Global ETD Search

1	Estudo morfométrico em foraminíferos planctônicos da margem continental brasileira / Morphometric study in planktonic foraminifera on Brazilian continental margin Iwai, Fabiane Sayuri 06 August 2015 (has links) A taxonomia de foraminíferos planctônicos é fonte de discussões desde o início de sua utilização na paleoceanografia, havendo discordância tanto na sua classificação como identificação. Estudos genéticos em foraminíferos planctônicos identificaram a presença de mais de uma espécie dentro das espécies aceitas atualmente. Estas espécies possuem diferenças ecológicas entre si, tendo implicações para interpretações paleoceanográficas. A morfometria e isótopos de carbono e oxigênio foram escolhidos como uma alternativa mais acurada e reproduzível para identificação de variações morfológicas e para explorar a ecologia dos indivíduos estudados, respectivamente. Foram utilizadas amostras de sedimento da margem continental brasileira para explorar o potencial da morfometria como ferramenta paleoceanográfica. Foi possível observar a dominância do genótipo pink de Globigerinoides ruber na margem continental brasileira. Para Globigerinoides sacculifer, foi possível identificar comportamentos de migração vertical distintos entre os morfotipos identificados. Em Globorotalia menardii observa-se que as relações morfométricas-ambientais para o grupo todo se mantém quando a análise é feita separadamente em cada morfotipo, confirmando o potencial da correlação direta da morfometria com variáveis ambientais. Já em Globigerinella siphonifera, a diferença morfológica é atribuída a presença de diferentes espécies e não a influências do ambiente sobre a morfologia. / Although taxonomy in planktonic foraminifera has been subject of debates since the beginning of its use in paleoceanography, disagreement in their classification and identification remains. Genetic investigations have identified the presence of more than one species for some of the species accepted nowadays. Hence, these species display ecological differences among them, resulting in implications for paleoceanographic interpretations. Morphometry and carbon and oxygen stable isotopes were chosen as a more precise and reproducible alternative to identify morphological variations and to explore the ecology of the specimens, respectively. Sediment samples from the Brazilian Continental Margin were used to explore morphometry\'s potential as a paleoceanography tool. It was possible to observe the dominance of the pink genotype in Globigerinoides ruber at Brazilian continental margin. For Globigerinoides sacculifer, it was possible to identify distinct vertical migration behaviour in each identified morphotype. In Globorotalia menardii it is possible to observe that the morphometric-enviromental relations for the whole group is maintained when each morphotype is analysed separately, confirming the potential of direct correlations between morphometry and environmental variables. Meanwhile, in Globigerinella siphonifera the morphological difference is attributed to the presence of two different species and not to the influence of the environment on the morphology. Cryptic species Espécies crípticas Isótopos estáveis Morfoespécies Morfologia Morfometria Morphology Morphometry Morphospecies Paleoceanografia Paleoceanography Stable isotopes
2	Estudo morfométrico em foraminíferos planctônicos da margem continental brasileira / Morphometric study in planktonic foraminifera on Brazilian continental margin Fabiane Sayuri Iwai 06 August 2015 (has links) A taxonomia de foraminíferos planctônicos é fonte de discussões desde o início de sua utilização na paleoceanografia, havendo discordância tanto na sua classificação como identificação. Estudos genéticos em foraminíferos planctônicos identificaram a presença de mais de uma espécie dentro das espécies aceitas atualmente. Estas espécies possuem diferenças ecológicas entre si, tendo implicações para interpretações paleoceanográficas. A morfometria e isótopos de carbono e oxigênio foram escolhidos como uma alternativa mais acurada e reproduzível para identificação de variações morfológicas e para explorar a ecologia dos indivíduos estudados, respectivamente. Foram utilizadas amostras de sedimento da margem continental brasileira para explorar o potencial da morfometria como ferramenta paleoceanográfica. Foi possível observar a dominância do genótipo pink de Globigerinoides ruber na margem continental brasileira. Para Globigerinoides sacculifer, foi possível identificar comportamentos de migração vertical distintos entre os morfotipos identificados. Em Globorotalia menardii observa-se que as relações morfométricas-ambientais para o grupo todo se mantém quando a análise é feita separadamente em cada morfotipo, confirmando o potencial da correlação direta da