A major argument for incorporating deciduous tree species in coniferous forest stands is their role in the amelioration and stabilisation of biogeochemical cycles. Current forest management strategies in central Europe aim to increase the area of mixed stands. In order to formulate statements about the ecological effects of mixtures, studies at the stand level are necessary. In a mixed stand of Norway spruce (Picea abies (L.) Karst.) and silver birch (Betula pendula Roth) in the Ore Mountains (Saxony, Germany), the effects of these two tree species on chemical and microbial parameters in the topsoil were studied at one site in the form of a case study. Samples were taken from the O layer and A horizon in areas of the stand influenced by either birch, spruce or a mixture of birch and spruce. The microbial biomass, basal respiration, metabolic quotient, pH-value and the C and N contents and stocks were analysed in the horizons Of, Oh and A. Significantly higher contents of microbial N were observed in the Of and Oh horizons in the birch and in the spruce-birch strata than in the stratum containing only spruce. The same was found with respect to pH-values in the Of horizon and basal respiration in the Oh horizon. Compared to the spruce stratum, in the birch and spruce-birch strata, significantly lower values were found for the contents of organic C and total N in the A horizon. The findings of the case study indicated that single birch trees have significant effects on the chemical and microbial topsoil properties in spruce-dominated stands. Therefore, the admixture of birch in spruce stands may distinctly affect nutrient cycling and may also be relevant for soil carbon sequestration. Further studies of these functional aspects are recommended.

Das Forschungsvorhaben hat das Ziel den Einfluss einzelbaumweise eingemischter einheimischer Eichen in Wäldern der Gemeinen Kiefer (Pinus sylvestris L.) zu untersuchen. Die Untersuchungsflächen befinden sich im Nordosten Deutschlands (Brandenburg). Im Untersuchungsgebiet Jerischke hat die Kiefer ein Alter von 78 Jahren und die Eiche ein Alter von 205 Jahren. Im Untersuchungsgebiet Weidmannsruh befindet sich die Kiefer im Alter 70 und die Eiche im Alter 108. In Abhängigkeit von der Entfernung zur Eiche werden folgende Effekte der Eiche nach Intensität und Qualität untersucht: - Effekte auf Mikroklima und Strahlung, - Effekte auf Humusmorphologie und bodenchemische Parameter und - Effekte auf Bodenvegetation und epigäische Fauna. Grundsätzlich sind die positiven Effekte der Eiche auf 10 bis 15 m vom Eichenstamm in den Kiefernbestand hinein begrenzt.

Made available in DSpace on 2015-03-26T13:53:22Z (GMT). No. of bitstreams: 1 texto completo.pdf: 1008368 bytes, checksum: 5118654c44cc454b278b6c83f243c20f (MD5) Previous issue date: 2012-02-15 / Conselho Nacional de Desenvolvimento Científico e Tecnológico / In Brazil, eucalyptus is planted in regions with different climate and soil conditions, which affect harvest residue decomposition and the rate of carbon transference fromresidues (Cres) to the soil, hence influencing soil organic matter (SOM) content. SOM is considered one of the most important indicators of soil quality and forest production sustainability. Debarking the trunk of the tree during forest harvesting contributes to increase the amount of residues left in the site, but it may impair residues decomposition due to the large C:N ratio of the material. However, very little is known about bark influence on eucalypt residue decomposition and its role on SOM formation under the Brazilian conditions. It is a common practice the use of low rates of N fertilizers in eucalypt plantations in Brazil, which would contribute to slower microbial activity in the decomposition process. Leaving the residues on the surface without incorporating them into the soil would also help to limit Cres to SOM. Hence, this study has the objective to determine the influence of leaving on soil surface eucalypt bark and slash in site after harvesting operations as wells as the effect of nitrogen fertilizer addition on the rates of residue decomposition and Cres to the SOM, and how climate and soil characteristics affect these processes. The experiment was installed in four sites (Eunápolis, Aracruz, Virginópolis and Belo Oriente) belonging to the Atlantic Forest biome and six (João Pinheiro, Três Marias, Curvelo, Itamarandiba e Luiz Antônio) belonging to the Cerrado (savanah type of vegetation), all varying in climate and soil characteristics. In each site the 12 treatments (3 slash handling systems, i.e., complete removal, left in the site with trunk debarking or not; 2 slash disposal systems, i.e., left on soil surface or incorporated into the soil; and 2 rates of N, i.e., zero or 200 kg ha-1) were laid out in random blocks with 4 replications. The remaining dry matter was quantified at 0, 3, 6, 12 and 36 months later, and estimated the decomposition constant (k) and half life time (t0,5) of the residues. Residue Ct to SOM was assessed 12 months after starting the trial, by soil physical fractionation to separate particulate organic matter (POM) and organic matter associated to soil minerals (MAM). The influence of climate and soil characteristics on t0,5 and Cres was estimated by linear multiple regression. Residue incorporation into the soil favored both the decomposition and Cres. Decomposition was enhanced by debarking but less Cres was detected. Nitrogen addition had little effect on the residues decomposition rate and SOM stabilization. Residues t0,5 was shorter in sites with