Basalt flows occur in the Snowville area of north-central Utah and the Table Mountain and Holbrook areas of south-central Idaho. All basalt flows are aphanitic in groundmass, and contain olivine, plagioclase, augite, and opaque oxides. They can be distinguished by texture. Snowville basalt has predominantly subophitic to intergranular textures. Table Mountain basalt is fine grained, with stumpy groundmass plagioclase and equant ilmenite crystals. Holbrook basalt has pilotaxitic to intergranular textures, with the presence of plagioclase phenocrysts and characteristic exsolution lamellae in Fe-Ti oxides. The olivine grains in Holbrook area are intensely oxidized to Fe-Ti oxides. Snowville basalt contains olivine phenocrysts (Fo88 -Fo44 ) in a groundmass of olivine (Fo63 -Fo47), augite (Wo42 -Wo36), and plagioclase (An77-An52). The lower flow unit of Table Mountain basalt contains olivine phenocrysts (Fo88-?) in a groundmass of augite (Wo44 En44 Fs17), and plagioclase (An58-An48). The upper flow unit of Table Mountain basalt has olivine phenocrysts (Fo82-Fo65), plagioclase phenocrysts (An73-An67), and plagioclase groundmass (An64-An55). The Holbrook basalt is composed of olivine phenocrysts (Fo67-Fo57)and plagioclase phenocrysts (An68-An43 ) in a groundmass of olivine (Fo59Fos53) augite (Wo39 En44 Fs17), and plagioclase (An67-An35). The basalts of the Snowville and Holbrook areas, represent petrographic, mineralogical, and chemical characteristics of both olivine-tholeiitic basalt and alkali-olivine basalt, whereas Table Mountain upper and lower flow units show their affinity with alkali-olivine basalt. Chemically, basalts from these three areas are consistently high in silica, magnesium, and alkali content. The Snowville basalt has a high Ba content and high strontium isotope ratio. Fractional crystallization models indicate that the basalt flows from the three different areas are genetically unrelated. The testing also suggests that the upper and lower flow units of the Table Mountain area are not genetically related. The basalts of the three areas also can not be evolved from the basalts found at Kelton, the Rozel Hills or Black Mountain. Basalts of the Snowville area have consistently higher magnesium and silica contents than Snake River basalt, Kelton area basalt, and Rozel Hills and Black Mountain basalt, indicating that they may represent what was initially a very primitive basaltic lava. High Ba content and strontium isotope ratio indicate that the Snowville basalt was contaminated by crustal material. Table Mountain and Holbrook basalt may have formed as a result of partial melting from a pyrolite or garnet peridotite mantle.

Sagebrush shrubsteppe ecosystems have increasingly garnered attention as an endangered ecosystem. Ninety nine percent of all sagebrush ecosystems are thought to have been impacted by humans, and over 50% of grassland and shrubsteppe species are believed to be in decline. Most of the research on sagebrush ecosystems has been conducted at lower elevations and in large expanses of sagebrush. A considerable amount of sagebrush is found at higher elevations, often in meadows found within a forest matrix. The role of this high-elevation habitat is poorly understood. We conducted bird, small mammal, vegetation, and soil surveys in sagebrush-dominated mountain meadows within the Wasatch-Cache National Forest in northeastern Utah. Meadows ranged from 0.6 to 782 hectares in size and included an impressive list of associated plant species. We detected two sagebrush-obligate species and numerous shrubsteppe-associated species. Each species appears to respond to different habitat characteristics, but all species that showed a significant relationship with meadow size were more likely to occur in larger meadows. Many species showed no relationship with size, suggesting that while larger meadows were preferred by some species, small meadows could also play an important role as habitat. While sagebrush-dominated mountain meadows were important for some species, we also failed to detect a number of species of interest. In particular, Sage Thrasher, Sage Sparrow, and pygmy rabbit were not found within the study area. North American Breeding Bird data suggests that Sage Thrashers can be found nearby. It is likely that these birds are only found in large expanses, and none of our meadows were large enough to support them. Sagebrush-dominated mountain meadows appear to be an important supplement to large expanses of sagebrush shrubsteppe habitat, but are not substitutable for all species.

Five groups of 400 hatchery rainbow trout, (Salmo gairdneri), were stocked in a small, mountain stream at )-week intervals from June to September, 1972. A fish trap captured any fish moving out of a 500 m study section. Fish began moving at high levels during the first day of each stocking and continued at high rates for 5-8 days (Early Phase), after which movement decreased to low levels for 6-9 days (Late Phase). Early Phase fish moved primarily at night, possibly due to their disoriented state and high subjectivity to stream conditions. Fish moving during Late Phase did so mainly during daylight, probably in response to diurnal periodicity of a day-active food organism in the drift. Forced movement due to social behavior did not seem to be an influencing factor, but the duration of visible light seemed important to moving fish.

