Magister Scientiae (Biodiversity and Conservation Biology) / Desertification deteriorates the landscape functionality of rangelands, affecting the resilience of biome boundaries which have a cascade effect not only in vegetation composition and characteristics but also in animal communities. The balance between plants and small mammals are essential in maintaining the functionality (i.e. nutrient cycling, soil stability, and water infiltration) of rangeland which includes the arid Steinkopf communal rangeland. However, the landscape functionality of rangelands has not been extensively studied, especially in communal areas where desertification is a serious concern since many people farm with livestock to survive. The aim of this study was to assess landscape functionality along an arid biome boundary and its relationship to small mammal assemblages. This study further aimed to provide a local landscape perspective of the current desertification process and projected expansion of the Desert Biome into more mesic biomes in South Africa. Three replicates for three Desert Biome, ecotone and Succulent Karoo Biome sites (27 sites in total) were selected to assess landscape functionality and survey small mammal assemblages over a one-year period. The change in landscape functionality between the three areas was assessed to determine the resilience of the biome boundary to desertification. For each small mammal survey, 216 live Sherman traps were set up during the summer and winter seasons to account for breeding, mortality, and possible migration. With the data, the body condition index, population density, and diversity were quantified.

Most Australian studies on estuarine plankton have examined distribution and abundance in relation to hydrological changes, primary productivity and associated nutrient dynamics. Relatively few have examined the complex interactions between zooplankton grazers and the type and quality of food available, or the role of zooplankton grazers in structuring phytoplankton communities, or their contribution to the nutrient pool. The ecological role of zooplankton grazers in the Swan River estuary, Perth, Western Australia, was examined as part of a collaborative research project directed by the Western Australian Estuarine Research Foundation, which was established in response to concern about increasing intensity and persistence of algal blooms. The present study focussed on one component of the zooplankton, the Copepoda, as model zooplankton grazers. A regular zooplankton monitoring programme, undertaken over a two year period, provided data on seasonal patterns of abundance and distribution of zooplankton over a broad spectrum of physical conditions. Relationships were identified between habitat variables, such as algal biomass, dissolved oxygen, salinity and suspended solids and zooplankton distribution, relative abundance and species composition. Prior to the inception of this study, it was assumed that copepod species composition, abundance and richness in the Swan River estuary may have changed over time, in response to long-term declines in water quality. Comparison of historical copepod monitoring data with current data did not detect any such change and it was concluded that there was greater variation in copepod species composition, abundance and richness within years than between years and that no significant change had occurred between 1966 and 1997. / However, an absence or reduction in abundance of copepods in areas of very high algal biomass (>80 pg chlorophyll a.L-1) suggests that local loss of water quality may have an impact on copepods over a small spatial scale within the estuary. Different aspects of the interactions between zooplankton grazers and phytoplankton were studied. Zooplankton grazing rates were measured in situ during algal blooms and in the laboratory under controlled conditions to determine the potential for zooplankton grazers to reduce algal biomass. Field and laboratory experiments supported the hypothesis that copepods and other zooplankton can exert 'top-down control' over phytoplankton biomass, but that the type and biomass of phytoplankton present affected their ability to exert this control. The results of the field and laboratory grazing experiments, along with literature data, were used to provide input data for a model of zooplankton and phytoplankton dynamics during a dinoflagellate bloom in the Swan River estuary. The model was tested against biomass measurements of zooplankton and phytoplankton to determine how well it predicted actual changes in the plankton community. The simulated output closely followed the measured Page x field data and fitted regression curves and provided information about diurnal patterns of phytoplankton production, respiration and migration and hydrodynamic transport, which was not available from field data. It was shown that zooplankton grazing, particularly grazing by microzooplankton, was the process contributing most to the observed decline in dinoflagellate biomass. Nutrient availability is one of several factors determining productivity of phytoplankton. Nutrients within copepod faecal pellets are relocated by faecal deposition to sediments, where microbial activity leads to the remineralisation of these nutrients. / Quantification of metabolic excretion of nutrients by copepods and the rate at which pellets are produced by copepod grazers, the concentration of nutrients within faecal pellets and the rate at which these nutrients are released indicated that copepods may play an important role in nutrient regeneration during summer and autumn when allochthonous nutrients are unavailable. At other times of the year, it is unlikely that copepods play an important role in nutrient regeneration. The research has provided a more detailed level of understanding of the interactions between zooplankton, phytoplankton and their environment. The data is ideally suited for use in a computer model to predict the effects of management actions on the Swan River estuary. This would allow pre-emptive management strategies to be developed and lessen the focus on reactive management.