higher rainfall regimes and mean annual temperatures and residues incorporation, mainly in sandy soils. The mean residue t0,5 was 1.28 year. Cres is reduced with increasing mean temperature and soil acidity and enhanced by increased clay content and deficit of C saturation of the soil. In average, the rate of C recovering was 15 % from the decomposed residues, in 12 months. / No Brasil as plantações de eucalipto são estabelecidas em regiões com diferentes condições edafoclimáticas que afetam a decomposição de resíduos da colheita e a transferência de C (Cres) deste material para a matéria orgânica do solo (MOS), que é considerada um dos principais indicadores da qualidade do solo e sustentabilidade da produção. Nestas plantações, o descascamento das árvores no campo proporciona maior aporte de compostos recalcitrantes e maior relação C:N dos resíduos, embora pouco se sabe sobre a influência da presença da casca na decomposição deste material e formação da MOS. Além disso, as baixas doses de N aplicadas nestas plantações e a manutenção dos resíduos na superfície do solo podem limitar a sua decomposição e Cres estabilizado na MOS e, por isso, a aplicação de N e incorporação deste material ao solo poderiam acelerar sua taxa de decomposição e favorecer a transferência de C para o solo. Os principais objetivos do presente estudo foram determinar a influência da manutenção da casca, da aplicação de N e da incorporação dos resíduos para a decomposição e transferência de C para a MOS. Além disso, objetivou-se determinar os principais fatores edafoclimáticos que controlam estes processos em condições de campo. Os experimentos foram instalados em áreas de Mata Atlântica, nos municípios de Eunápolis (BA), Aracruz (ES), Virginópolis (MG), Belo Oriente (MG) e áreas de Cerrado, nos municípios de João Pinheiro (MG), Três Marias (MG), Curvelo (MG), Itamarandiba (MG), Vazante (MG) e em Luiz Antônio (SP). Foi utilizado o delineamento experimental em blocos casualizados com quatro repetições. Em cada local o experimento consistiu de 12 tratamentos em esquema fatorial completo 3 X 2 X 2, incluindo os fatores resíduos da colheita de eucalipto (remoção e manutenção de resíduos sem e com casca), manejo dos resíduos (superficial e incorporados) e aplicação de N (0 e 200 kg ha-1). Para o referido estudo foi avaliada a massa de matéria seca remanescente após 0, 3, 6, 12 e 36 meses, estimados a taxa de decomposição (k) e os valores de tempo de meia-vida (t0,5) dos resíduos de eucalipto. A transferência de C deste material para a MOS foi avaliada após 12 meses, por meio do fracionamento físico do solo para a separação da matéria orgânica particulada (MOP) e associada aos minerais (MAM). A influência das variáveis climáticas e propriedades do solo sobre o t0,5 e Cres para o solo foi determinada por regressão linear múltipla. A incorporação dos resíduos favorece tanto a decomposição quanto a transferência de C dos resíduos para o solo. A partir da manutenção da casca a decomposição dos resíduos é acelerada e relativamente ocorre menor recuperação de C no solo. Tanto a decomposição quanto a estabilização da MOS são pouco influenciadas pela aplicação de N. O t0,5 dos resíduos de eucalipto é reduzido quando ocorrem maiores volumes de precipitação, temperaturas mais elevadas e incorporação dos resíduos, principalmente em solos arenosos. Em média, o t0,5 dos resíduos de eucalipto foi de 1,28 anos. A Cres para o solo é reduzida pelo aumento da temperatura e redução da acidez do solo e favorecida por maiores teores de argila e déficit de saturação de C. Em média, a taxa de recuperação de C dos resíduos no solo em 12 doze meses foi de 15 % do total decomposto.

A major argument for incorporating deciduous tree species in coniferous forest stands is their role in the amelioration and stabilisation of biogeochemical cycles. Current forest management strategies in central Europe aim to increase the area of mixed stands. In order to formulate statements about the ecological effects of mixtures, studies at the stand level are necessary. In a mixed stand of Norway spruce (Picea abies (L.) Karst.) and silver birch (Betula pendula Roth) in the Ore Mountains (Saxony, Germany), the effects of these two tree species on chemical and microbial parameters in the topsoil were studied at one site in the form of a case study. Samples were taken from the O layer and A horizon in areas of the stand influenced by either birch, spruce or a mixture of birch and spruce. The microbial biomass, basal respiration, metabolic quotient, pH-value and the C and N contents and stocks were analysed in the horizons Of, Oh and A. Significantly higher contents of microbial N were observed in the Of and Oh horizons in the birch and in the spruce-birch strata than in the stratum containing only spruce. The same was found with respect to pH-values in the Of horizon and basal respiration in the Oh horizon. Compared to the spruce stratum, in the birch and spruce-birch strata, significantly lower values were found for the contents of organic C and total N in the A horizon. The findings of the case study indicated that single birch trees have significant effects on the chemical and microbial topsoil properties in spruce-dominated stands. Therefore, the admixture of birch in spruce stands may distinctly affect nutrient cycling and may also be relevant for soil carbon sequestration. Further studies of these functional aspects are recommended.

