Coloradia pandora (Blake) is a phytophagous insect that defoliates Pinus ponderosa (Dougl. ex Laws.) in south-central Oregon. Little is known about the extent of damage this insect inflicts upon its host trees during an outbreak. In this paper, we present stem analyses on four dominant Pinus ponderosa trees that enable us to determine the amount of volume lost during each Coloradia pandora outbreak on this site for the past 450 years. We found that on average an outbreak inhibits radial growth so that an individual tree produces 0.057 m³ less wood volume than the potential growth for the duration of an individual outbreak. A total of 0.549 m³ of growth per tree was inhibited by 10 outbreaks during the lifetime of the trees, which, in this stand, equates to 9.912 m³/ha (1,700 board feet/acre) of wood suppressed over the last 450 years throughout the stand. Our results do not support previous findings of a lag in suppression onset between the canopy of the tree versus the base. Crossdating of stem analysis samples is paramount to definitively examine the potential for a lagged response throughout the

Disturbances are ubiquitous features of most northern forest ecosystems. The subsequent response of plant assemblages on both short (resistance or not) and long term (recolonization or not) will depend on a number of factors operating at several spatial scales. In boreal forest ecosystems, bryophyte assemblages are a conspicuous and species rich group of plants for which these processes are poorly understood. Using a combination of experimental and observational approaches this thesis explores these questions for closed-canopy bryophyte assemblages in relation to a) microtopography (both for the initial and long-term response), b) environmental constrains during post-logging succession and c) disturbance type. My results clearly show that the shade and shelter provided by microtopographic surface structures can increase survival rates of bryophytes following clear-cut logging by decreasing mortality from microclimatic stress and mechanical disturbance. Following clear-cutting, the recovery of forest floor and dead wood living bryophytes seems to be a relatively steady and progressive process without any major bottleneck episodes in the young or semi-mature forest stages with much of the pre-disturbance composition recovered after 50 years. Although boulders were found to increase the initial survival on clear-cuts and hence increase disturbance resistance, we found no evidence that boulders influenced the subsequent recolonization process. Lastly, strong compositional dissimilarities were found in young forests (40 years) following clear-cut logging, wildfire and insect outbreak, indicating divergent trajectories to occur following different disturbances. Hence, early seral stages of forest ecosystems regenerating after natural disturbances seem to compliment young managed forests in maintaining landscape level diversity. / At the time of the doctoral defense, the following papers were unpublished and had a status as follows: Paper 2: Submitted. Paper 3: Manuscript. Paper 4: Manuscript.

Climate-triggered forest disturbances are increasing either by drought or by other climate extremes. Droughts can change the structure and function of forests in long-term or cause large-scale disturbances such as tree mortality, forest fires and insect outbreaks in short-term. Traditional approaches such as dendroclimatological surveys could retrieve the long-term responses of forest trees to drought conditions; however, they are restricted to individual trees or local forest stands. Therefore, multitemporal satellite-based approaches are progressing for holistic assessment of climate-induced forest responses from regional to global scales. However, little information exists on the efficiency of satellite data for analyzing the effects of droughts in different forest biomes and further studies on the analysis of approaches and large-scale disturbances of droughts are required. This research was accomplished for assessing satellite-derived physiological responses of the Caspian Hyrcanian broadleaves forests to climate-triggered droughts from regional to large scales in northeast Iran. The 16-day physiological anomalies of rangelands and forests were analysed using MODIS-derived indices concerning water content deficit and greenness loss, and their variations were spatially assessed with monthly and inter-seasonal precipitation anomalies from 2000 to 2016. Specifically, dimensions of forest droughts were evaluated in relations with the dimensions of meteorological and hydrological droughts. Large-scale effects of droughts were explored in terms of tree mortality, insect outbreaks, and forest fires using field observations, multitemporal Landsat and TerraClimate data. Various approaches were evaluated to explore forest responses to climate hazards such as traditional regression models, spatial autocorrelations, spatial regression models, and panel data models. Key findings revealed that rangelands’ anomalies did show positive responses to monthly and inter-seasonal precipitation anomalies. However, forests’ droughts