Bibliography: leaves 105-121. This study reports the first thorough investigation of palatability variation in Atriplex vesicaria Heward ex Benth. (bladder saltbush). Intensive small-plot dietary trials, supported by a paddock dietary experiment, cross-fence comparisons and cafeteria trials, showed that sheep preferentially grazed female saltbushes over male and bisexual ones. Sheep avoided male saltbushes due to a chemical deterrent, and used visual (male flower spike) and olfactory cues to detect male plants. The effect of this selective grazing was to reduce the size and reproductive output of female shrubs. Sheep also tended to return to shrubs they had grazed previously. In the semi-arid regions of South Australia, where bladder saltbush is grazed year-round, physical protection is recommended to maximise survival and reproductive output of heavily grazed shrubs.

Excessive bacterial levels are a major water quality concern. Better methods are needed to quantify the proportion of bacterial loading contributed by various sources, and best management practices are needed to restore water quality. This study assessed the ability of alternative water supplies and grazing management to reduce E. coli loading from cattle and evaluated the ability of quantitative polymerase chain reaction analysis of total and bovine-associated Bacteroides markers (AllBac and BoBac, respectively) to determine the percentage of bovine-associated fecal contamination. Runoff from seven small watersheds, representing ungrazed, properly stocked, and overstocked conditions, was analyzed for E. coli, AllBac, and BoBac to assess grazing management impacts on E. coli runoff and the effectiveness of Bacteroides markers. To determine the effectiveness of alternative water, instream E. coli levels and cattle movement were evaluated before and after alternative water was provided. The study found that when alternative off-stream water was provided, the amount of time cattle spent in the creek was reduced 43 percent and the direct deposition of E. coli into Clear Fork of Plum Creek was estimated to be reduced from 1.11E 07 to 6.34E 06 colony forming units per animal unit per day. Observed pre- and post-treatment E. coli loads suggested similar reductions; however, this study could not conclusively attribute observed E. coli loading reductions to providing alternative water because of the lack of statistical significance of these observations, possibly due to decreased streamflow during Year 2 (due to drought) and a corresponding increase in E. coli levels. The study found that rotational stocking, if timed appropriately, was very effective at reducing E. coli runoff. The impact of grazing timing in relation to runoff events was more significant than the impact of grazing management (i.e. ungrazed properly stocked or overstocked) or stocking rate. When runoff occurred more than two weeks following grazing, E. coli levels in runoff were decreased more than 88 percent. Finally, data suggest that AllBac and BoBac markers are good indicators of recent fecal contamination from cattle. However, although elevated BoBac/AllBac ratios generally aligned well with cattle presence, this ratio appeared to underestimate the percentage of bovine-associated fecal contamination.

This research was conducted in the Mamora cork oak forest of Morocco to: (1) Describe the impacts of sheep grazing (none, 35% use, 70% use) in March, April, May and June of 1987 and 1988 on seasonal forage production; (2) evaluate the effects of these factors on annual changes in herbaceous foliar cover and botanical composition; and (3) determine their effects on seasonal changes in forage quality of the herbaceous vegetation. Neither the month of 1987 nor the degree of use had significant effects on the rate of herbage accumulation. In 1988, the month of grazing, the degree of use and their interaction had significant effects on all rates of herbage accumulation except that between May and June for the March grazed treatment. For the April, May and June treatments, grazing depressed the forage production of the subsequent months. The interaction of the month of grazing and the degree of use had no significant effect on the botanical composition of all groups of species. Averaged over the season, the other forbs were the dominant group of species. Total canopy cover was significantly affected by the year only. The 1988 total vegetative ground cover was significantly higher than the 1987 and 1989 ones as a result of the impacts of the intensity and distribution of rains on vegetative growth, development and distribution. The degree of use had a significant effect on the forage nutrients only for some grazing treatments. It seemed that the effect of the grazing intensity on forage nutrients of the subsequent months is more evident when grazing occurs early in the season. This study developed the first information for this area related to the response of the vegetation to the time and the intensity of grazing. The results suggest that scheduling intensity and timing of grazing livestock might be a key factor in community dynamics. Combined research on grazing management, fertilization and seeding should be undertaken and implemented in the next Mamora management plan. / Graduation date: 1992