<p>Predmet&nbsp; istraživanja&nbsp; ove&nbsp; doktorske&nbsp; disertacije&nbsp; je&nbsp; procena&nbsp; potencijalno&nbsp; biodostupne&nbsp; frakcije odabranih&nbsp; prioritetnih&nbsp; organskih&nbsp; polutanata&nbsp;&nbsp; (pentahlorbenzena,&nbsp; heksahlorbenzena,&nbsp; lindana, trifluralina, 4-oktilfenola i 4-nonilfenola) u sedimentu, ispitivanje biodegradacionog potencijala ovih&nbsp; jedinjenja&nbsp; u&nbsp; sedimentu&nbsp; u&nbsp; različitim&nbsp; uslovima,&nbsp; kao&nbsp; i&nbsp; procena&nbsp; potencijala&nbsp; stabilizacije<br />sedimenta&nbsp; dodatkom&nbsp; ugljeničnih&nbsp; sorpcionih&nbsp; agenasa.&nbsp; U&nbsp; cilju&nbsp; razvoja&nbsp; metoda&nbsp; za&nbsp; određivanje biodostupnosti odabranih organskih polutanata ispitivane su i&nbsp; optimizovane metode vi&scaron;estepene i jednostepene parcijalne ekstrakcije primenom sledećih hemijskih agenasa:&nbsp; Tenaks&nbsp; smole, XAD-4&nbsp; smole&nbsp; i&nbsp; rastvora&nbsp; ciklodekstrina&nbsp; (2-hidroksipropil-&beta;-ciklodekstrina,&nbsp; &beta;-ciklodekstrina&nbsp; i&nbsp; metil-&beta;-ciklodekstrina).&nbsp; Optimalni&nbsp; agens&nbsp; za&nbsp; procenu&nbsp; biodostupne&nbsp; frakcije&nbsp; odabranih&nbsp; organskih zagađujućih&nbsp; materija&nbsp; sa&nbsp; sedimenta&nbsp; XAD-4&nbsp; smola,&nbsp; a&nbsp; optimalno&nbsp; vreme&nbsp; ekstrakcije&nbsp; primenom jednostepenih&nbsp; ekstrakcija&nbsp; je&nbsp; oko&nbsp; 8&nbsp; h.&nbsp; Procena&nbsp; biodegradacionog&nbsp; potencijala&nbsp; odabranih prioritetnih&nbsp; organskih&nbsp; zagađujućih&nbsp; materija&nbsp; u&nbsp; sedimentu&nbsp; ispitana&nbsp; je&nbsp; u&nbsp; različitim&nbsp; aerobnim&nbsp; i anaerobnim uslovima uz optimizaciju uslova putem biostimulacije i bioaugmentacije. Pokazano je&nbsp; da&nbsp; svih&nbsp; &scaron;est&nbsp; odabranih&nbsp; jedinjenja&nbsp; poseduju&nbsp; značajan&nbsp; potencijal&nbsp; biodegradacije&nbsp; u&nbsp; anaerobnoj sredini&nbsp; pri&nbsp; čemu&nbsp; u&nbsp; slučaju&nbsp; lindana&nbsp; i&nbsp; trifluralina&nbsp; dolazi&nbsp; do&nbsp; potpunog&nbsp; uklanjanja&nbsp; biodostupne količine&nbsp; jedinjenja&nbsp; primenom&nbsp; sva&nbsp; četiri&nbsp; ispitana&nbsp; inokuluma.&nbsp; Aerobni&nbsp; potencijal&nbsp; u&nbsp; ispitanim uslovima&nbsp; pokazali&nbsp; su&nbsp; samo&nbsp; alkil&nbsp; fenoli,&nbsp; gde&nbsp; je&nbsp; takođe&nbsp; uklonjena&nbsp; skoro&nbsp; celokupna&nbsp; količinabiodostupne&nbsp; frakcije&nbsp; jedinjenja&nbsp; u&nbsp; sedimentu&nbsp; (78-85%).&nbsp; Potencijal&nbsp; remedijacije&nbsp; sedimenta dodatkom&nbsp; ugljeničnih&nbsp; materijala&nbsp; ispitan&nbsp; je&nbsp; sa&nbsp; aspekta&nbsp; odabira&nbsp; ugljeničnih&nbsp; sorpcionih&nbsp; agenasa (aktivni&nbsp; ugalj,&nbsp; biougalj&nbsp; i&nbsp; humus);&nbsp; određivanja&nbsp; optimalne&nbsp; količine&nbsp; materijala;&nbsp; isptivanja dugoročnih i kratkoročnih efekata dodatka ovih agenasa na biodostupnost organskih zagađujućih materija&nbsp; kako&nbsp; bi&nbsp; se&nbsp; ispitao&nbsp; efekat&nbsp; starenja &nbsp; i&nbsp; toksičnosti&nbsp; dobijenih&nbsp; sme&scaron;a.&nbsp; Rezultati&nbsp; stabilizacije zagađujućih&nbsp; supstanci&nbsp; u&nbsp; sedimentu&nbsp; pokazuju&nbsp; da:&nbsp; (1)&nbsp; povećanje&nbsp; doze&nbsp; sva&nbsp; tri&nbsp; sorpciona&nbsp; agensa dovodi&nbsp; do&nbsp; povećanja&nbsp; efikasnosti&nbsp; imobilizacije&nbsp; i&nbsp; smanjenja&nbsp; biodostupne&nbsp; frakcije&nbsp; odabranih organskih jedinjenja; (2) starenjem sme&scaron;a sedimenta i&nbsp; sorbenata u toku prvih 90 dana dolazi do daljeg smanjenja biodostupne frakcije svih jedinjenja, nakon čega se biodostupna koncentracija ispitivanih&nbsp; jedinjenja&nbsp; primenom&nbsp; aktivnog&nbsp; uglja&nbsp; ne&nbsp; menja,&nbsp; dok&nbsp; primenom&nbsp; biouglja&nbsp; i&nbsp; humusa&nbsp; dolazi&nbsp; do&nbsp; porasta&nbsp; biodostupne&nbsp; frakcije&nbsp; jedinjenja.&nbsp; Testovi&nbsp; fitotoksičnosti&nbsp; su&nbsp; pokazali&nbsp; da&nbsp; je&nbsp; <em>Zea mays&nbsp; </em>akumulirao&nbsp; značajno&nbsp; veće&nbsp; količine&nbsp; ispitivanih&nbsp; jedinjenja&nbsp; iz&nbsp; netretiranog&nbsp; sedimenta&nbsp; u poređenju sa&nbsp; <em>Cucurbita pepo</em>&nbsp; i&nbsp;<em> Lactuca sativa. </em>Toksičnost sme&scaron;a sedimenta sa aktivnim ugljom i humusom&nbsp; procenjena&nbsp; na&nbsp; osnovu&nbsp; inhibicije&nbsp; luminiscencije&nbsp; na&nbsp; <em>Vibrio&nbsp; fischeri</em>&nbsp; kao&nbsp; i&nbsp; ispitivanjem <em>Zea&nbsp; mays&nbsp; </em>germinacije&nbsp; i&nbsp; produkcije&nbsp; biomase&nbsp; je&nbsp; pokazala&nbsp; značajno&nbsp; smanjenje&nbsp; u&nbsp; odnosu&nbsp; na netretirani&nbsp; sediment.&nbsp; Akumulacija&nbsp; ispitivanih&nbsp; jedinjenja&nbsp; u&nbsp; biomasi&nbsp; <em>Zea&nbsp; mays&nbsp; </em>u&nbsp; netretiranom sedimentu je bila značajno veća u odnosu na sve sme&scaron;e sedimenta i aktivnog uglja i humusa. Sva tri&nbsp; sorbenta&nbsp; pokazala&nbsp; su&nbsp; veliki&nbsp; remedijacioni&nbsp; potencijal&nbsp; za&nbsp; sediment&nbsp; zagađen&nbsp; organskim zagađujućim supstancama, ali je aktivni ugalj pokazao najbolje performance.</p> / <p>The&nbsp; aim&nbsp; of&nbsp; the&nbsp; research&nbsp; in&nbsp; this&nbsp; PhD&nbsp; dissertation&nbsp; is&nbsp; the&nbsp; assessment&nbsp; of&nbsp; the&nbsp; potentially biodegradable&nbsp; fraction&nbsp; of&nbsp; selected&nbsp; organic&nbsp; pollutants&nbsp;&nbsp; (pentachlorbenzene,&nbsp; hexachlorobenzene, lindane,&nbsp; trifluraline,&nbsp; 4-octylphenol&nbsp; and&nbsp;&nbsp;&nbsp; 4-nonylphenol)&nbsp; in&nbsp; the&nbsp; sediment,&nbsp; estimation&nbsp; of&nbsp; the biodegradation potential of these compounds in sediment in different conditions, as well as the assessment&nbsp; of&nbsp; the&nbsp; stabilization&nbsp; potential&nbsp; sediment&nbsp; by&nbsp; the&nbsp; sediment&nbsp; amendment&nbsp; with&nbsp; of&nbsp; carbon rich sorption agents. In order to develop and optimise&nbsp; methods for the bioavailability assessment of&nbsp; the&nbsp; selected&nbsp; organic&nbsp; pollutants,&nbsp; methods&nbsp; of&nbsp; multistage&nbsp; and&nbsp; single-step&nbsp; non&nbsp; exhaustive extraction&nbsp; were&nbsp; studied&nbsp; using&nbsp; the&nbsp; following&nbsp; chemical&nbsp; agents:&nbsp; Tenax&nbsp; resin,&nbsp; XAD -4&nbsp; resin&nbsp; and&nbsp; a cyclodextrin&nbsp; solution&nbsp; (2-hydroxypropyl- &beta;-cyclodextrin,&nbsp; &beta;-cyclodextrin&nbsp; and&nbsp; methyl-&beta;-cyclodextrin).