morfometria com variáveis ambientais. Já em Globigerinella siphonifera, a diferença morfológica é atribuída a presença de diferentes espécies e não a influências do ambiente sobre a morfologia. / Although taxonomy in planktonic foraminifera has been subject of debates since the beginning of its use in paleoceanography, disagreement in their classification and identification remains. Genetic investigations have identified the presence of more than one species for some of the species accepted nowadays. Hence, these species display ecological differences among them, resulting in implications for paleoceanographic interpretations. Morphometry and carbon and oxygen stable isotopes were chosen as a more precise and reproducible alternative to identify morphological variations and to explore the ecology of the specimens, respectively. Sediment samples from the Brazilian Continental Margin were used to explore morphometry\'s potential as a paleoceanography tool. It was possible to observe the dominance of the pink genotype in Globigerinoides ruber at Brazilian continental margin. For Globigerinoides sacculifer, it was possible to identify distinct vertical migration behaviour in each identified morphotype. In Globorotalia menardii it is possible to observe that the morphometric-enviromental relations for the whole group is maintained when each morphotype is analysed separately, confirming the potential of direct correlations between morphometry and environmental variables. Meanwhile, in Globigerinella siphonifera the morphological difference is attributed to the presence of two different species and not to the influence of the environment on the morphology. Espécies crípticas Isótopos estáveis Morfoespécies Morfologia Morfometria Paleoceanografia Cryptic species Morphology Morphometry Morphospecies Paleoceanography Stable isotopes
3	Plant- and arthropod diversity of vegetable gardens along a socio-economic gradient within the Tlokwe Municipal Area / Nicola Botha Botha, Nicola January 2012 (has links) Globally urbanization has increased to such an extent that more than half of the human population currently resides in cities. In the years to come, urban expansion will especially take place in developing countries through efforts to improve economic growth and poverty alleviation. This may have a negative effect on native biodiversity within and surrounding urban environments. However, residential areas with a high proportion of gardens form a significantly large part of urban environments and these domestic gardens contribute to the maintenance and preservation of biodiversity in cities. Although the preservation of biodiversity in these gardens is important in the overall conservation of urban green spaces, little is known about how these gardens can possibly contribute to conservation purposes in urban areas. Bearing in mind that anthropogenic activities are possible drivers of urban biodiversity, it is vital to quantify socio-economic aspects within urban ecological research. In developing countries, such as South Africa, the inclusion of socio-economic aspects are especially important because there is a wider gap between poor and wealthy households. There are also a larger number of people that are dependent on their gardens for subsistence purposes, such as vegetable gardening. In the Municipal Area of Tlokwe, South Africa, there exists a definite socio-economic gradient from the poorer western to the more affluent eastern part of the city. Five socio-economic status (SES) classes, primarily based on % unemployment, were used in this study. The ultimate aim of this study was therefore to determine the plant- and arthropod diversity within urban domestic gardens along a socio-economic gradient. Vegetable gardens within domestic gardens were selected to quantify plant- and arthropod biodiversity. Biodiversity of adjacent lawns were also sampled for comparative purposes. The study also attempted to determine to what extent socio-economic aspects of city residents may be possible drivers of biodiversity within the gardens. Various other factors that might have an effect on the plant and/or arthropod diversity were included such as soil characteristics, specific management factors of the gardens and other land-uses surrounding domestic gardens. Arthropod diversity was surveyd by means of pitfall traps and suction sampling in eight 0.25 m2 squares along an 8 m transect in each representative garden. Arthropods were identified up to morphospecies level. Vegetation was surveyed along the same transect and total species composition was determined. Plants were identified up to species level. The plant and arthropod surveys were conducted in both the vegetable gardens and lawns of all SES classes. For the soil samples a 1:2.5 water analysis was conducted. A social survey