An ecological study was conducted at Cascade Springs, Wasatch county, Utah from August 1965, to June 1967. The algae and aquatic moss species were determined and ecological information about these organisms was obtained. Physical data were also collected: bicarbonates, carbonates, free carbon dioxide, hydroxides, oxygen, pH, and temperature. The physical environment of the water was found to be very constant. Tests taken at the different study sites were consistently similar, not only during each day, but all year around. No great variation of any kind was found in the water. In contrast to the water environment, the topography of the spring varied greatly. Variations were noted in bottom composition, sunlight, water depth, velocity, and volume. But although Cascade Springs appears to have a variety of ecological niches, the number of species of algae and mosses were low. Twelve genera and fourteen species of algae, and ten genera and thirteen species of mosses were collected. The terrace region, both edges and pools, were occupied by all algae species. All but Vaucheria were found in the terrace pools, and all but Chara were found on the terrace edge, and none were found in the terrace pools. The only species of algae found in the spring basin were Chara and Spirogyra, and all moss species but one were found in the spring basin. The algae species were most abundant in the spring and summer months, and least abundant in the winter, although the water temperatures remained constant. The mosses were found to be perennials, and they were constant in both abundance and location. One species of algae was found unique to the spring: Plectonema tomasinianum. This was an abundant species that had not been collected elsewhere in Utah. The other species, both algae and mosses, were common in Utah.

Mosses are some of the first colonizers to disturbed sites, yet their roles in early plant community structuring are not well understood. The primary succession zones of volcanoes provide opportunities to conduct natural experiments into how mosses contribute to early plant community formation, as well as how the unique environments found in such zones affect plant traits, particularly those associated with stress tolerance. Though plant community changes have been well-documented since Mount St. Helens (MSH) volcano erupted in 1980, the volcano's moss assemblages, their influence on other plants, and their potential roles in chemical-mediated competition and biogeochemical cycling have garnered little attention. Using a natural stress gradient from primary to secondary succession zones on MSH, and in control and nutrient manipulated test plots, I sought to elucidate how populations of three dominant moss species, Polytrichum juniperinum, Ceratodon purpureus, and Racomitrium canescens, respond to abiotic stress, as well as to provide life history and interaction data on establishment stages of these stress tolerant taxa. I first analyzed possible tradeoffs in survival strategies of four moss communities in test plots along an abiotic stress gradient. In P. juniperinum, seta specific density (mg/mm) increased significantly in response to nitrogen (N) addition. Differences in both vegetative and sexual reproductive morphological measurements were dependent on site and did not correlate with abiotic stress. In C. purpureus, the percentage of total spores germinated increased with N addition. Site dependent responses in nutrient allocation to vegetative and reproductive structures may be a result of phenotypic plasticity alone or may be a result of local adaptation. In mosses adapted to environmental stress, the allocation of nitrogen must be balanced between growth and survival. Efficient nitrogen uptake confers a competitive advantage if allocated to the higher dispersal of quickly germinating spores. Second, my results show the moss R. canescens may be able to inhibit the germination rate of co-occurring moss spores when spores were germinated in moss gametophyte infusions. R. canescens may also inhibit the germination of the co-occurring vascular plant Lupinus lepidus when seeds are germinated within intact moss patches. By uncovering chemical-mediated interactions between mosses on the germination and initial growth of neighboring mosses and vascular plants we can gain a better understanding of the mechanisms stress tolerant plants may use to limit resource competition. Such advantages offer insight into how mosses effectively colonize and affect primary succession landscapes.