&nbsp; Results&nbsp; showed&nbsp; that&nbsp; optimal&nbsp; agent&nbsp; for&nbsp; estimating&nbsp; the&nbsp; bioavailable&nbsp; fraction&nbsp; of selected organic pollutants from the sediment is XAD-4 resins, and that the optimum extraction time using single-step extraction is about 8 h. The assessment of the biodegradation potential of selected&nbsp; priority&nbsp; organic&nbsp; pollutants&nbsp; in&nbsp; the&nbsp; sediment&nbsp; was&nbsp; examined&nbsp; in&nbsp;&nbsp; various&nbsp; aerobic&nbsp; and anaerobic&nbsp; conditions,&nbsp; with&nbsp; the&nbsp; optimization&nbsp; of&nbsp; conditions&nbsp; through&nbsp; biostimulation&nbsp; and bioaugmentation.&nbsp; It&nbsp; has&nbsp; been&nbsp; shown&nbsp; that&nbsp; all&nbsp; six&nbsp; selected&nbsp; compounds&nbsp; possess&nbsp; significant biodegradation&nbsp; potential&nbsp; in&nbsp; the&nbsp; anaerobic&nbsp; environment,&nbsp; where&nbsp; in&nbsp; the&nbsp; case&nbsp; of&nbsp; lindane&nbsp; and trifluraline there is complete removal of the bioavailable amount of the compound&nbsp;&nbsp; using all four inoculum tested. Aerobic potential under the applied conditions has been showen only for alkyl phenols,&nbsp; where&nbsp; almost&nbsp; all&nbsp; of&nbsp; the&nbsp; bioavailable&nbsp; fraction&nbsp; of&nbsp; the&nbsp; compound&nbsp; in&nbsp; the&nbsp; sediment&nbsp; was removed (78-85%). The potential of sediment remediation with the amendment of sediment withcarbon&nbsp; rich&nbsp; materials&nbsp; was&nbsp;&nbsp; examined&nbsp; from&nbsp; the&nbsp; aspect&nbsp; of&nbsp; selecting&nbsp; carbon&nbsp; sorption&nbsp; agents (activated carbon, biochar and humus); estimation the optimal material doses; the long -term and short-term effects of the addition&nbsp; of these agents on the bioavailability of organic pollutants&nbsp; in order to examine the effect of aging and the toxicity of the resulting mixtures. The results of the stabilization of pollutants in the sediment show that: (1)&nbsp; increasing the dose of all three&nbsp;&nbsp; sorption agents&nbsp; leads&nbsp; to&nbsp; an&nbsp; increase&nbsp; in&nbsp; the&nbsp; immobilization&nbsp; efficiency&nbsp; and&nbsp; reduction&nbsp; of&nbsp; the&nbsp; bioavailable fraction of the selected&nbsp; organic compounds; (2) aging of the amended sediment during the first 90 days results in a further reduction of the biodegradable fraction of all compounds, after which the&nbsp; biodegradable&nbsp; concentration&nbsp; of&nbsp; the&nbsp; selected&nbsp; compounds&nbsp; remain&nbsp; the&nbsp; same&nbsp; in&nbsp; the&nbsp; case&nbsp; of activated carbon amendment, while the bioavailable fraction of&nbsp; the compound increases with the use&nbsp; of&nbsp; biohar&nbsp; and&nbsp; humus.&nbsp; Phytotoxicity&nbsp; tests&nbsp; showed&nbsp; that&nbsp; <em>Zea&nbsp; mays&nbsp;</em> accumulated&nbsp; significantly higher amount of selected organic pollutants from unamended sediment, comparing to&nbsp; <em>Cucurbita pepo&nbsp;</em> and <em>Lactuca sativa</em>. Toxicity of activated carbon and humus amended sediment assessed by <em>Vibrio&nbsp; fischeri&nbsp;</em> luminescence&nbsp; inhibition&nbsp; test&nbsp; and&nbsp; by&nbsp; measuring&nbsp; <em>Zea&nbsp; mays&nbsp; </em>germination&nbsp; and biomass yield was significantly reduced in the amended sediment samples. Accumulation of the selected&nbsp; organic&nbsp; pollutants&nbsp; in&nbsp; the&nbsp; <em>Zea&nbsp; mays</em>&nbsp; biomass&nbsp; in&nbsp; the&nbsp; unamended&nbsp; sediment&nbsp; were&nbsp; a significantly&nbsp; higher&nbsp; than&nbsp; in&nbsp; the&nbsp; humus&nbsp; and&nbsp; activated&nbsp; carbon&nbsp; amended&nbsp; sediment.&nbsp; Both&nbsp; sorbents show&nbsp; potential&nbsp; to&nbsp; be&nbsp; used&nbsp; as&nbsp; remediation&nbsp; agents&nbsp; for&nbsp; organically&nbsp; contaminated&nbsp; sediment,&nbsp; but activated carbon exhibited the better performance.</p>

In einem für das Erzgebirge (Sachsen, Südostdeutschland) typischen Wirtschaftswald aus ca. 60-jähriger Fichte (Picea abies [L.] KARST.), in den Sand-Birken (Betula pendula Roth) eingemischt sind, sollte die Wirkung der Baumartenmischung auf den Oberbodenzustand erfasst und bewertet werden. Die Befunde stammen aus einem Bestand, der sowohl hinsichtlich der Altersstruktur, des Bodentyps als auch der Stoffeinträge und der Kalkung für das Erzgebirge repräsentativ ist. Analysiert wurden photosynthetisch aktive Strahlung, Bodenvegetation, Streumenge und -verteilung, Dynamik der Streuzersetzung, Humuskörpermorphologie sowie chemische und mikrobiologische Oberbodeneigenschaften. Dabei fand ein einzelbaumweiser Ansatz in Kopplung mit Transekten Anwendung. Es wurden eindeutige Effekte durch einzelbaumweise eingemischte Birken nachgewiesen. Mit Hilfe des Einzelbaumansatzes und davon ausgehenden Transekten wurden reine Fichtenbereiche und Mischungsbereiche untersucht. Zusätzlich wurden für die Auswertung drei Straten gebildet. Die Zuordnung dafür war die folgende: Probepunkte unter der Birkenkrone ergaben das Birken-Stratum, Probepunkte