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The variations in discharge and water chemistry among and within headwater catchments are not well understood. Developing a better understanding of the processes that control these variations is crucial to determining how headwater catchments will respond to changes in climate and land use. This dissertation explores how hydrologic processes in headwater catchments may be better understood by utilizing a hydropedological framework, where similar soils are grouped together and considered to be representative of and developed by similar hydrologic and biogeochemical processes. In the first chapter, soil groups, called hydropedological units (HPUs) are found to be indicative of distinct water table regimes characterized by the interquartile range and median of shallow groundwater levels, the percent time water table exists in the soil, and the level of catchment storage at which groundwater responds. The second chapter explores the hydrological processes that may lead to the formation of HPUs in the catchment. By examining water table records and unsaturated water potential from tensiometers we found that lateral unsaturated flow regimes may be partially responsible for the patterns of lateral translocation observed in HPUs. Finally, the third chapter identifies two HPUs in the catchment as sources of streamwater dissolved organic carbon (DOC). While near-stream areas have typically been found to be DOC sources in headwater catchments, the HPUs identified as sources occur at high elevations in the catchment, near channel heads. Overall, these findings will be useful to better explain runoff generation, soil formation, and DOC export from headwater catchments. Headwater streams source water to larger bodies of water that are valuable natural resources. Therefore, explaining these processes is critical to predicting and responding to changes in climate and land use that may affect important water supplies. / Ph. D.

I studied how geology, land use and rainfall, correlated with peak flow responses in 110 headwater stream sites during a drought year. Highest discharges were observed in the most developed catchments and in the most poorly drained soils, but specific responses were variable depending on both geology and land disturbance. Redundancy analysis indicated that both surficial geology and land disturbance were important predictors of discharge and that rainfall was in general a poor predictor of discharge. I conclude that responses of headwater streams to individual storms are unpredictable from data generated using GIS, but increased peak flows occur associated with human development, mitigated by surficial geology. The headwater streams that are most vulnerable to flow alterations occur on poorly drained soils, and where urbanization tends to concentrate. Much greater attention to managing water is required if further degradation of stream ecosystems is to be prevented from our future land use.

I studied how geology, land use and rainfall, correlated with peak flow responses in 110 headwater stream sites during a drought year. Highest discharges were observed in the most developed catchments and in the most poorly drained soils, but specific responses were variable depending on both geology and land disturbance. Redundancy analysis indicated that both surficial geology and land disturbance were important predictors of discharge and that rainfall was in general a poor predictor of discharge. I conclude that responses of headwater streams to individual storms are unpredictable from data generated using GIS, but increased peak flows occur associated with human development, mitigated by surficial geology. The headwater streams that are most vulnerable to flow alterations occur on poorly drained soils, and where urbanization tends to concentrate. Much greater attention to managing water is required if further degradation of stream ecosystems is to be prevented from our future land use.

The aims of this theses were (i) to evaluate trends in spring yields in the Czech Republic (period 1971-2007) and trends in runoff from small forested catchments of the GEOMON network during the period 1994-2011, (ii) evaluate the impact of climate change on streamwater chemistry in acid-sensitive catchment (Lysina), and (iii) estimate the impact of anticipated climate change projected according to different CO2 emission scenarios on flow patterns forested (GEOMON) and alpine headwater (Skalnaté Lake) catchments in the periods 2021-2050 and 2071-2100. Significant negative trends on the annual level were observed at 18% of springs while positive trends at only less than 0.5% of 157 tested objects and 4 of 18 regions revealed significant decreasing trend. To these regions belonged the zones of Carpathian Cretaceous and Paleogene sediments in the southeast and northeast of the Czech Republic, the zones of tertiary and Cretaceous basins in the southeast and Proterozoic and Paleozoic crystalline rocks in the central part of the Czech Republic. Most spring yields were decreasing within the summer months June- August (on average 24%) and least between February and April (on average 15%) (Paper I). There were no general patterns found indicating either significant increases or decreases in runoff on either...