was conducted in all representative gardens by means of a questionnaire and a SPOT 5 satellite imagery was used to determine the land-use types in the areas surrounding the participating gardens. All the above mentioned factors were compared between the different SES classes. Diversity indices for the arthropods, multivariate statistical analyses and ANOVA analyses were applied to test for meaningful variables between socio-economic status classes as well as vegetable gardens and lawns. From the results it was evident that the more affluent SES classes had significantly higher arthropod diversity values, whilst the lower income classes had higher plant diversity. The factor analysis between the plants and arthropods with the surrounding land-uses revealed two significant factors. Firstly, arthropod diversity was influenced by domestic gardens in the surrounding landscape and there was a positive correlation between these two variables. This indicates that a high percentage of surrounding domestic gardens were possible drivers of arthropod diversity. No correlations were evident between plant and arthropod diversity. Secondly, the other significant factor showed that one SES class had a significantly higher percentage of woodlands and grasslands as opposed to two of the other classes that had a significantly higher percentage of built structures within the surrounding area. Differences were also apparent between the SES classes concerning management regimes, financial stability and level of education. The two more affluent SES classes had obtained a higher level of education and income and had management practices that were uncommon in the three poorer SES classes. This study proposes that domestic gardens are a means to conserve biodiversity in cities. Vegetable gardens in domestic gardens will also be able to harbour a larger diversity of plants and arthropods than the lawns. The socio-economic status of residents also had a significant effect on biodiversity and therefore it should be included in studies on urban domestic gardens. This study also provides additional knowledge to the fundamentals of the field of urban ecology and the importance of using domestic gardens as an urban green space for conservation purposes. / Thesis (Master of Environmental Sciences)--North-West University, Potchefstroom Campus, 2013 Urban ecology Domestic gardens Socio-economic status Morphospecies Vegetable gardens Lawns
4	Plant- and arthropod diversity of vegetable gardens along a socio-economic gradient within the Tlokwe Municipal Area / Nicola Botha Botha, Nicola January 2012 (has links) Globally urbanization has increased to such an extent that more than half of the human population currently resides in cities. In the years to come, urban expansion will especially take place in developing countries through efforts to improve economic growth and poverty alleviation. This may have a negative effect on native biodiversity within and surrounding urban environments. However, residential areas with a high proportion of gardens form a significantly large part of urban environments and these domestic gardens contribute to the maintenance and preservation of biodiversity in cities. Although the preservation of biodiversity in these gardens is important in the overall conservation of urban green spaces, little is known about how these gardens can possibly contribute to conservation purposes in urban areas. Bearing in mind that anthropogenic activities are possible drivers of urban biodiversity, it is vital to quantify socio-economic aspects within urban ecological research. In developing countries, such as South Africa, the inclusion of socio-economic aspects are especially important because there is a wider gap between poor and wealthy households. There are also a larger number of people that are dependent on their gardens for subsistence purposes, such as vegetable gardening. In the Municipal Area of Tlokwe, South Africa, there exists a definite socio-economic gradient from the poorer western to the more affluent eastern part of the city. Five socio-economic status (SES) classes, primarily based on % unemployment, were used in this study. The ultimate aim of this study was therefore to determine the plant- and arthropod diversity within urban domestic gardens along a socio-economic gradient. Vegetable gardens within domestic gardens were selected to quantify plant- and arthropod biodiversity. Biodiversity of adjacent lawns were also sampled for comparative purposes. The study also attempted to determine to what extent socio-economic aspects of city residents may be possible drivers of biodiversity within the gardens. Various other factors that might have an effect on the plant and/or arthropod diversity were included such as soil characteristics, specific management factors