Alpine and arctic tree line expansion depends on the establishment of tree seedlings above the current tree line, which is expected to occur with climate warming. However, tree lines often fail to respond to higher temperatures. Other environmental factors are therefore likely important for tree seedling establishment. Above the tree line, establishing seedlings encounter existing vegetation such as bryophytes, which often dominate in arctic and alpine tundra. Bryophytes modify their environment in various ways and may mediate climate change effects on establishing tree seedlings, and with that tree line expansion. The aim of this thesis was to understand if and how the environment, in particular bryophytes, mediates the impact of climate change on tree seedling establishment at the alpine and arctic tree line. This was explored by reviewing literature on tree seedling establishment at alpine and arctic tree lines globally. In addition, tree seedling survival and growth of Betula pubescens and Pinus sylvestris were assessed experimentally. Here, individuals were planted into mono-specific mats of different bryophytes species and exposed to warming and different precipitation regimes. The literature review revealed that besides from temperature, tree seedling establishment is affected by a wide range of abiotic and biotic factors including water, snow, nutrients, light, disturbance and surrounding vegetation. Furthermore the review revealed that for example vegetation can change tree seedling responses to climate change. The experiments showed that especially tree seedling survival was adversely affected by the presence of bryophytes and that the impacts of bryophytes were larger than those of the climate treatments. Seedling growth, on the other hand, was not hampered by the presence of bryophytes, which is in line with earlier findings that seedling survival, growth and seed germination do not respond similarly to changes in environmental conditions. Moreover, we found several indications that vegetation above the tree line, including bryophytes, mediated tree seedling responses to warming and precipitation or snow cover. This thesis shows that temperature alone should not be used to predict future tree seedling establishment above the alpine and arctic tree line and that extrapolations from climate envelope models could strongly over or under estimate tree line responses to warming. This underlines the value of multi-factorial studies for understanding the interplay between warming and other environmental factors and their effects on tree seedling establishment across current tree lines.

Alpine and arctic tree line expansion depends on the establishment of tree seedlings above the current tree line, which is expected to occur with climate warming. However, tree lines often fail to respond to higher temperatures. Other environmental factors are therefore likely important for tree seedling establishment. Above the tree line, establishing seedlings encounter existing vegetation such as bryophytes, which often dominate in arctic and alpine tundra. Bryophytes modify their environment in various ways and may mediate climate change effects on establishing tree seedlings, and with that tree line expansion. The aim of this thesis was to understand if and how the environment, in particular bryophytes, mediates the impact of climate change on tree seedling establishment at the alpine and arctic tree line. This was explored by reviewing literature on tree seedling establishment at alpine and arctic tree lines globally. In addition, tree seedling survival and growth of Betula pubescens and Pinus sylvestris were assessed experimentally. Here, individuals were planted into mono-specific mats of different bryophytes species and exposed to warming and different precipitation regimes. The literature review revealed that besides from temperature, tree seedling establishment is affected by a wide range of abiotic and biotic factors including water, snow, nutrients, light, disturbance and surrounding vegetation. Furthermore the review revealed that for example vegetation can change tree seedling responses to climate change. The experiments showed that especially tree seedling survival was adversely affected by the presence of bryophytes and that the impacts of bryophytes were larger than those of the climate treatments. Seedling growth, on the other hand, was not hampered by the presence of bryophytes, which is in line with earlier findings that seedling survival, growth and seed germination do not respond similarly to changes in environmental conditions. Moreover, we found several indications that vegetation above the tree line, including bryophytes, mediated tree seedling responses to warming and precipitation or snow cover. This thesis shows that temperature alone should not be used to predict future tree seedling establishment above the alpine and arctic tree line and that extrapolations from climate envelope models could strongly over or under estimate tree line responses to warming. This underlines the value of multi-factorial studies for understanding the interplay between warming and other environmental factors and their effects on tree seedling establishment across current tree lines.

Nathaniel Hawthorne’s collection of short stories, Mosses from an Old Manse, serves as his contribution to the philosophical discussions on Transcendentalism in Concord, MA in the early 1840s. While Ralph Waldo Emerson, Henry David Thoreau, and the other individuals involved in the Transcendental club often seem to readily accept the positions presented in Emerson’s work, it is never so simple for Hawthorne. Repeatedly, Hawthorne’s stories demonstrate his difficulty in trying to identify his own opinion on the subject. Though Hawthorne seems to want to believe in the optimistic potential of the spiritual and intellectual ideal presented in Emersonian Transcendentalism, he consistently dwells on the evil and blackness that may be contained in the human heart. The collection of short stories written while Hawthorne lived in Concord and surrounded himself with those dominant literary figures represents the clearest articulation of his ambivalent position on Transcendentalism.