were highly associated with increases in temperatures and evapotranspiration and were slightly associated with the decreases in precipitation and surface water level. The hazard intensity of droughts has affected the water content of forests higher than their greenness properties. The stages of moderate to extreme dieback of trees were significantly associated with the hazard intensity of the deficit of forests’ water content. However, the stage of severe defoliation was only associated with the hazard intensity of forests’ greenness loss. Climate hazards significantly triggered insect outbreaks and forest fires. Although maximum temperatures, precipitation deficit, availability of soil moisture and forest fires of the previous year could significantly trigger insect outbreaks, the maximum temperatures were the only significant triggers of forest fires from 2010‒2017. In addition to climate factors, environmental and anthropogenic factors could control fire severity during a dry season. The overall evaluation indicated the evidence of spatial associations between satellite-derived forest disturbances and climate hazards. Future studies are required to apply the approaches that could handle big-data, use the satellite data that have finer wavelengths for large-scale mapping of forest disturbances, and discriminate climate-induced forest disturbances from those that induced by other biotic and abiotic agents. / Klimagbedingte Waldstörungen nehmen entweder durch Dürre oder durch andere Klimaextreme zu. Dürren können langfristig die Struktur und Funktion der Wälder verändern oder kurzfristig große Störungen wie Baumsterben, Waldbrände und Insektenausbrüche verursachen. Traditionelle Ansätze wie dendroklimatologische Untersuchungen könnten die langfristigen Reaktionen von Waldbäumen auf Dürrebedingungen aufzeigen, sie sind aber auf einzelne Bäume oder lokale Waldbestände beschränkt. Daher werden multitemporale satellitengestützte Ansätze zur ganzheitlichen Bewertung von klimabedingten Waldreaktionen auf regionaler bis globaler Ebene weiterentwickelt. Es gibt jedoch nur wenige Informationen über die Effizienz von Satellitendaten zur Analyse der Auswirkungen von Dürren in verschiedenen Waldbiotopen. Daher sind weitere Studien zur Analyse von Ansätzen und großräumigen Störungen von Dürren erforderlich. Diese Forschung wurde durchgeführt, um die aus Satellitendaten gewonnenen physiologischen Reaktionen der im Nordosten Irans gelegenen kaspischen hyrkanischen Laubwälder auf klimabedingte Dürren auf lokaler und regionaler Ebene zu bewerten. Auf der Grundlage der aus MODIS-Daten abgeleiteten Indizes wurden die 16-tägigen physiologischen Anomalien von Weideland und Wäldern in Bezug auf Wassergehaltsdefizit und Grünverlust analysiert und ihre Variationen räumlich mit monatlichen und intersaisonalen Niederschlagsanomalien von 2000 bis 2016 bewertet. Insbesondere wurden die Dimensionen der Walddürre in Verbindung mit den Dimensionen der meteorologischen und hydrologischen Dürre bewertet. Großräumige Auswirkungen von Dürren wurden in Bezug auf Baumsterblichkeit, Insektenausbrüche und Waldbrände mit Hilfe von Feldbeobachtungen, multitemporalen Landsat- und TerraClimate Daten untersucht. Verschiedene Ansätze wurden ausgewertet, um Waldreaktionen auf Klimagefahren wie traditionelle Regressionsmodelle, räumliche Autokorrelationen, räumliche Regressionsmodelle und Paneldatenmodelle zu untersuchen. Die wichtigsten Ergebnisse zeigten, dass die Anomalien von Weideland positive Reaktionen auf monatliche und intersaisonale Niederschlagsanomalien aufweisen. Die Dürren in den Wäldern waren jedoch in hohem Maße mit Temperaturerhöhungen und Evapotranspiration verbunden und standen in geringem Zusammenhang mit dem Rückgang von Niederschlägen und des Oberflächenwasserspiegels. Die Gefährdungsintensität von Dürren hat den Wassergehalt von Wäldern stärker beeinflusst als die Eigenschaften ihres Blattgrüns. Die Stufen mittlerer bis extremer Baumsterblichkeit waren signifikant mit der Gefährdungsintensität des Defizits des Wassergehalts der Wälder verbunden. Das Ausmaß der starken Entlaubung hing jedoch nur mit der Gefährdungsintensität des Grünverlustes der Wälder zusammen. Die Klimagefahren haben zu deutlichen Insektenausbrüchen und Waldbränden geführt. Obwohl Maximaltemperaturen, Niederschlagsdefizite, fehlende Bodenfeuchte und Waldbrände des Vorjahres deutlich Insektenausbrüche auslösen konnten, waren die Maximaltemperaturen die einzigen signifikanten Auslöser von Waldbränden von 2010 bis 2017. Neben den Klimafaktoren können auch umweltbedingte und anthropogene Faktoren den Schweregrad eines Brandes während einer Trockenzeit beeinflussen. Die Gesamtbewertung zeigt Hinweise auf räumliche Zusammenhänge zwischen aus Satellitendaten abgeleiteten Waldstörungen und Klimagefahren. Weitere Untersuchungen sind erforderlich, um Ansätze anzuwenden, die mit großen Datenmengen umgehen können, die Satellitendaten in einer hohen spektralen Auflösung für die großmaßstäbige Kartierung von Waldstörungen verwenden und die klimabedingte Waldstörungen von denen zu unterscheiden, die durch andere biotische und abiotische Faktoren verursacht werden.