of the gardens and other land-uses surrounding domestic gardens. Arthropod diversity was surveyd by means of pitfall traps and suction sampling in eight 0.25 m2 squares along an 8 m transect in each representative garden. Arthropods were identified up to morphospecies level. Vegetation was surveyed along the same transect and total species composition was determined. Plants were identified up to species level. The plant and arthropod surveys were conducted in both the vegetable gardens and lawns of all SES classes. For the soil samples a 1:2.5 water analysis was conducted. A social survey was conducted in all representative gardens by means of a questionnaire and a SPOT 5 satellite imagery was used to determine the land-use types in the areas surrounding the participating gardens. All the above mentioned factors were compared between the different SES classes. Diversity indices for the arthropods, multivariate statistical analyses and ANOVA analyses were applied to test for meaningful variables between socio-economic status classes as well as vegetable gardens and lawns. From the results it was evident that the more affluent SES classes had significantly higher arthropod diversity values, whilst the lower income classes had higher plant diversity. The factor analysis between the plants and arthropods with the surrounding land-uses revealed two significant factors. Firstly, arthropod diversity was influenced by domestic gardens in the surrounding landscape and there was a positive correlation between these two variables. This indicates that a high percentage of surrounding domestic gardens were possible drivers of arthropod diversity. No correlations were evident between plant and arthropod diversity. Secondly, the other significant factor showed that one SES class had a significantly higher percentage of woodlands and grasslands as opposed to two of the other classes that had a significantly higher percentage of built structures within the surrounding area. Differences were also apparent between the SES classes concerning management regimes, financial stability and level of education. The two more affluent SES classes had obtained a higher level of education and income and had management practices that were uncommon in the three poorer SES classes. This study proposes that domestic gardens are a means to conserve biodiversity in cities. Vegetable gardens in domestic gardens will also be able to harbour a larger diversity of plants and arthropods than the lawns. The socio-economic status of residents also had a significant effect on biodiversity and therefore it should be included in studies on urban domestic gardens. This study also provides additional knowledge to the fundamentals of the field of urban ecology and the importance of using domestic gardens as an urban green space for conservation purposes. / Thesis (Master of Environmental Sciences)--North-West University, Potchefstroom Campus, 2013 Urban ecology Domestic gardens Socio-economic status Morphospecies Vegetable gardens Lawns
5	Exploring the taxonomy of a facultative selfing, polymorphic land snail: the genus Rumina, Gastropoda Pulmonata / Etude de la taxonomie d'un escargot terrestre polymorphique et capable de se reproduire par l'autofécondation: le genre Rumina, Gastropoda Pulmonata Prévot, Vanya 14 September 2011 (has links) Le genre Rumina Risso, 1926 (Subulinidae) est constitué de gastéropodes terrestres, hermaphrodites et capables de réaliser de l’autofécondation ainsi que de la fécondation croisée. Plusieurs espèces ont été décrites sur base de subtiles différences morphologiques telles que la forme et la taille de la coquille et la coloration du corps. Trois espèces sont actuellement toujours reconnue dans la littérature: Rumina decollata (Linnaeus, 1758), R. saharica Pallary, 1901 et R. paivae (Lowe 1861). Cependant, uniquement le statut spécifique de R. decollata et R. saharica a été confirmé par la morphologie de la coquille et l'anatomie génitale. L’objectif de la thèse est de clarifier la taxonomie du genre Rumina par une approche de taxonomie intégrative en associant des caractères moléculaires, morphologiques et anatomiques ainsi que les méthodes issues de la génétique des populations. Quatre gènes mitochondriaux et deux gènes nucléaires ont été séquencés pour reconstruire la phylogénie de Rumina. Les résultats confirment le statut d’espèce de R. saharica, excluent R. paivae en tant qu’espèce et proposent une nouvelle classification pour R. decollata en 6 espèces phylogénétiques. R. saharica est aussi confirmé en tant qu’espèce morphologique suite à une étude morphomètrique, cependant les nouvelles espèces de R. decollata n’ont pas pu être différenciées ni par les variables morphologiques testées, ni par des particularités dans l’anatomie génitale. Néanmoins, deux des espèces