High-severity disturbances are the primary drivers of Engelmann spruce-subalpine fir ecosystems in the southern Rocky Mountain. Recently, an unprecedented, landscape-wide (at least 250 km2) spruce beetle outbreak killed virtually all the Engelmann spruce on the Markagunt Plateau in southwestern Utah, USA. Results from dendroecological analyses suggested the combination of antecedent disturbance history and drought-driven stand development was responsible for creating suitable host conditions prior to the recent outbreak. Multiple and consistent lines of evidence suggested mixed- and high-severity fires shaped the development of the Markagunt Plateau. Subsequent stand development, influenced by species-specific differential tree-ring response to drought, resulted in the gradual increase of Engelmann spruce dominance across the landscape. Spatiotemporal outbreak dynamics included the early, independent and spatially synchronous building of beetle populations in moist sites with large Engelmann spruce across the landscape. As the outbreak evolved over time, it is likely temperature anomalies accelerated beetle population growth, leading to more rapid spruce mortality. In the wake of the spruce beetle outbreak, results from simulated potential fire behavior suggested there was a reduction in probability of active crown fire for one or two decades on near-pure Engelmann spruce sites after the outbreak. This counterintuitive result suggested extreme fire behavior is not an inevitable consequence of spruce beetle outbreaks. Regardless of the occurrence of fire, forest response is likely to be dominated by advance regeneration in the seedling bank. Furthermore, because spruce was virtually absent from the understory, forest reorganization is likely to be dominated by subalpine fir. In response to recent outbreaks such as the Markagunt Plateau, silviculturists are questioning what they can do to limit the loss from these likely inevitable spruce beetle outbreaks. Concepts of resistance and resilience can be used in planning vegetation management intended to indirectly control beetle populations by manipulating their habitat (vegetation). Resilient landscapes will ideally have spruce age class diversity and size class diversity in spatially discontinuous patches.

Growth and food habits of 399 Age 0 mountain whitefish from the Logan River, Utah, were studied between March, 1970, and April, 1971. At the end of their first six months of life, whitefish were 86-96 mm total length and weighed 6-8 grams (wet). Total temperature experience was 2,950-3,430 degree- days above 32° F. The length-weight relationship for Age 0 mountain whitefish was best described by three stanzas with slopes of 4.3333 for fish 12.5-17.0 mm total length, 3.4437 for fish 17.0-55.0 mm total length, and 2.8043 for fish 50.0-112.0 mm total length. Scalation commenced at 30-35 mm total length and was complete at 40-50 mm total length. Feeding began before yolk-sac absorption was complete and 85 percent of the total diet in numbers was chironomid larvae. Age 0 mountain whitefish in the Logan River fed during daylight and evening hours, and selected chironomid larvae and other food organisms 2-4 mm long.

Temperatures selected seasonally by adult mountain whitefish were measured in the laboratory in a horizontal gradient. Final preferendum estimates, based on acute (3-hour) preference tests conducted with fish acclimated to 5, 10, and 15 C each season, were 17.7 C (pre-spawning), 11.9 C (post-spawning), 9.9 C (winter), and 16.3 C (spring). Seasonal influence on temperature selection was evident on the basis of differences in final preferenda, covariance analysis of responses of laboratory-acclimated fish, and temperature selection by fish held at ambient river temperatures. Post-spawning and winter groups selected lower temperatures than did pre-spawning and spring groups. Pre-spawnine fish selected temperatures unsuitable for embryo survival. Reproductive status as reflected by gonad size was evidently not a factor that influenced seasonal temperature selection of adult whitefish.

Two basalt flows and an andesite fissure eruption occur in the Rozel Hills - Black Mountain area in Box Elder County, Utah. Both basalt flows are aphanitic, and contain olivine, plagioclase, augite, and opaque oxides. They may be distinguished both chemically and on the basis of their textures. Unit 2 basalt is finer grained, and appears to be associated with a fault in the Black Mountain area. Chemically, it is similar to high-iron lavas in the Craters of the Moon, Idaho area. It has higher alkali, total iron, and titanium contents that Unit 1 basalt. Unit 1 basalt is similar to other tholeiitic basalts in the Basin and Range province. It is coarser-grained and has higher silicon, magnesium, aluminum, and calcium contents than Unit 2 basalt. While the basalts have characteristics of alkali-olivine basalts, such as absence of calcium-poor pyroxene, and a high alkali to silica ratio, both units are hypersthene- and olivine-nonnative and classified as olivine tholeiites. Subsurface basalt in the area appears to represent both lava types, but extensive alteration makes comparison difficult. Andesite from the area is aphanitic, and contains plagioclase, orthopyroxene, and opaque oxides. Temperatures of equilibration calculated from co-existing magnetite and ilmenite range f rom 1148°C to 745°C for basalt. Temperatures calculated from co-existing olivine and clinopyroxene range from 1009°C to 994°C. While the two basalt units cannot be related by fractional crystallization at surface conditions, some parent - daughter relations are likely within each unit, and the andesite is most likely a differentiation product of Unit 1 basalt. Partial melting of pyrolite and spinel lherzolite mantle compositions to produce Unit 1 and Unit 2 basalts was investigated. Since no unique temperature and pressure of equilbration for these melts and residu~l material from each mantle type was found, it is concluded that partial melting of a mantle of pyrolite or spinel lherzolite composition did not produce these lavas. Unit 2 basalt may be derived from Unit 1 basalt by fractionation of high pressure (8 kb) phases. This mechanism, similar to that proposed for the Craters of the Moon - Snake River Plain system, may account for an evolved basalt (Unit 2) with a lower silica content than a less differentiated basalt (Unit 1).