<p>Fish communities in agricultural headwater streams are known to be impacted by a variety of factors, including water chemistry, habitat modification, and hydrology. Little research has been conducted on how climate change influences these communities, yet the effects of climate on lake and river fish have been well documented. I hypothesized that fish community metrics would be reduced by the effects of climate change. I examined the effects of climate and hydrology metrics on fish communities at nine sites in the Saint Joseph River, Indiana and Michigan and at 18 sites in the Upper Big Walnut Creek, Ohio watersheds, from 2006 to 2019. Air temperature, water temperature, precipitation, water discharge, width, velocity, and depth metrics were calculated seasonally for each sampling year. Fish were examined seasonally with backpack electrofishing and seine netting and identified to species level. Principal component analyses were used to create axes which represented gradients of climate and hydrology metrics. Linear mixed effect and logistic regression modeling suggested that hydrology is a stronger predictor than climate, but that both influence fish communities. Percent Percidae, percent herbivore, and percent open substrate spawner were positively correlated with precipitation and water temperature. Presence herbivore was negatively correlated with precipitation and positively correlated with water temperature. My data only somewhat supported the hypothesis that climate would reduce fish community metrics. Gradients of hydrology were observed to be stronger predictors than gradients of climate. However, one must acknowledge relationships between climate and hydrology and the potential for climate to have indirect effects on fish communities through influences on hydrology. This study increases understanding of how fish communities in agriculturally dominated headwater streams are influenced, and emphasizes the need for further research on how these fishes will be impacted by a changing climate. </p>

North American prairie headwater streams are highly threatened and relatively poorly studied. Most studies on prairie streams have occurred at the Konza Prairie Biological Station, a tallgrass prairie Long Term Ecological Research site in the Flint Hills ecoregion near Manhattan, KS. According to the Stream Biome Gradient Concept, several ecosystem factors vary along a gradient from more allochthonous forested streams to more autochthonous desert streams, with grassland streams often intermediate in several key ecological factors including litter inputs, primary production, and invertebrate abundance and biomass. However, few studies have examined the degree of variation that exists within prairie headwater streams, and whether this variation occurs along a longitudinal gradient as well, with more mesic tallgrass prairie streams differing from more xeric shortgrass prairie streams, and mixed-grass sites intermediate between the two. I examined thirteen prairie headwater stream sites in the central United States from 2014 to 2017. My objective was to determine whether basal resource composition – including standing stocks of coarse particulate organic matter (CPOM), fine particulate organic matter (FPOM), and very fine particulate organic matter (VFPOM), sestonic and benthic chlorophyll-a levels, and sources of CPOM – differed significantly among streams in tallgrass, mixed-grass, and shortgrass prairie regions. In addition, I examined whether invertebrate communities differed among tallgrass, mixed-grass, and shortgrass prairie regions, and whether this was reflected in the functional feeding group composition, habit composition, voltinism, and dispersal ability of invertebrate communities. There were no significant differences in total CPOM, FPOM, and VFPOM standing stocks among regions. However, CPOM composition did differ with region, with tallgrass sites having higher standing stocks of leaf litter, but lower standing stocks of grass litter and macrophyte litter than the other regions. Benthic chlorophyll-a did not differ significantly among regions, but there were lower sestonic chlorophyll-a levels in tallgrass sites. Given higher light availability and nutrient levels in shortgrass and mixed-grass streams, lack of stable substrata may be limiting benthic algae in these regions. Invertebrate abundance and biomass were highest in mixed-grass sites and lowest in tallgrass sites, with shortgrass sites intermediate. Mixed-grass sites also had significantly higher Shannon diversity and taxa richness than tallgrass sites. A NMDS revealed that sites differed in overall community structure. Functional feeding group composition did not differ significantly across regions, with collector-gatherers, followed by predators, dominating in all sites. High Predator-Prey Index (predator biomass: other invertebrate biomass) values in each region indicates strong top-down pressure and high turnover rates of prey taxa. While there was a weak correlation between leaf litter AFDM and invertebrate community structure, the correlation between latitude and longitude and invertebrate community structure was much higher, indicating that basal resources may not be the main drivers in these systems. Invertebrate habit composition did not differ with region; all regions were dominated by taxa preferring fine substrata (burrowers and sprawlers). All regions showed selection against semivoltine invertebrates and were dominated by high dispersing insect taxa, though the proportion of the insect community that consisted of high dispersers did not differ significantly with region. My results suggest that generalizing about prairie streams based on studies from one or a handful of sites may not be prudent, at least for some aspects. The higher proportion of autochthonous inputs in shortgrass and mixed-grass regions may drive increased invertebrate abundance, biomass, richness, and diversity, but the relative hydrologic stability of the mixed-grass sites might also explain these results or have an interactive or additive relationship with primary production. Overall, the link between basal resources and communities across prairie types was somewhat weak, and all regions were dominated by collector-gatherers with rapid life-cycles and high dispersal abilities, indicating that disturbance may be a more important community filter than basal resource composition. Streams in all three regions have highly variable hydrology, and this may be an overriding factor that results in similarity in communities.

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