The group of pleurocarpous mosses comprises approximately 5000 species, which corresponds to about half of all mosses. The pleurocarpous mosses (i.e. “the Core Pleurocarps”) form a monophylum, which consists typically of perennial mosses with creeping stems and abundant lateral branches. In pleurocarpous mosses the archegonium and thus also sporophyte development is restricted to the apices of short, specialized lateral branches, in contrast to most other mosses, where archegonia and sporophytes develop terminally on the main axis (acrocarpous) or on major branches (cladocarpous). Traditionally, pleurocarpous mosses have been divided into three orders based mainly on their sporophytic characters. Brotherus described the Neckeraceae in 1925 and placed it into the Leucodontales, later the family has alternatively been divided into two or three separate families: the Thamnobryaceae, the Neckeraceae and the Leptodontaceae. These families have been placed even in different orders (Neckeraceae and Leptodontaceae among the leucodontalean mosses and Thamnobryaceae among hypnalean mosses) according to their peristome structure and the grade of peristome reduction. A growing amount of evidence indicates that a grouping based on sporophytic characters is artificial and based on convergent evolution. According to the latest phylogenetic studies of pleurocarpous mosses, based on molecular data, the Neckeraceae belong to the order Hypnales and share a sister group relationship with the Lembophyllaceae. In the most recent comprehensive classification 28 genera were included in the Neckeraceae family. This classification was based on both morphological and molecular data, but done with limited taxon sampling that did not cover all species of the family. Some previous studies based on molecular data have challenged the family concept of the Neckeraceae, indicating the need for a revision of the family. Here the family concept of the Neckeraceae is revisited, the closest relatives of the family are resolved and its position within pleurocarpous mosses is shown. In addition, new insights into the morphological evolution of the family are provided. Previous phylogenetic studies indicated that branch lengths among pleurocarpous mosses are usually extremely short. Therefore we chose to use mainly non-coding DNA sequences from rapidly evolving DNA regions. The phylogenetic reconstructions are based on extensive sequence data from all genomes: plastid trnS-trnF and rpl16, nuclear ITS1 & 2 and mitochondrial nad5. Both parsimony (PAUP and PRAP2) and Bayesian statistics (MrBayes) were employed for phylogenetic reconstructions. In order to use the information provided by length mutations indels were included in the analyses as binary data using a simple indel coding approach. No severe conflicts appeared between the different methods used, but the indel coding affected the support values of the inferred topologies. Therefore, all support values resulting from different methods are shown along the phylogenetic trees. The morphological features are studied and synapomorphies for each clade formed in the phylogenetic analyses are interpreted. A new delimitation of the family makes it necessary to reconsider the relevance of the morphological description and the morphological features characteristic of the family need to be reconsidered. Due to new groupings, some changes in the morphological circumscriptions of the genera are necessary, resulting in two new genera and several new combinations. Chapter 1 gives a broad overview of the relationships of the pleurocarpous mosses and shows the need for changes in the definition of genera, families and the corresponding nomenclature in this group. Chapter 2 is a population genetic study on the genus Thamnobryum. The main aim of this chapter is to test the species concept in Thamnobryum that are endemic to strictly restricted regions showing only minor differences in the morphological features in comparison to some more common species. In Chapter 3 the monophyly of the Neckeraceae is tested. In addition, in this chapter the ancestral character states of some morphological characters within the Neckeraceae are reconstructed. Chapters 4 and 5 resolve the genus composition and the relationships within the family in more detail. The results of this thesis show that the Neckeraceae need re-circumscription; this includes changes in the genus composition. The Lembophyllaceae is confirmed to be the sister group of the Neckeraceae. In addition to the new phylogeny, the potential evolution of several characters as a result of environmental selection pressures is analyzed. From the ancestral state reconstructions made (using BayesTraits) for both the habitat and a selection of morphological characters, character state distributions and habitat shift appear congruent, peristome reduction being a good example. However, some character states do not correlate with the habitat, suggesting very complex evolutionary patterns underlying these morphological characters. Many widely distributed genera that are composed of several species and seem to be morphologically coherent (Echinodium, Homalia, Thamnobryum, partly Neckera), are shown in this thesis to be polyphyletic. They are replaced by smaller, geographically more restricted genera that at least in some cases (e.g. Thamnomalia, Homalia s.str., Neckera s.str.) seem to form morphologically heterogeneous genera. In other words, morphology can be misleading in the family Neckeraceae even at the genus level and convergent evolution in both morphological and sequence level characters are common within the family. Special habitat conditions have been shown to result in similar morphological structures also in several other moss groups. This kind of convergent evolution occurs in aquatic mosses, and seems to have occurred among the neckeraceous species Thamnobryum alopecurum and its allies. However, similar morphological structure in similar aquatic habitats can also be due to true phylogenetic relationships as is the case within the Neckeraceae for Handeliobryum sikkimense and Hydrocryphae wardii, or the members of Touwia. The geographical grouping seems to be more strongly correlated with the phylogenetic grouping than thought before.