unter der Fichte das Fichten-Stratum und Probepunkte im Übergangsbereich zwischen Birke und Fichte das Fichten-Birken-Stratum. Im Bereich der Birkenkronen waren überwiegend höhere Strahlungswerte festzustellen als unter den Fichtenkronen. Gezeigt wurde aber auch, dass die höheren Strahlungswerte, die unter den Birkenkronen auftraten, auch in lichteren bzw. lückigeren Partien des reinen Fichtenbestandes zu finden waren. Die Sippenanzahl der Bodenvegetation erhöhte sich sowohl unter reiner Fichte als auch in Mischung von Fichte und Birke vor allem mit zunehmender Annäherung zum Kronenrand. Insgesamt waren aber bei der Mischung von Fichte und Birke ähnlich hohe Sippenanzahlen wie im reinen Fichtenbereich zu finden. Allerdings gab es Sippen, die nur auf einer der beiden Flächen auftraten. Es waren höhere Deckungsgrade bei der Mischung von Fichte und Birke im Vergleich zur reinen Fichte sichtbar. Die Befunde der Streusammler ließen erkennen, dass in einer Entfernung von > 16 m vom Birkenstamm kaum noch Birkenstreu auftrat. Nennenswerte Trockenflussdichten wurden nur in einem Radius von 10 m festgestellt. In diesem Bereich ist ein Effekt der Birkenstreu auf die Oberbodeneigenschaften zu erwarten. Die Gesamttrockenmasse einer relativ vitalen Birke mit rund 38 cm Brusthöhendurchmesser betrug im Untersu-chungsjahr rund 8 kg. Die Modellierung unter Hinzunahme weniger vitaler Birken reduzierte die Gesamttrockenmasse auf 6,3 kg. Der Massenverlust der Birkenblätter in den Litter-Bags war in allen drei Straten bei der Mischung der Birkenblätter mit Fichtennadeln im Verhältnis 5:1 am größten. Bei den Fichtennadeln zeigten sich die höchsten Trockenmassenverluste in den reinen Nadel-Litter-Bags. Die Litter-Bags unter der Birkenkrone wiesen für alle Varianten die beste Zersetzung der Birkenblätter und der Fichtennadeln auf. Die C/N-Verhältnisse der Streu ließen keine eindeutigen positiven Effekte der Birkenkrone erkennen. Sie waren aber bei den Birkenblättern in allen Varianten, in allen Straten und zu allen Terminen deutlich enger als die der Fichtennadeln. Desweiteren war der Abfall der C/N-Kurven bei den Birkenblättern steiler als bei den Fichtennadeln. Die Mächtigkeit des Of-Horizontes wies sowohl in der Mischsituation als auch in der reinen Fichtenfläche deutlichere räumliche Muster auf als im Oh-Horizont. Im Of waren dadurch deutliche, vom Einzelbaum abhängige, Muster zu erkennen. Die höchsten Mächtigkeiten traten in Stammnähe auf. Die größten Gesamtmächtigkeiten existierten unter reiner Fichte. Beim Vergleich der Mächtigkeiten in den Straten war in der Of-Lage vor allem das Birken-Stratum signifikant verschieden vom Fichten-Stratum. Für die Oh-Mächtigkeit und somit auch die Gesamtmächtigkeit zeigte zusätzlich das Fichten-Birken-Stratum signifikante Unterschiede im Vergleich zum Birken-Stratum. Als Humusform trat der rohhumusartige Moder auf. Dabei überwog im Fichtenstratum der feinhumusreiche rohhumusartige Moder, im Fichten-Birken-Stratum und im Birken-Stratum dagegen der feinhumusarme rohhumusartige Moder. Signifikante Unterschiede zwischen Straten konnten in der Of-Lage für den pH-Wert, Cges-Vorrat, Ccarbonat-Gehalt, Nges-Gehalt, Nges-Vorrat, und den metabolischen Quotienten erfasst werden. In der Oh-Lage waren es die nachstehenden Oberbo-deneigenschaften: Corg/N-Verhältnis, Cges-Gehalt, Nmik-Gehalt, Cmik/Nmik-Verhältnis, Basalatmungsaktivität, sowie der metabolische Quotient, die sich zwischen den Straten signifikant unterschieden. Im A-Horizont zeigten sich signifikante Unterschiede der Straten beim Corg/N-Verhältnis, Cges-Gehalt, Ccabonat-Gehalt, Corg-Gehalt sowie dem Nges-Vorrat. Mit Hilfe der multivariaten Ordination (mit verschiedenen Elementgehalten und pH-Werten) ist die Stratenbildung erneut bestätigt worden. Dabei hatten die Elemente C, Mg, Ca und N den größten Einfluss. Die Auswertung der Daten der effektiven Kationenaustauschkapazität ließ vor allem für den A-Horizont Unterschiede zwischen den Straten erkennen. Dabei war die Akeff im Fichtenstratum am kleinsten und der prozentuale Anteil der sauren Kationen an der Akeff am größten. Die höchste Akeff besaß das Birken-Stratum. Eine Reihe der untersuchten Größen ließen eindeutige Baumarteneffekte erkennen. Alles in allem wurde der Nachweis erbracht, dass die Effekte in dem Bereich zu finden sind, der durch die Laubstreu der Birken geprägt ist. Bei den Oberbodeneigenschaften wurden keine Veränderungen an Punkten, die mehr als 10 m vom Birkenstamm entfernt waren, festgestellt. Aus den Ergebnissen war abzuleiten, dass für ähnliche Bedingungen wie im Untersuchungsbestand ein Mischungsanteil der Birke von mindestens 10 % zu empfehlen ist. Die Ergebnisse sind aber durch zukünftige Forschung noch zu verifizieren. Dafür sollten vor allem die Produktionsziele und die waldbaulichen Behandlungskonzepte für Mischbestände aus Fichte und Birke in Deutschland erneut konkretisiert werden. Um dafür eindeutige ökologische und ökonomische Kriterien festlegen zu können, wird eine Zusammenführung wichtiger Ergebnisse aus den verschiedenen forstlichen Forschungsbereichen (z. B. Standortskunde, Vegetationskunde, Waldwachstumskunde, Ökonomie, Zoologie, Waldbau) empfohlen. / In a typical managed forest of the Ore Mountains (Saxony, SE Germany), consisting of 60 years old Norway spruce (Picea abies [L.] KARST.) admixed with silver birches (Betula pendula Roth) the impact of the admixture on topsoil properties was quantified and evaluated. This study was conducted in a stand which is representative for the Ore Mountains regarding the soil type, deposition and liming. Photosynthetic