Whitebark pine (Pinus albicaulis Engelmann) is a treeline species in the central Rocky Mountains. Its occupation of high elevations previously protected whitebark pine from long-term mountain pine beetle outbreaks. The mountain pine beetle, however, is currently reaching outbreaks of record magnitude in high-elevation whitebark pine. We used a factorial laboratory experiment to compare mountain pine beetle (Dendroctonus ponderosae Hopkins) life history characteristics between a typical host, lodgepole pine (Pinus contorta Engelmann), and whitebark pine. We tested the effects of natal host and brood host on beetle fecundity, offspring size, and brood sex-ratio. We reared mountain pine beetles from whitebark pine and from lodgepole pine, and infested half of them into their natal host and half into the other host. Fecundity was greater overall in lodgepole pine brood hosts. Among lodgepole brood hosts, beetles from whitebark pine had greater fecundity. Fecundity was also significantly related to phloem thickness, which was greater in lodgepole pine. Offspring were larger from whitebark brood hosts than from lodgepole, regardless of their parents’ natal host. Finally, sex-ratio was closer to 1:1 in lodgepole than in whitebark brood hosts. We conclude that host species affects life history of mountain pine beetle with consequences for individual beetle fitness.

Over the last two decades, mountain pine beetle (Dendroctonus ponderosae) populations reached epidemic levels across much of western North America, including high elevations where cool temperatures previously limited beetle persistence. Many high-elevation pine species are susceptible hosts and experienced high levels of mortality in recent outbreaks, but co-occurring Great Basin bristlecone pine (Pinus longaeva), the longest-living non-clonal organism, were not attacked. I assessed Great Basin bristlecone pine resistance to mountain pine beetle by evaluating mountain pine beetle host selection behavior and reproductive success in this species. To evaluate mountain pine beetle host selection preference for Great Basin bristlecone pine, I used no-choice 48-hour attack box experiments that confined pioneering female beetles onto pairs of living Great Basin bristlecone and limber pine (P. flexilis), a susceptible host tree species. To investigate the effect of induced tree defenses on host selection behavior, I repeated the tests on paired sections of Great Basin bristlecone and limber pines that had been recently cut, thereby removing their capacity for induced defensive reactions to an attack. Mountain pine beetles avoided Great Basin bristlecone pine relative to limber pine, suggesting that Great Basin bristlecone pine has a high level of resistance to mountain pine beetle due at least in part to stimuli that repel pioneering females from initiating attacks, even when induced defenses are compromised. To investigate mountain pine beetle reproductive success in Great Basin bristlecone pine, I compared the mating success, fecundity, and brood production of mountain pine beetle parents placed in cut Great Basin bristlecone pine bolts with that of mountain pine beetles placed in cut bolts of limber pine and lodgepole pine (P. contorta), two susceptible species. Initial reproductive development was similar in all three tree species, but nearly all brood in Great Basin bristlecone pine died before emerging. The extensive offspring mortality observed in Great Basin bristlecone pine may be a key evolutionary driver behind mountain pine beetle aversion to the species. These findings suggest that Great Basin bristlecone pine is a highly resistant species with low vulnerability to climate-driven increases in mountain pine beetle outbreaks at high elevations.

Climate change is likely to disrupt the timing of developmental events (phenology) in insect populations in which development time is largely determined by temperature. Shifting phenology puts insects at risk of being exposed to seasonal weather extremes during sensitive life stages and losing synchrony with biotic resources. Additionally, warming may result in loss of developmental synchronization within a population, making it difficult to find mates or mount mass attacks against well-defended resources at low population densities. It is unknown whether genetic evolution of development time can occur rapidly enough to moderate these effects. The work presented here is largely motivated by the need to understand how mountain pine beetle (MPB) populations will respond to climate change. MPB is an important forest pest from both an economic and ecological perspective, because MPB outbreaks often result in massive timber loss. Recent MPB range expansion and increased outbreak frequency have been linked to warming temperatures. We present a novel approach to modeling the evolution of phenology by allowing the parameters of a phenology model to evolve in response to selection on emergence time and density. We also develop a temperature-dependent phenology model for MPB that accounts for multiple types of developmental variation: variation that persists throughout a life stage, random variation, and variation due to the MPB oviposition mechanism. This model is parameterized using MPB development time data from constant temperature laboratory experiments. We use Laplace's method to approximate steady distributions of the evolution model under stable temperatures. Here the mean phenotype allows for parents and offspring to be oviposited at exactly the same time of year in consecutive generations. These results are verified numerically for both MPB and a two-stage model insect. The evolution model is also applied to investigate the evolution of phenology for MPB and the two-stage model insect under warming temperatures. The model predicts that local populations can only adapt to climate change if development time can adapt so that individuals can complete exactly one generation per year and if the rate of temperature change is moderate.