de R. decollata correspondent à deux morphotypes (clair et foncé) auparavant décrits et étudiés dans la région de Montpellier. L’étude de ces deux espèces phylogénétiques avec des microsatellites et allozymes permettent de confirmer le statut spécifique des deux morphotypes et révèlent que R. decollata pratique l’autofécondation croisée à un taux supérieur à celui rapporté dans la littérature, défiant ainsi la règle selon laquelle les hermaphrodites pratiqueraient exclusivement l’autofécondation ou exclusivement la fécondation croisée. L’espèce correspondante au morphotype foncé a été introduite en plusieurs endroits du monde et semble être l’espèce possédant la plus grande capacité invasive parmi les Rumina. Ainsi, nos résultats suggèrent que le genre Rumina, auparavant décrit comme étant composé de trois espèces, est en fait un complexe de sept espèces, qui doivent être davantage étudiées de façon à confirmer leur statut d’espèce par d’autres concepts d’espèce. / Rumina spp. Risso, 1826 (family Subulinidae) is a hermaphroditic terrestrial snail, performing both selfing and outcrossing. Several nominal species have been described based on subtle differences in the shape and size of the shell, and body coloration. Currently, three taxa are still recognized, viz. R. decollata (Linnaeus, 1758), R. saharica Pallary, 1901 and R. paivae (Lowe, 1860). Yet, species-specific differentiation has only been confirmed for R. decollata and R. saharica, based on shell and genital morphology. This work aims at resolving the taxonomy of the genus Rumina through an integrative taxonomic approach by combining molecular, morphological and anatomical characters, as well as population genetic methods. Four mitochondrial and two nuclear genes were used to infer Rumina’s phylogeny. Results suggest that R. saharica is a phylogentic species, R. paivae is not a phylogenetic species and R. decollata is composed by 6 phylogenetic species. The specific status of R. saharica was confirmed by a morphometric analysis, however the remaining phylogenetic species of R. decollata could not be differentiated neither by the shell characters analyzed nor by the genital anatomy. Nevertheless, two phylogenetic species of Rumina representing the dark and light colored strains previously described in the Montpellier region. The study of both these strains with microsatellites and allozymes confirmed their specific status and revealed that outcrossing might be more prevalent than was previously suggested in the literature, therefore defying the alleged rule that hermaphroditic species should be either strict self-fertilizers or strict outcrosser. The dark strain was introduced in several places through the world and seems to be the one with highest invasive character within the genus Rumina. Therefore, our results suggest that the genus Rumina, previously described as having three species, is in fact a complex of seven species that need to be further explored in order to confirm their species status under other species concepts. / Doctorat en Sciences / info:eu-repo/semantics/nonPublished Biologie Sciences exactes et naturelles Intersexuality in animals Gastropoda -- Classification Gastropoda -- Reproduction Hermaphrodisme chez les animaux Gastéropodes -- Classification Gastéropodes -- Reproduction espèces phylogénetiques espèces morphologiques taxonomie species delimitation phylogenetic species morphometry biological species morphospecies taxonomy morphométrie délimitation d’espèces espèces biologiques Rumina
6	Methodological investigations on vegetation typology and phytogeography of rain forests of tropical Africa Senterre, Bruno B.M.L. 17 June 2005 (has links) I. An original methodological discussion is proposed on the problem of the typology of tropical rain forest’s plant communities, based on the study of forest types across gradients of continentality and elevation, within Atlantic central Africa. These investigations were based on the statement that the main problems in forest typology are related to the non-zonal or zonal character of the different vegetation types and to non considering the relations and differences between forest strata. II. Field data consisted in phytosociological homogeneous sample plots localized within different recognized phytogeographical entities, in a region of tropical Africa where these entities are known to be well conserved. A total of 37 such plots were inventoried in the region extending from the littoral forests of Ndoté, Equatorial Guinea, which are wet evergreen forests, to the continental forests of the Dja, Cameroon, known as evergreen seasonal forests. The studied region also included the oriental Atlantic forests of Equatorial Guinea, known as moist evergreen forests or caesalp forests. In various parts of this continentality gradient, some plots