Seasonal trends in forage production and environmental parameters for five plant community types within a northeastern Oregon riparian zone were described and modeled using correlation and path analysis. Wet meadows produced the greatest amount of herbage biomass, followed by moist bluegrass meadows, gravel bars, forests and dry bluegrass meadows. Trends in soil moisture generally increased and then declined from spring to fall. Depth to the water table declined and then increased. Soil temperatures steadily increased. Variables driving seasonal forage production varied by community type. Soil moisture was most important in dry bluegrass meadows and least important in wet meadows. Depth to the water table was most important in wet meadows and least important in dry bluegrass meadows. The amount of herbage production which had already occurred was also an important variable in describing biomass production. Streamflow levels and the amount of production having occurred were driving variables in the gravel bar communities. Preference for grazing different riparian vegetation community types and forage intake by cattle was monitored over a three-week grazing period occuring at the end of summer. Concurrent to preference and intake, vegetative and nutritional characteristics of the forage available for grazing were monitored and relationships between these variables and both community preference and intake described through correlation and path analyses. Grazing cattle initially favored communities with highly digestibile forage, hence communities dominated by Kentucky bluegrass were most preferred. Late in the grazing period community preference was best associated with community abundance, indicating that cattle were grazing communities in proportion to their abundance in the pasture. Intake levels were greater during the first year of the study than the second (2.15 versus 1.81 percent of body weight). Daily grazing time declined as livestock neared the end of the grazing period. Intake was correlated with in vitro dry matter digestibility and the amount of time spent grazing, but poorly related to the amount of forage available. The indirect effect of the amount of forage available on intake was greater than the direct effect and functioned through increases in grazing time as a result of increased availability of highly digestible forage. / Graduation date: 1992

A study investigating the effect of sheep stocking density and grazing duration on forage intake, grazing efficiency, dietary selectivity and subsequent forage accumulation in tall fescue(Festuca arundinacea)- subclover(Trifolium subtaranium) pastures was conducted near Corvallis, Oregon during early and late spring in 1988 and 1989. Grazing treatments were 2, 6, and 10-day duration and corresponding stocking densities 380, 130, 78 and 1390, 460 and 280 ewes/ha during early and late trials each year, respectively. Average daily intake and grazing efficiency were highest (P< 0.05) in the 10-day duration and lowest in the 2-day duration. During the first 2 days of all duration treatments, average daily intake decreased as stocking density increased (P< 0.05). Manure cover and crowding stress may explain lower average daily intake under the shorter duration/high density treatments. However, stocking density had little effect on grazing efficiency. This was largely due to the high amount of forage destroyed under the low density treatments which offset the effect of higher forage intake of that treatment. Within the 10-day duration treatment, average daily intake was the same over time (P> 0.05), while grazing efficiency decreased as grazing progressed (P< 0.05). The low grazing efficiency during the early stages of grazing reflected high initial forage destruction probably caused by the movement of animals at the start of grazing to establish bedding and habitual use areas. Growth rate of forage after grazing was highest in the longer duration paddocks and lowest in the shorter duration paddocks (P< 0.05), but the yield was similar under all treatments (P> 0.05). Although short duration/high density grazing is considered to be non-selective, sheep were equally or more selective under very short duration/very high density compared to longer duration/lower density treatments in this study. The 2 days duration was not an attractive management option since the intake and grazing efficiency were low, and the animals were selective. / Graduation date: 1993