De ekologiskt betydelsefulla småbiotoperna blir allt färre i det moderna jordbruket på grund av intensifieringen av jordbrukslandskapet som resulterat i ett allt mer fragmenterat landskap. För att kunna ge småbiotoper, såsom åkerholmar, ett optimalt skydd behövs flera studier om organismerna på holmarna, deras diversitet och spridning. I denna undersökning studeras artrikedomen av mossor och lavar på busk- och trädbekädda samt öppna åkerholmar i ett öppet- och ett skogsdominerat landskap. Analysen av artrikedom i studien tyder på att de finns fler arter av mossor och lavar i det öppna landskapet i förhållande till det skogsdominerade landskapet samt på de åkerholmarna som saknade busk- och trädvegetation. Dessa miljöer består av torr och stenig mark och växer igen relativt långsamt, vilket är gynnsamt för vissa mossor och lavar. Det fanns ingen signifikant korrelation mellan diversitet och storleken på åkerholmarna eller mellan diversitet och holmarnas avstånd till skog. Antalet arter i ett habitat kan bero på regionala artpoolen samt antalet potentiella kolonisatörer men även lokala processer. Vidare är det viktigt att nämna att studien tyder på att diversitet av mossor och lavar påverkas av förhållandena på holmarna, såsom busk- och trädtäckningen. Detta i kombination med en variation inom dessa små biotoper kan skapa goda förutsättningar för ett landskap med hög diversitet. / The ecologically important small remnant habitats are becoming fewer in modern agriculture due to the intensification of agricultural landscapes that have resulted in an increasingly fragmented landscape. In order to provide an ideal conservation framework for small habitats, such as midfield islets, several studies about the organisms on the islets, their diversity and distribution are needed. In this study I investigated the diversity of mosses and lichens on open midfield islets with and without shrubs and trees in an open and a forest-dominated landscape. The analysis of species diversity in the study suggests that, overall; there are more species of mosses and lichens in the open landscape in relation to the forest-dominated landscape, and on the opened islets lacking shrub and tree vegetation. These environments consist of dry and rocky soils and overgrow relatively slowly, which is favourable for mosses and lichens. There was no significant correlation between diversity and size of the midfield islets or their distance to forest. The number of species in a habitat may depend on regional species pool and the number of potential colonizers but also local processes. Furthermore, it is important to mention that the study suggests that the diversity of mosses and lichens is influenced by the conditions on the midfield islands, such as shrub and tree cover. These conditions, combined with a variation within these small habitats can create good conditions for a landscape with high diversity.