were localized within climax non-zonal formations, namely the submontane rain forests. The emphasis was put on the vegetation of the Monte Alén National Park. The sampling methodology was willing to be as "complete ", including all strata, "quantitative ", enumerating all individuals, and "representative ", within each stratum, as possible. These multi-layers plots were realised using nested sub-plots, with a sampling size of 100 individuals for every ligneous stratum recognized (dominant trees, dominated trees and shrubs) and a sampling size of 200m² for the herbaceous and suffrutex stratum. Forest types were defined independently for each stratum and the differences were analysed. A method was proposed for the simultaneous analysis of all floristic data, converting and standardizing the values from ligneous strata, on the one hand, and from understorey strata, on the other hand. III. Ten forest types were described using IndVal and discussed in the general context of the guineo-congolian region, from a syntaxonomic view point (agglomerative classification) and from a phytogeographical view point (divisive classification). Homologies between these two approaches are described. The proposed phytogeographical system is based on an "open " conception of hierarchical classifications, combining advantages of agglomerative and divisive classifications. In concrete terms, the non-zonal criteria, for example the submontane variants, are categorised separately and in analogy with the zonal criteria, related to the usual phytochoria. Analysis of ecological relationships for the 10 communities showed that the main variables related to the floristic variability in our mainland rain forests are elevation, rainfall, hygrometry (estimated using bryophytes cover levels) and distance to the ocean. The two extremes on the vertical microclimatic gradient, dominant trees stratum and herbaceous stratum, give similar typologies, however canonical analysis showed that for the herbaceous layer, non-zonal variables (hygrometry and elevation) were gaining more importance when the influence of the two zonal variables was attenuated. In every case, spatial autocorrelation was less important than the environment in explaining floristic variability but its role increased in the spatial arrangement of understorey species, whose dispersal capacity is generally lower than canopy trees. The phytosociological, phytogeographical and ecological description of forest types is accompanied by a physiognomical description using biological types spectrum, as well as architectural models, leaf sizes, etc. With regard to diversity, we have demonstrated that species richness was higher from upper to lower strata because of the accumulation in lower strata of species from various strata. On the other hand, the proper stratum diversity, i.e. the structural set, decreased from dominant trees to shrubs. The proper diversity of the herb layer showed relatively high figures mainly due to the higher individual density in relation to the existence of microstrata. Within the 37 sample plots, 1,050 taxa have been identified to species or morpho-species levels, for a total of 25,750 individuals. These taxa represent 442 genus among 104 families. The richest forest type is found on the foothills of the Niefang range, on the windward side. This forest type is also characterised by a high number of oligotypic genus and by species belonging to functional types indicators of glacial refuges. These functional types are defined on the basis of the dispersal capacity and on kind of stand needed for effective germination. We formulated the hypothesis that this kind of "foothills refuge ", characterised by his zonal nature, could have been one of the rare refuges for species from mainland rain forests, while montane and fluvial refuges would mainly have preserved species from non-zonal forest types: (sub)montane and riverine. Based on indicator species of submontane forests, a potential distribution map of this forest type has been realised at the Atlantic central African scale. More than 400 submontane forest localities have been mapped. These forests begin at 400m of altitude near the ocean, and progressively at higher altitude for increasing distance to the ocean. Many lowland localities also comprised submontane species, which could indicate the existence of ecological transgressions. These transgressions would allow migratory tracks for submontane species between isolated mountain ranges, not only during glacial periods, through heights at the northern and southern borders of the congo basin, but also contemporarily through the lowland riverine forest network, in the centre of this basin. Finally, a special attention has been attributed to littoral forests and to some cases of choroecological transgressions, coupled