active radiation, the layer of herbs and mosses, the mass and distribution of the litter fall, the dynamics of litter decom-position, and morphology of the humus layer as well as the chemical and microbiological topsoil properties were all analyzed. The study was based on a single-tree-approach where samples were collected along transects from selected sample trees of birch and spruce. Transects were established within areas of pure spruce as well as within areas of spruce and birch mixture. Three different strata were defined based on the situation of the examined points in relation to the crowns of birch and spruce: the birch stratum, the spruce stratum and the spruce-birch stratum with measurements from underneath the crowns of birches, spruces and the transition area between spruce and birch crowns, respectively. Higher photosynthetic active radiation values were detected in the area underneath the crowns of birch crowns in comparison with the area underneath the crowns of spruce. Higher values of photosynthetic active radiation were also registered within areas of light foliage and in gaps in the pure spruce stand. The number of ground vegetation species underneath pure spruce and in the mixture of spruce and birch was negatively correlated with the distance to the crown edge. But altogether the same number of ground vegetation species was found in the mixed stand and in the pure spruce stand. A few species were, however, specific for each stand type. A higher coverage of ground vegetation was found in the mixture of spruce and birch in comparison to the pure spruce. The results of the litter traps show marginal densities of birch leaves at a distance of more than 16 m from the birch stem. The majority of the dry flux density was found within a radius of 10 m around the birch stem. Within this area an effect of the birch litter on top soil properties was expected. The whole dry litter mass was measured in the study time of a relative vital birch with a diameter at breast high of ca. 38 cm and was ca. 8 kg. Modeling the amount of litter for birches with lower vitality indicated a dry litter mass of 6.3 kg. The mass lost of the birch leaves in the litter bags was in each stratum the highest at a ratio of 5 to 1 of birch leaves to spruce needles. The greatest loss of dry mass for spruce needles was registered in litter-bags with pure needles. The litter-bags under the birch crowns exhibited the best decomposition rate of birch leaves and spruce needles for all variants. The C/N-ratios of the litter showed no clear positive effects of the birch crown. But they were clearly smaller for the birch leaves in contrast to the spruce needles in all variants, in each stratum and at all dates. Furthermore the C/N-curves drop away more for the birch leaves in contrast to the spruce needles. The thickness of the Of-horizon in contrast to the Oh-horizon showed clear spatial patterns for the mixed situation and the pure spruce area. Thus in the Of-horizon a clear spatial pattern in connection to the single tree was apparent. The highest thickness was found close to the stem. The highest total thickness of the Of- and Oh-horizon was found underneath the pure spruce stand. When comparing the thickness of the humus layers between the stratums, significant differences where found for the Of-layer between the birch stratum and the spruce stratum. In addition, for the thickness of the Oh-layer as well as for the total thickness of all layers, significant differences were observed between the spruce-birch stratum and the birch stratum. The humus form was a mor-like moder. In the spruce stratum more fine-humus-rich variants of the mor-like moder was found, in the spruce-birch stratum and in the birch stratum it was more fine-humus-poor variants of the mor-like moder. Significant differences between the stratums in the Of-layer were found with respect to acidification, the pool of Ctot and Ntot, the content of Ccarbonat and Ntot and the metabolic quotient. For the Oh-layer significant differences between the stratums were found for the ratio of Corg and N, the content of Ctot and Nmic, the ratio of Cmic and Nmic, the basal respiration activity and the metabolic quotient. In the A-horizon significant differences between the stratums were found for the ration of Corg and N, the content of Ctot, Ccarbonat and Corg and the pool of Ntot. The results of multivariate ordination confirm the formation of stratums again. Thereby the contents of C, Mg, Ca and N had the greatest influence. The data analysis of the effective cation exchange capacity (ECEC) showed differences between the stratums mainly for the A horizon. In the spruce stratum the ECEC was lowest and the percentage of acid cations at the ECEC was greatest. The highest ECEC was in the birch stratum. Different results showed clear tree effects. These were only found in areas, which are influenced by birch leaves. In this study, a minimum tree admixture of 10% was required before an impact on the ground conditions of the whole stand was observed. Therefore, based on the results of this study, an admixture of birch of 10% is recommended. These results however must be verified through future research. These findings can be used to assist in meeting the goals of silvicultural management for mixed spruce and birch forests in Germany. Definition of explicit ecological and economic criteria created from a combination of results from different forest research sectors (e. g. Site ecology, Vegetation science, Forest Growth, Economy, Zoology, Silviculture) is recommended.

