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<p>Arthropods have a rich fossil record spanning the Phanerozoic. Biomineralized forms such as the extinct trilobites are particularly common and are proven index fossils for biostratigraphy. Forms with an unmineralized cuticle are more rare, preserved only in so called <i>konservat lagerstätten</i>. Cambrian strata of Greenland have yielded rich trilobite faunas with potential for intercontinental correlation of Cambrian strata, but also an exceptionally preserved fauna, the Sirius Passet Lagerstätte. The first part of this thesis is concerned with trilobite biotratigraphy of the provisional Cambrian Series 2 in Greenland. The second part is concerned with exceptionally preserved arthropods from the Sirius Passet Lagerstätte, but also from 'Orsten' deposits from the Cambrian of Sweden.</p><p><i>Perissopyge phenax occurs</i> in the Henson Gletscher and Paralleldal formations spanning the Series 2 and 3 boundary interval in North Greenland. It also occurs in the Sekwi Formation of Yukon Territory, demonstrating that the species may hold potential for correlation within Laurentia. An indeterminate species of <i>Perissopyge</i> is shown to occur in the Ella Island Formation of North-East Greenland together with <i>Olenellus</i> cf. <i>hanseni</i>, which is similar to <i>Olenellus</i> cf. <i>truemani</i> described from the Henson Gletscher Formation. If this correlation is further corroborated it would offer a first tie-point for the An t'Sron Formation of North-West Scotland which yields <i>Fritzolenellus</i> <i>lapworthi</i>, herein reported for the first time from the Bastion Formation which underlies the Ella Island Formation.</p><p><i>Oelandocaris</i> <i>oelandica</i> from ‘Orsten’ deposits in the Cambrian series 3 and 4 boundary interval in Sweden is an early representative of the Crustacean stem lineage. <i>Kiisortoqia</i> <i>avannaarsuensis</i> is a new arthropod from the Sirius Passet Lagerstätte with robust antennulae strikingly similar to the 'raptorial' limb of the problematic anomalocaridids. The ventral morphology of the 'bivalved' <i>Isoxys</i> <i>volucris</i> is described for the first time and compared with other species assigned to <i>Isoxys</i> from Cambrian lagerstätten around the world. Finally, <i>Siriocaris</i> <i>trolla</i>, is a new arthropod that similarities with trilobites and certain ‘trilobitomorphs’ but seems to lack important synapomorphies of these taxa, though this may be due to preservational limitations in the material at hand.</p>

Approximately 600 to 500 million years ago, a major evolutionary radiation called the “Cambrian Explosion” gave rise to nearly all of the major animal phyla known today. This radiation is recorded by various fossil lagerstätten, such as the Burgess Shale in Canada, where soft-bodied animals are preserved in exquisite detail. Many Cambrian fossils are enigmatic forms that are morphologically dissimilar to their modern descendants, but which still provide valuable information when interpreted as stem-group taxa because they record the actual progression of evolution and give insight into the order of character acquisitions and homologies between living taxa. One such group of fossils is the anomalocaridids, large presumed predators that have had a complicated history of description. Their body has a trunk with a series of lateral lobes and associated gills, and a cephalic region with a pair of large frontal appendages, a circular mouth apparatus, stalked eyes and a cephalic carapace. Originally, two taxa were described from the Burgess Shale, Anomalocaris and Laggania, however data presented herein suggests that the diversity of the anomalocaridids was much higher. Newly collected fossil material revealed that a third Burgess Shale anomalocaridid, Hurdia, is known from whole-body specimens and study of its morphology has helped to clarify the morphology and systematics of the whole group. Hurdia is distinguished by having mouthparts with extra rows of teeth, a unique frontal appendage, and a large frontal carapace. Two species, Hurdia victoria and Hurdia triangulata were distinguished based on morphometric shape analysis of the frontal carapace. A phylogenetic analysis placed the anomalocaridids in the stem lineage to the euarthropods, and examination of Hurdia’s well-preserved gills confirm the homology of this structure with the outer branches of limbs in upper stem-group arthropods. This homology supports the theory that the Cambrian biramous limb formed by the fusion of a uniramous walking limb with a lateral lobe structure bearing gill blades. In this context, new evidence is present on the closely allied taxon Opabinia, suggesting that it had lobopod walking limbs and a lateral lobe structure with attached Hurdia-like gills. The diversity of the anomalocaridids at the Burgess Shale is further increased by two additional taxa known from isolated frontal appendages. Amplectobelua stephenensis is the first occurrence of this genus outside of the Chengjiang fauna in China, but Caryosyntrips serratus is an appendage unique to the Burgess Shale. To gain a better understanding of global distribution, a possible anomalocaridid is also described from the Sirius Passet biota in North Greenland. Tamisiocaris borealis is known from a single appendage, which is similar to Anomalocaris but unsegmented, suggesting this taxon belongs to the arthropod stem-lineage, perhaps in the anomalocaridid clade. Thus, the anomalocaridids are a widely distributed and highly diverse group of large Cambrian presumed predators, which provide important information relevant to the evolution of the arthropods.