A study was conducted on an annual cropped field near Lanigan, Saskatchewan over two years (2005-2006, 2006-2007) to evaluate the effects of three extensive winter feeding systems (bale grazing (BG), swath grazing (SG) and straw-chaff grazing (ST-CH)) and one intensive winter feeding system (drylot (DL)) on cow performance, soil nutrients, crop yield and system cost of production.<p> Differences in BW (P<0.05) were observed during the 2005-2006 study period with the greatest difference occurring with cows in the SG feeding system. Cows grazing swaths (SG) had a BW loss of 8.0 kg over the 78 d trial period, however these cows consumed 15% less DM and 13% less TDN than cows bale grazing, grazing crop residue or fed in drylot pens. Differences in BW change (P<0.05) were also observed during Yr 2 between the cows fed drylot and cows grazing barley straw-chaff, 32.9 and 6.5 kg, respectively. This difference in body weight change (BW∆) and lower TDN consumption may be attributed to inaccessibility of the straw-chaff feed in the field, due to inclement weather and would suggest a lengthy acclimation period for extensive field grazing systems.<p> The effects of extensive winter feeding system on soil nutrients and soil structure were determined the following spring after winter grazing. NO3-N levels at the low slope position in the 0-15 cm depth were 53% higher on the BG sites than the ST-CH sites. This may be attributed to the larger concentration of feed, thus feed nutrients, in the BG feeding system. Phosphorus levels on the BG wintering sites were 34% higher than levels in the SG or ST-CH sites. Crop biomass measured on the BG sites was consistent with soil nutrients captured, resulting in a 15% increase in biomass compared to ST-CH and SG sites. Soil nutrient and crop biomass distribution was consistent among winter grazing sites with the ST-CH sites having the most uniform distribution of nutrients and crop biomass, and the BG sites having the least.

Three studies were undertaken from 2006 to 2007 to examine litter (dead plant material) in southwestern Manitoba pastures. First, the relationship between litter and soil microclimate was tested across five pasture sites. The amount of litter biomass was not strongly related to soil moisture, though near-surface soil temperatures were reduced when litter was present. Second, the effect of four simulated grazing strategies on the litter layer was measured in six pastures. It was found that after three years of simulated grazing, litter was present in largest quantities in the least-frequently grazed treatments. Finally, a field survey was conducted assessing the quantity of litter present in native pastures across Manitoba. Litter was quite variable and averaged 1902 kg/ha over two years. This research confirmed the value of litter as an indicator of sustainable pasture management, though it remains unclear whether litter is important to pastures from the perspective of soil microclimate. / October 2008

A study was conducted on an annual cropped field near Lanigan, Saskatchewan over two years (2005-2006, 2006-2007) to evaluate the effects of three extensive winter feeding systems (bale grazing (BG), swath grazing (SG) and straw-chaff grazing (ST-CH)) and one intensive winter feeding system (drylot (DL)) on cow performance, soil nutrients, crop yield and system cost of production.<p> Differences in BW (P<0.05) were observed during the 2005-2006 study period with the greatest difference occurring with cows in the SG feeding system. Cows grazing swaths (SG) had a BW loss of 8.0 kg over the 78 d trial period, however these cows consumed 15% less DM and 13% less TDN than cows bale grazing, grazing crop residue or fed in drylot pens. Differences in BW change (P<0.05) were also observed during Yr 2 between the cows fed drylot and cows grazing barley straw-chaff, 32.9 and 6.5 kg, respectively. This difference in body weight change (BW∆) and lower TDN consumption may be attributed to inaccessibility of the straw-chaff feed in the field, due to inclement weather and would suggest a lengthy acclimation period for extensive field grazing systems.<p> The effects of extensive winter feeding system on soil nutrients and soil structure were determined the following spring after winter grazing. NO3-N levels at the low slope position in the 0-15 cm depth were 53% higher on the BG sites than the ST-CH sites. This may be attributed to the larger concentration of feed, thus feed nutrients, in the BG feeding system. Phosphorus levels on the BG wintering sites were 34% higher than levels in the SG or ST-CH sites. Crop biomass measured on the BG sites was consistent with soil nutrients captured, resulting in a 15% increase in biomass compared to ST-CH and SG sites. Soil nutrient and crop biomass distribution was consistent among winter grazing sites with the ST-CH sites having the most uniform distribution of nutrients and crop biomass, and the BG sites having the least.