Sands Reef, and smaller patch reefs (Four-Mile and Seven-Mile Reefs) in the same general area. In this thesis extensive use has been made of Hutton's uniformitarian principles. Hutton's doctrine is particularly relevant to the study of depositional processes and relict shorelines. Coastal processes and weather patterns during the late Pleistocene were broadly similar to modem conditions enabling direct comparisons to be made. A computer-aided facies analysis model has been developed based on textural statistics and compositional features of carbonate-cemented coastal sandstones. Many attempts have been made to distinguish different ancient sedimentary depositional environments, most workers in this field having little success. The new method of facies reconstruction is based on: (1) underwater observations of sedimentary structures and general reef morphology; (2) a petrographic study of the reef-base enabling flve facies: aeolianite, backbeach, forebeach, swash, and welded bar facies to be recognised, which control the geomorphology of Two-Mile Reef; (3) cluster and discriminant analysis comparing graphic settling statistics of acid-leached reef-base samples with those of modem unconsolidated dune/beach environments. The results of this analysis demonstrated that the beachrocks and aeolianites on the shelf formed during a regression and that late Pleistocene coastal facies are similar to modem northern Zululand coastal environments, which have been differentiated into aeolian, backbeach, forebeach, swash, & welded bar. A late Pleistocene and Holocene history of the shelf shows that during the late Pleistocene, post Eemian regressions resulted in deposition and cementation of coast-parallel beachrocks and aeolianites, which define a series of four distinct palaeocoastline episodes with possible ages between 117 000 and 22 000 years B.P. The beachrock/aeolianites formed on the shelf during stillstands and slow regressions, and the gaps between these strandline episodes represent periods of accelerated sealevel regression or a minor transgressive phase which hindered deposition and cementation. The formation of these lithologies generated a considerable sediment sink in the nearshore zone. This reduced sediment supply and grain transport in the littoral zone during the Holocene, and probably enhanced landward movement of the shoreline during the Flandrian transgression. Prior to the Last Glacial Maximum, the beachrock/aeolianite sedimentary sequence was emergent and blanketed by shifting aeolian sands. The Pongola River, which flowed into Lake Sibaya, reworked the unconsolidated sediments on the shelf, and exploited the route of least resistance: along White Sands and Wright Canyon axes. The erosion resulting from fluvial denudation in Wright Canyon has caused this canyon to erode some of the beachrock/aeolianite outcrops which form palaeocoastline episode 2 and entrench the canyon to a deeper level; this eroded the shelf to a distance of 2km offshore. During the Flandrian transgression the unconsolidated sediment cover was eroded, exposing and submerging the beachrock/aeolianite sequence. Flandrian stillstands caused erosional features such as wave-planed terraces, potholes, and gullies to be incised into beachrock and aeolianite outcrops; these are seen at present depths of -47m, -32m, .26m, -22m, -17m to -15m, and -12m. High energy sediment transfers, in an onshore direction, resulted in the deposition of sand bars across the outlet of Lake Slbaya's estuary and the development of a 130m + coastal dune barrier on a pre-existlng, remnant Plelstocene dune stub. Sea-level stabilised at its present level 7 000-6 000 years B.P. and coral reef growth on the beachrock/aeolianite outcrops probably started at 5 000 years B.P. A minimum age for the formation of the northern Zululand coral reefs has been established at 3780 ± 60 years B.P. A mid Holocene transgression relating to the Climatic Optimum deposited a + 2m raised beach rock sequence. This transgression eroded the coastal dune barrier and caused a landward shoreline translation of approximately 40m. A minor transgression such as this can be used as a model for coastal erosion which will result from the predicted 1.5m rise in sea-level over the next century. This rise in sea-level could result in a 30m landward coastline translation of the present coastline, ignoring the influence that storms and cyclones will have on the coastline configuration. / Thesis (Ph.D.)-University of Natal, Durban, 1991.