Magistro darbe aprašomi samanų tyrimai Biržų girios skirtingų gupių miškuose. Darbo objektas - Biržų girios skirtingų grupių miškai. Darbo tikslas - ištirti Plunksninės pliusnės, Tridantės bazanijos, Tikrosios garbuonės, Plokščiosios pliusnės, Paprastosios šilsamanės ir Šilinės plunksnės gausą ir gyvybingumą Bižų girios skirtingų grupių miškuose. Darbo metodai – Tyrimams atlikti buvo pasirinkti skirtingų grupių miškai, po 2kv., apie 1ha. Šie suskirstyti į 6 objektus. Kiekviename objekte buvo tirta samanų įvairovė, gyvybingumas ir gausa pagal trys samanų grupes išskiriant po dvi rūšis iš kiekvienos grupės. Naudojamos samanų grupės: Specializuotosios buveinių rūšys; Indikatorinės rūšys ir Lyginamosios rūšys. Darbo rezultatai. Atlikus tyrimus, nustatyta, kad botaniniuose draustiniuose bendras specializuotųjų samanų rūšių padengimo proc. 27,5%, indikatorinių - 40%, o lyginamųjų 45%; Ūkiniuose miškuose: Specializuotųjų buveinių rūšių nerasta, indikatorinių apie 10%, lyginamųjų - 27%. Kertinėse miško buveinėse: Specializuotosios samanų rūšys užima 32%, indikatorinės- 31%, lyginamosios 52,5%. Surinkti duomenys rodo, kad vertingiausias objektas samanų rūšių gausumo ir gyvybingumo atžvilgiu KMB 267/260 kv. / Master’s work describes the research journal of forest moss gruops different woods. Work items - juornal of forests of different groups of forests.The aim – to explore Plunksninės pliusnės, Tridantės bazanijos, Tikrosios garbuonės, Plokščiosios pliusnės, Paprastosios šilsamanės and Šilinės plunksnės Biržų bundance and vitality of forests of different groups of forests. Working methods – the tests were carried out to select groups of different woods, after 2kv. about 1ha. These are divided into 6 sites. Each vitality and abundance of moss under the three groups after the exclusion of two species from each group. Used moss group: Specialized habitat types, indicator species, and comparative views. Results of the work. After the tests, found that the total reserves in its botanical specialized moss species percent coverage - 27,5%, indicator - 40%, and 45% of the comparative; Commercial forests: specialized habitant types were found, indicative of 10%, comparable - 27%. KEB: specialized moss species occupies - 32%, indicator - 31%, and 52,5% comparision; The data collected indicate that the most valuable object of moss species abundance and vitality in respect of KEB 267/260 kv.

The presence of mosses in black spruce forests is known to have an important impact on the availability and abundance of nutrients in this ecosystem. Mosses contribute to long-term accumulation of organic matter and storage of nutrients as well as to short-term nutrient release. In the boreal forest of northwestern Quebec, the effect of mosses on nutrient cycling was examined within the framework of a chronosequence ranging from 25 to 300 years of age. Laboratory and buried bag incubations, total nutrient digests, respirometric and root abundance measurements as well as moisture and temperature measurements were done to characterize the nutritional status of the organic matter profiles. In general, no change in moss accumulation, or nutrient storage or availability across the chronosequence could be detected. There were, however, differences between feather mosses and Sphagnum mosses, the latter having significantly lower levels of nutrients than the former in terms of mineralizable nitrogen and total carbon, nitrogen, phosphorus, magnesium and potassium content. The nutritional profile of moss cores showed different horizons, reflecting differences in organic matter quality down the organic layer. More specifically, nitrogen availability on a concentration basis as well as root abundance decreased with depth. Forest floor temperature seemed to confine the most biologically active horizon, referred to as the active layer, to a shallow depth, however neither temperature nor moisture seemed to explain the nutritional differences between feather mosses and Sphagnum mosses. These results may lead to practical consequences in that they show a clear distinction between the effects of feather mosses and Sphagnum mosses in nutrient cycling, suggesting that moss cover, might be useful as an indicator of site nutritional status. The results also show that accumulation of nutrients in organic surface horizons, as has been observed elsewhere, does not appear to occur in blac

Decomposition and MeHg concentration were measured for Sphagnum fuscum, Sphagnum angustifolium and Chamaedaphne calyculata in a poor fen in the low boreal forest zone of the Canadian Shield. Litter bags were transplanted reciprocally into hummocks, hollows and lawns and retrieved after 1, 2, 3,11 and 15 months. / Mass losses follow the trend: Chamaedaphne calyculata >> S. angustifolium > S. fuscum. Between species differences were far more significant than differences between locations or depths, indicating that litter quality is the major control on decomposition. / McHg concentrations generally increased during decomposition, particularly near the water table: for C. calyculata up to 13000%, for the two Sphagnum mosses up to 500%, suggesting that decomposition stimulates mercury methylation or McHg scavenging. / The discovery of a high McHg substance on C. calyculata leaf surfaces (25 ng g-1) suggests that much plant MeHg data may by skewed by the presence of a biologically active 'film'.