to the ecological equalization phenomenon. plant diversity foothills refuge glacial refuge environmental stability conservation rarefaction curves individuals-area curves species-area curves compensations écologiques Guinée Equatoriale Monte Alén Monts de Cristal Réserve de Ndoté rainforest transgression Monte Alén Cristal mountains strate forestière sous-bois ensemble structural groupe écologique modèles architecturaux guilde types fonctionnels synusie genres monotypiques genres paucispecifiques diversité végétale courbes aire-espèces courbes individus-espèces courbes de raréfaction courbes aire-constance aire minimale morpho-espèces pléistocène constancy-area curves minimal area structural ensemble understorey ecological group synusia oligotypic genus Réserve de Biosphère du Dja Cameroun Quaternaire glaciation refuge glaciaire refuge de pied de massifs montagneux stabilité environnementale conservation transgression forestière phytosociology plant community phytogeography open classification non-zonal nested plots forest types IndVal morphospecies pleistocene littoral forest rain forest afromontane submontane altitudinal belt glaciation Quaternary Equatorial Guinea ecological equalization continentality continentalité étage d’altitude forêt littorale forêt dense humide afromontagnard submontagnard IndVal types forestiers phytosociologie phytogéographie classification ouverte azonal relevés emboîtés multistrates monotypic genus architectural model guild plant functional type Ndoté Reserve Cameroon Dja Biosphere Reserve forest stratum forest layer understory
7	Recherches méthodologiques pour la typologie de la végétation et la phytogéographie des forêts denses d'Afrique tropicale Senterre, Bruno 17 June 2005 (has links) I. An original methodological discussion is proposed on the problem of the typology of tropical rain forest’s plant communities, based on the study of forest types across gradients of continentality and elevation, within Atlantic central Africa. These investigations were based on the statement that the main problems in forest typology are related to the non-zonal or zonal character of the different vegetation types and to non considering the relations and differences between forest strata.<p><p>II. Field data consisted in phytosociological homogeneous sample plots localized within different recognized phytogeographical entities, in a region of tropical Africa where these entities are known to be well conserved. A total of 37 such plots were inventoried in the region extending from the littoral forests of Ndoté, Equatorial Guinea, which are wet evergreen forests, to the continental forests of the Dja, Cameroon, known as evergreen seasonal forests. The studied region also included the oriental Atlantic forests of Equatorial Guinea, known as moist evergreen forests or caesalp forests. In various parts of this continentality gradient, some plots were localized within climax non-zonal formations, namely the submontane rain forests. The emphasis was put on the vegetation of the Monte Alén National Park.<p><p>The sampling methodology was willing to be as "complete ", including all strata, "quantitative ", enumerating all individuals, and "representative ", within each stratum, as possible. These multi-layers plots were realised using nested sub-plots, with a sampling size of 100 individuals for every ligneous stratum recognized (dominant trees, dominated trees and shrubs) and a sampling size of 200m² for the herbaceous and suffrutex stratum.<p><p>Forest types were defined independently for each stratum and the differences were analysed. A method was proposed for the simultaneous analysis of all floristic data, converting and standardizing the values from ligneous strata, on the one hand, and from understorey strata, on the other hand.<p><p>III. Ten forest types were described using IndVal and discussed in the general context of the guineo-congolian region, from a syntaxonomic view point (agglomerative classification) and from a phytogeographical view point (divisive classification). Homologies between these two approaches are described. The proposed phytogeographical system is based on an "open " conception of hierarchical classifications, combining advantages of agglomerative and divisive classifications. In concrete terms, the non-zonal criteria, for example the submontane variants, are categorised separately and in analogy with the zonal criteria, related to the usual phytochoria.