Das Forschungsvorhaben hat das Ziel den Einfluss einzelbaumweise eingemischter einheimischer Eichen in Wäldern der Gemeinen Kiefer (Pinus sylvestris L.) zu untersuchen. Die Untersuchungsflächen befinden sich im Nordosten Deutschlands (Brandenburg). Im Untersuchungsgebiet Jerischke hat die Kiefer ein Alter von 78 Jahren und die Eiche ein Alter von 205 Jahren. Im Untersuchungsgebiet Weidmannsruh befindet sich die Kiefer im Alter 70 und die Eiche im Alter 108. In Abhängigkeit von der Entfernung zur Eiche werden folgende Effekte der Eiche nach Intensität und Qualität untersucht: - Effekte auf Mikroklima und Strahlung, - Effekte auf Humusmorphologie und bodenchemische Parameter und - Effekte auf Bodenvegetation und epigäische Fauna. Grundsätzlich sind die positiven Effekte der Eiche auf 10 bis 15 m vom Eichenstamm in den Kiefernbestand hinein begrenzt.

About 100 years ago, groups of European beech were reintroduced into Norway spruce stands in some parts of Germany as a restoration approach. The aim of this approach was to maintain or rehabilitate the ecological function of soil fertility and the regeneration option of beech for the next generation. The effect of beech groups on surrounding spruce stands was studied in the Ore Mountains by recording natural regeneration of beech and humus layer thickness and form. Point pattern statistics were used to analyse the spread of beech regeneration and to determine factors influencing its establishment. It was found that the density of regeneration decreases with increasing distance from the beech group. However, beech regeneration was found up to distances of 69 m. Furthermore, it becomes evident that PAR radiation (maximum regeneration densities at PAR values of 35 W/m²) and fencing (3.41 times higher regeneration density compared to unfenced areas) against deer have a positive influence on beech regeneration density. Ordered categorical models were used to model humus form and non-linear models were used to model humus layer thickness. It could be proven that the most bioactive humus forms and lowest humus layer thicknesses were found within the beech group. With increasing distance to the beech group, the total humus layer thickness and the proportion of mormoder in the spruce stand increased. The positive influence of the beech group on the humus composition extends to about 40 m from the centre of the beech group. Due to the former arrangement of the beech groups in the terrain, the effects can also extend to the spruce stands in between. The hypotheses on the restoration approach of reintroducing groups of beech into spruce stands formulated by foresters 100 years ago can thus be confirmed. For future restoration approaches of spruce stands, groupwise mixtures of beech should be established with a distance of 40–50 m.

En dépit de son importance pour les écosystèmes aquatiques, l’origine et les mécanismes de production du carbone organique dissous (COD) sont toujours sujets à discussion. Cette thèse vise à mieux comprendre le rôle joué par l’hydrologie tant sur les mécanismes de production et la location des sources du COD que sur les flux et les processus de transfert du sol vers les cours d’eau. Un suivi haute fréquence des eaux du sol et de rivière a été réalisé sur la totalité d’un cycle hydrologique dans le bassin versant expérimental de Kervidy-Naizin (Morbihan, Observatoire de Recherche en Environnement (ORE) AgrHys). Le COD contenu dans ces eaux a été caractérisé par sa composition isotopique en carbone (δ13C), et ce suivi a été complété par l’analyse de la dynamique de la nappe. L’analyse des variations saisonnières de concentration et de composition du COD révèle l’existence d’un relais dans les sources et les mécanismes de production du COD dans les sols des zones de fond de vallée en phase avec les changements de régime hydrologique du bassin. Ainsi, lors de la période automnale de remontée de la nappe, le compartiment COD présente un caractère faiblement aromatique et une composition isotopique particulière, suggérant une origine microbienne. Ce réservoir de très faible taille (5% du flux annuel de COD exporté par le cours d’eau) est entièrement épuisé par les premières crues d’automne. La mise en charge de la nappe en versant conduit à la mobilisation d’un second réservoir de COD caractérisé par une aromaticité élevée et une signature isotopique proche de celle de la matière organique des sols. Ce compartiment correspond au fond humique « ancien » des sols et présente une taille nettement plus important que le premier (90% du flux annuel). Cependant ce compartiment ne se limite pas au COD produit dans sols des zones de bas-fond mais comprend aussi des apports de matières organiques issues des sols de bas de versant. Enfin, le rabattement de la nappe au milieu du printemps marque le début de la période d’assèchement du bassin versant et le retour dans les sols de bas-fond d’un COD faiblement aromatique et isotopiquement similaire à celui observé en automne. En conséquence, la teneur et la composition du COD véhiculé par le cours d’eau varient fortement à l’échelle saisonnière (variabilité temporelle des processus de production du COD). Les horizons superficiels des sols des zones de bas-fond sont la principale source de COD, par lesquels transitent entre 60 et 80% du flux de COD exporté lors des événements de crues. Concernant le réservoir humique, un résultat important est qu’il semble limité dans les sols de bas de versant, alors qu’il semble au contraire illimité dans les sols de bas fond. Au final, les résultats obtenus dans le cadre de cette thèse démontrent le contrôle majeur de la dynamique de la nappe sur les mécanismes de production de COD par le sol ainsi que sur la localisation spatiale et les flux de COD transférés de ces sols vers les cours d’eau à l’échelle de la saison. Ils valident également l’hypothèse émise selon laquelle l’exportation du COD par les cours d’eau ne résulterait pas d’un simple « flushing » du COD produit dans les sols de bas-fond mais mobiliserait plutôt un ensemble de sources localisées le long du continuum rivière-zone de bas-fond-zone de versant, et dont le caractère plus moins limité du réservoir expliquerait les dynamiques des concentrations en COD annuelles observées à l’exutoire de ces bassins. Un autre résultat majeur de cette thèse est la qualification des isotopes comme