Using metacommunity theory, I investigated the mechanisms of meta-assemblage structure and assembly among medium- to large-bodied mammals in North Texas. Mammals were surveyed with camera-traps in thirty property units of the LBJ National Grasslands (LBJNG). In Chapter II the dispersal and environmental-control based processes in community assembly were quantified within a metacommunity context and the best-fit metacommunity structure identified. A hypothesis-driven modelling approach was used in Chapter III to determine if the patterns of species composition and site use could be explained by island biogeography theory (IBT) or the habitat amount hypothesis (HAH). Islands were defined as the LBJNG property unit or the forest patch bounded by the property unit. Forest cover was selected as the focal habitat for the HAH. Seasonal dynamics were explored in both chapters. Metacommunity structure changed with each season, resulting in quasi-nested and both quasi and idealized Gleasonian and Clementsian structures. Results indicated that the anthropogenic development is, overall, not disadvantageous for this assemblage, that community assembly receives equal contributions from spatial and environmental factors, and that the metacommunity appears to operate under the mass effects paradigm. The patterns of species composition and site use were not explained by either IBT or HAH. Likely because this assemblage of generalist, dispersal-capable mammals are utilizing multiple habitat types both in the protected land and in the private land. This research highlights the versatility of these species and the potential value of rural countryside landscapes for wildlife conservation.

Understanding how habitat amount and configuration affect species richness, occurrence or abundance has been one of the major foci of research in ecology and biogeography, given its central importance for conservation planning and landscape management. We conducted bird point counts within clearcuts and mature pine stands of different sizes and configuration in working pine forest in north-central Mississippi to determine species associations. Early-successional and mature pine focal species showed varying response to the proportion and proximity of vegetation conditions in the landscape. While elements of configuration exhibited a greater influence on predicted avian abundance in this landscape, meaning many species require a mosaic of habitat conditions that come from both early-successional and mature vegetation types. Efforts to combine management of timber and conservation of songbirds must consider both species’ habitat requirements and the distribution of these requirements in the landscape.

The world-wide intensification of agriculture has led to a decline in species richness due to land use change, isolation, and fragmentation of natural and semi-natural habitats in agricultural and forestry landscapes. As a consequence, there is a current landscape management focus on the importance of green infrastructure to mitigate biodiversity decline and preserve ecosystem functions e.g. pollination services and pest control. Even though intensification in agriculture has been ongoing for several hundreds of years, remnant habitats from earlier management practices may still be remaining with a surprisingly high plant richness. Preserving these habitats could help conserving plant species richness in agricultural landscapes, as well as other organisms that are dependent on plants for food and shelter. In this thesis I focus on two small remnant habitats; midfield islets and borders between managed forest and crop field in southeastern Sweden. In the past, both habitats were included in the grazing system and therefore often still have remnant population of grassland specialist species left today. I have used these two remnant habitats as model habitats to investigate the effect of landscape factors and local factors on species richness of plants, flower morphologies and plants with fleshy fruits. Additively, I analysed the effect of surrounding landscape and local openness on the functions; pollination success, biological pest control of aphids and seed predation on midfield islets. One of my studies showed that spatial distribution and size of the habitat affected plant species richness. Larger habitat size and higher connectivity between habitats increased species richness of plants in the habitats. Openness of the habitats was shown to be an important factor to increase species richness and richness of flower morphologies, both on midfield islets and in forest borders. Even though midfield islets had the highest species and morphology richness, both habitat types are needed for habitat complementary as forest borders have more plants with fleshy fruits and a higher richness of plant species that flowers in spring/early summer. It was also shown that a more complex forest border, not just with gaps in the canopy, but also with high variation in tree stem sizes increases plant species richness in the field layer. The conclusion is that by managing small remnant habitats to remain or become more semi-open and complex in their structure, would increase species richness of plants, grassland specialist species, and flower morphologies. It would also increase some ecosystem functions as seed predation and biologic pest control of aphids are more effective close to trees. If both midfield islets and forest borders would be managed to be semi-open, the area and connectivity of semi-open habitat would increase in the agricultural landscape, which may also improve pollination success as the connectivity between populations has a possibility to increase. Grassland specialist species are clearly abundant in the small remnant habitats. As the decline of semi-natural grasslands is causing a decline in grassland specialists’ species, not only plants, I recommend that small remnant habitats are included in conservation and management plans and strategies to improve habitat availability and connectivity for grassland species in agricultural landscapes. / <p>Research funder Ekoklim. Project:4339602.</p><p>At the time of the doctoral defense, the following paper was unpublished and had a status as follows: Paper 4: Manuscript.</p>