<p><p>Analysis of ecological relationships for the 10 communities showed that the main variables related to the floristic variability in our mainland rain forests are elevation, rainfall, hygrometry (estimated using bryophytes cover levels) and distance to the ocean. The two extremes on the vertical microclimatic gradient, dominant trees stratum and herbaceous stratum, give similar typologies, however canonical analysis showed that for the herbaceous layer, non-zonal variables (hygrometry and elevation) were gaining more importance when the influence of the two zonal variables was attenuated. In every case, spatial autocorrelation was less important than the environment in explaining floristic variability but its role increased in the spatial arrangement of understorey species, whose dispersal capacity is generally lower than canopy trees. The phytosociological, phytogeographical and ecological description of forest types is accompanied by a physiognomical description using biological types spectrum, as well as architectural models, leaf sizes, etc.<p><p>With regard to diversity, we have demonstrated that species richness was higher from upper to lower strata because of the accumulation in lower strata of species from various strata. On the other hand, the proper stratum diversity, i.e. the structural set, decreased from dominant trees to shrubs. The proper diversity of the herb layer showed relatively high figures mainly due to the higher individual density in relation to the existence of microstrata. Within the 37 sample plots, 1,050 taxa have been identified to species or morpho-species levels, for a total of 25,750 individuals. These taxa represent 442 genus among 104 families. The richest forest type is found on the foothills of the Niefang range, on the windward side. This forest type is also characterised by a high number of oligotypic genus and by species belonging to functional types indicators of glacial refuges. These functional types are defined on the basis of the dispersal capacity and on kind of stand needed for effective germination. We formulated the hypothesis that this kind of "foothills refuge ", characterised by his zonal nature, could have been one of the rare refuges for species from mainland rain forests, while montane and fluvial refuges would mainly have preserved species from non-zonal forest types: (sub)montane and riverine.<p><p>Based on indicator species of submontane forests, a potential distribution map of this forest type has been realised at the Atlantic central African scale. More than 400 submontane forest localities have been mapped. These forests begin at 400m of altitude near the ocean, and progressively at higher altitude for increasing distance to the ocean. Many lowland localities also comprised submontane species, which could indicate the existence of ecological transgressions. These transgressions would allow migratory tracks for submontane species between isolated mountain ranges, not only during glacial periods, through heights at the northern and southern borders of the congo basin, but also contemporarily through the lowland riverine forest network, in the centre of this basin. Finally, a special attention has been attributed to littoral forests and to some cases of choroecological transgressions, coupled to the ecological equalization phenomenon.<p> / Doctorat en sciences agronomiques et ingénierie biologique / info:eu-repo/semantics/nonPublished Sciences exactes et naturelles Agronomie générale Phytogeography -- Africa, Sub-Saharan Forest surveys -- Africa, Sub-Saharan Phytogéographie -- Afrique noire Inventaires forestiers -- Afrique noire submontane afromontane rain forest littoral forest pleistocene morphospecies IndVal forest types nested plots non-zonal open classification phytogeography plant community phytosociology transgression forestière conservation stabilité environnementale refuge de pied de massifs montagneux refuge glaciaire glaciation Quaternaire Cameroun Réserve de Biosphère du Dja oligotypic genus synusia ecological group understorey structural ensemble minimal area constancy-area curves pléistocène morpho-espèces aire minimale courbes aire-constance courbes de raréfaction courbes individus-espèces courbes aire-espèces diversité végétale genres paucispecifiques genres monotypiques synusie types fonctionnels guilde modèles architecturaux groupe écologique ensemble structural sous-bois strate forestière Cristal mountains Monte Alén rainforest transgression Réserve de Ndoté Monts de Cristal Guinée Equatoriale compensations écologiques species-area curves individuals-area curves rarefaction curves environmental stability glacial refuge foothills refuge plant diversity altitudinal belt Quaternary Equatorial Guinea ecological equalization continentality continentalité étage d’altitude forêt littorale forêt dense humide afromontagnard submontagnard types forestiers phytosociologie phytogéographie classification ouverte azonal relevés emboîtés multistrates monotypic genus architectural model guild plant functional type Ndoté Reserve Cameroon Dja Biosphere Reserve forest stratum forest layer understory

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