outil de traçage des sources et de la dynamique du COD dans les bassins versants. Un point clé de ce travail est la différenciation isotopiques des réservoirs COD des sols de bas-fond et des sols de bas de versant, qui a permis pour la première fois de révéler la participation de l'un et de l’autre au flux exporté à l’exutoire et d’estimer quantitativement leur contribution respective à ce même flux. / Despite its importance to aquatic ecosystems, origin and production mechanisms of dissolved organic carbon (DOC) are still subject to discussion. This thesis aims to better understand the role of hydrology in controlling both DOC production in soils and export by stream waters. High frequency monitoring of soils and streams waters was carried out during an entire hydrological cycle in the Kervidy-Naizin catchment (Morbihan). The DOC was characterized by its stable carbon isotopic composition, and groundwater table dynamic was also analyzed during the same period. The analysis of seasonal variations in both DOC concentration and composition reveals a succession in the sources and mechanisms of production of DOC in soils of valley bottom in line with changes in the hydrological regime of the basin. Thus, during the autumn period of rising water-table, the COD compartment has a low aromatic character and a particular isotopic composition, suggesting a microbial origin. This reservoir of very small size (5% of the annual flux of DOC exported by the stream) is fully exhausted by the first autumn storm events. The increase of water-table in the upland domain leads to the mobilization of a second DOC pool characterized by a high aromaticity and a isotopic signature similar to those of soil organic matter. This compartment corresponds to "old" humic fraction of the soil and has a size much larger than the first (90% of annual flux). However, this compartment includes contributions of DOC from riparian soils and also upland soils. Finally, the water-table drawdown in the middle of spring marks the beginning of the drying period of the catchment and the return to the soil of the “fresh” DOC pool. As a result, concentration and composition of DOC transported by the stream vary greatly across a seasonal scale (temporal variability of production processes DOC). The surface horizons of soils riparian soils are the main source of DOC, which handle between 60 and 80% of stream DOC flux during flood events. An important result is that the DOC pool seems limited in upland soils, while its seems no-limited in riparian soils. Finally, the results obtained in this thesis demonstrate the major control of the dynamics of the water-table on the mechanisms of production of DOC from the soil as well as the spatial location of DOC sources and DOC flux transferred from the soil to stream through the hydrological cycle. They also validate the hypothesis that DOC export by rivers mobilizes rather a set of sources located along the upland-riparian-stream continuum. Another major result of this thesis is the validation of stable carbon isotopes as a tool for tracing sources and dynamics of DOC in the catchment.

Orientador: Enner Herênio Alcântara / Banca: Thanan Walesza Pqueno Rodrigues Guimarães / Banca: Nilton Nobuhiro Imai / Resumo: A matéria orgânica dissolvida (DOM) é a componente da água que pode ser usada como indicativo de sua qualidade, pois possui duas fontes: uma alóctone, relacionada com descargas de material terrestre, estando vinculada aos ácidos húmicos, e outra autóctone, associada às descargas fluviais ou produção própria do corpo hídrico estudado, estando relacionada aos ácidos fúlvicos. A matéria orgânica dissolvida colorida (CDOM) é a fração colorida da DOM, que pode ser usada como proxy para a observação desta última em águas interiores. O reservatório hidrelétrico de Funil (FHR) foi o corpo hídrico escolhido como área de estudo deste trabalho. Neste contexto, o objetivo geral desta pesquisa foi identificar e avaliar as mudanças no coeficiente de absorção da CDOM (aCDOM) na superfície da água ao longo do tempo (1995 - 2010), bem como entender a sua relação com mudanças no uso e cobertura da terra (LULC) na bacia de contribuição do FHR. Para alcançar tal objetivo foram realizados: (i) o mapeamento histórico de LULC (1995 - 2010, com 5 anos de intervalo) para detecção de mudanças; (ii) o estudo de um conjunto de modelos bio-ópticos baseados na literatura, bem como de um novo modelo empírico desenvolvido para estimar aCDOM via reflectância simulada (Rrs_simulated) para o sensor Thematic Mapper (TM); (iii) a distribuição espaço-temporal do aCDOM por meio da aplicação de um modelo bio-óptico em imagens TM/Landsat-5 de 1995 a 2010, e (iv) a análise das fontes possíveis de CDOM/DOM, assim com... (Resumo completo, clicar acesso eletrônico abaixo) / Abstract: The dissolved organic matter (DOM) is a water compound related to water quality, since it has two sources: one allochthonous, related to terrestrial discharges that can be linked to humic acids, and another autochthonous, associated with river input and itself production, so related to or fulvic acids. The colored dissolved organic carbon (CDOM) is the colored fraction of DOM that could be used as a proxy for its occurrence in inland waters. The Funil hydroelectric reservoir (FHR) was chosen as the study site for this work. In this context, the general aim of this research was to identify and to evaluate the changes in CDOM absorption coefficient (aCDOM) at the water surface over time (1995 - 2010), and to understand its relationship with land cover land use (LULC) changes in FHR watershed. For match this goal, (i) a LULC historical mapping (1995 - 2010, with 5 years of interval) was made to change detection, (ii) a bio-optical model set and a new model were studied in order to estimate aCDOM from simulated reflectance (Rrs_simulated) for Thematic Mapper (TM) sensor, (iii) a aCDOM spatial and temporal distribution was obtained by applying a bio-optical model in TM/Landsat-5 imagery from 1995 to 2010, and (iv) the possible CDOM/DOM sources in FHR were found, as well aCDOM historical behavior/distribution over time was analysed. Thus, the first study was the support vector machine algorithm (SVM) parameterization according to study area characteristics to LULC supervised classi... (Complete abstract click electronic access below) / Mestre