Arthropods have a rich fossil record spanning the Phanerozoic. Biomineralized forms such as the extinct trilobites are particularly common and are proven index fossils for biostratigraphy. Forms with an unmineralized cuticle are more rare, preserved only in so called konservat lagerstätten. Cambrian strata of Greenland have yielded rich trilobite faunas with potential for intercontinental correlation of Cambrian strata, but also an exceptionally preserved fauna, the Sirius Passet Lagerstätte. The first part of this thesis is concerned with trilobite biotratigraphy of the provisional Cambrian Series 2 in Greenland. The second part is concerned with exceptionally preserved arthropods from the Sirius Passet Lagerstätte, but also from 'Orsten' deposits from the Cambrian of Sweden. Perissopyge phenax occurs in the Henson Gletscher and Paralleldal formations spanning the Series 2 and 3 boundary interval in North Greenland. It also occurs in the Sekwi Formation of Yukon Territory, demonstrating that the species may hold potential for correlation within Laurentia. An indeterminate species of Perissopyge is shown to occur in the Ella Island Formation of North-East Greenland together with Olenellus cf. hanseni, which is similar to Olenellus cf. truemani described from the Henson Gletscher Formation. If this correlation is further corroborated it would offer a first tie-point for the An t'Sron Formation of North-West Scotland which yields Fritzolenellus lapworthi, herein reported for the first time from the Bastion Formation which underlies the Ella Island Formation. Oelandocaris oelandica from ‘Orsten’ deposits in the Cambrian series 3 and 4 boundary interval in Sweden is an early representative of the Crustacean stem lineage. Kiisortoqia avannaarsuensis is a new arthropod from the Sirius Passet Lagerstätte with robust antennulae strikingly similar to the 'raptorial' limb of the problematic anomalocaridids. The ventral morphology of the 'bivalved' Isoxys volucris is described for the first time and compared with other species assigned to Isoxys from Cambrian lagerstätten around the world. Finally, Siriocaris trolla, is a new arthropod that similarities with trilobites and certain ‘trilobitomorphs’ but seems to lack important synapomorphies of these taxa, though this may be due to preservational limitations in the material at hand.

The Pachycormiformes were a successful group of stem-teleosts. Although they persisted for more than 100 million years in the Mesozoic seas and occupied a significant space of the ecosystem, little is known about this most diverse group. One of the earliest pachycormiformes is the lower Jurassic Pachycormus bollensis. A reconstruction of the phylogeny with the early P. bollensis together with more derived and earlier studied species has yielded important information about the relationships within the group. Both a parsimony analysis and a Bayesian analysis were performed. Three exceptionally complete specimens of P. bollensis from the Holzmaden-locality were used for the study. The resulting trees strongly supported pachycormiform monophyly. Three major ecomorphological clades were returned from the analyses: the filter-feeders, hyper carnivores and a more generalist predator radiation that included P. bollensis. Despite, node support within the pachycormiformes are generally weak. The tooth-structure and the phylogenetic position of P. bollensis might indicate an intermediate grade between the filter-feeders and the hyper carnivores. / Pachycormiformer var en mycket framgångsrik ordning utav stam-teleoster. Trots att de överlevde i mer än 100 miljoner år i de Mesosoiska haven och erövrat ett flertal olika platser i ekosystemet, är denna mågfacetterade grupp som helhet tämligen okänd. Pachycormus bollensis från yngre jura är en av de tidigste arterna i denna grupp. En fylogenetiskt rekonstruktion av den tidiga P. bollensis tillsammans med yngre, mer utvecklade arter kan ge viktig information om släktskapen inom gruppen. En parsimonisk analys och en Bayesian analys utfördes. Tre exceptionellt bevarade exemplar av Pachycormus bollensis från Holzmaden användes i studien. Resultatet visar på ett stark monofyletiskt sammanband inom pachycormiformerna. Båda analyserna visade robusta resultat för tre övergripande grupper; filtrerare, karnivorer och en mer generell predator linje som inkluderar P.  bollensis. Dock hade de interna släktskapen inom varje större grupp svagt stöd. Tand-stukturen tillsammans med den fylogenetiska placeringen av P. bollensis kan tyda på en intermediär grupp mellan filterarna och karnivorerna.

This study aims to unravel the ecological and evolutionary dynamics within the calcareous nannofossil communities at the Eocene/Oligocene (E/O) transition and during the Oligocene time when Cenozoic 'icehouse' conditions were established. The main question this study aims to answer is whether the changes in the nannofossil assemblages were controlled by intrinsic evolutionary trends or if the changes were controlled by environmental factors such as changes in temperature and nutrient availability in the surface water. These questions are addressed with detailed analyses of the taxonomy, biostratigraphy and fluctuations in abundance and diversity of calcareous nannofossil assemblages from a continuous latest Eocene through Oligocene sediment section from the ODP Site 1218 (8°53.38´N, 135°22.00´W), paleo-equatorial Pacific Ocean. An improved nannofossil taxonomy and biostratigraphy has been established. At the E/O transition, changes in the nannofossil preservation mimics changes in calcium carbonate content. A detailed investigation of late Eocene and Oligocene sediments yields age estimates for ten nannofossil bioevents. Morphometric studies of the Reticulofenestra umbilicus-R. hillae show that these cannot be subdivided into two different morphospecies. Based on different morphometry and stratigraphic ranges, the Oligocene Sphenolithus lineage appears to be the result of a combined anagenetic and cladogenetic evolution. A new nannolith species is described: Triquetrorhabdulus longus. High-resolution nannofossil data indicate changes in the composition, abundance and diversity of the mid-Oligocene assemblages. Intervals of high diversity index coincide with Oi-glaciation events. However, visual examination of the variations in abundance of nannofossil taxa do not appear to correlate with changes in either oxygen or carbon isotopes. This presumably indicates that a dynamic equilibrium did not exist between these Oligocene nannoplankton assemblages and changes in surface water temperature or productivity conditions. / Doctoral Thesis in Marine Geoscience at Stockholm University, Sweden 2009