A família Accipitridae compreende 67 gêneros e 256 espécies globalmente distribuídas e, representada por águias, gaviões e abutres do Velho Mundo, figura como uma das linhagens mais representativas de aves não-passeriformes atuais. O relacionamento filogenético entre os gêneros de Accipitridae consiste no assunto mais debatido em trabalhos dedicados à sistemática da família, com inúmeros esforços voltados ao reconhecimento de subgrupos que, em sua maioria, foram fundamentados em similaridade global entre os táxons. Apesar de haver diversos estudos sobre anatomia comparada do grupo, estudos morfológicos com base em metodologias cladistas estão restritos a uma única investigação, que resultou em uma compreensão limitada sobre a filogenia do grupo. Em contrapartida, na última década surgiram inúmeras hipóteses com base em dados moleculares, que trataram diversos níveis taxonômicos da família. Alguns destes estudos demonstraram que a grande maioria dos agrupamentos tradicionais não corresponde a grupos monofiléticos e propuseram um novo arranjo intrafamiliar para o grupo. Neste contexto, o presente estudo propõe uma hipótese filogenética para os Accipitridae, com base no estudo do esqueleto craniano e pós-craniano, utilizando-se de uma ampla representatividade taxonômica do grupo. Para tal, foram examinados 433 espécimes, pertencentes a coleções brasileiras e norte-americanas, os quais representaram 113 espécies e 59 gêneros de accipitrídeos, além de quatro táxons utilizados como grupos externos, pertencentes à ordem Accipitriformes. A partir do estudo comparado deste material, foram codificados 161 caracteres osteológicos, dos quais 116 são binários e 45 multiestados não-ordenados, atribuídos a 117 táxons terminais. A matriz de caracteres foi submetida às análises de parcimônia, o que resultou em 36 árvores igualmente parcimoniosas. A topologia de consenso estrito mostrou-se bem resolvida e consideravelmente congruente às hipóteses filogenéticas prévias sobre Accipitridae, com base em dados moleculares, uma vez que 11 das 14 subfamílias propostas naqueles estudos foram aqui recuperadas e apresentaram suporte significativo. Assim, com base na topologia obtida, bem como no conhecimento acumulado sobre a filogenia do grupo, foi proposto um rearranjo taxonômico para a família Accipitridae, composto por 10 subfamílias, duas das quais incluem três tribos cada: Gypaetinae (Gypaetini, Polyboroidini e Pernini), Macheiramphinae, Aegypiinae, Circaetinae, Elaninae, Harpiinae, Aquilinae, Haliaeetinae, Milvinae e Buteoninae (Buteonini, Circini e Accipitrini). Adicionalmente, foram feitas recomendações para a atualização da nomenclatura de alguns gêneros e espécies de Accipitridae. / The family Accipitridae comprises 67 genera and 256 species with a global distribution and, represented by hawks, eagles and Old World vultures, it figures as one of the largest lineages of non-passerine modern birds. Phylogenetic relationships among accipitrid genera have been a contentious issue in studies on the family systematics, with many efforts aimed at the recognition of subgroups, most of which have been founded on overall similarity between taxa. Although there have been several studies on the comparative anatomy of the group, morphological studies based on cladistic methods are restricted to a single investigation which provided only limited insights into the phylogeny of the group. In contrast, in the last decade, several hypotheses based on molecular data have been put forward that treat with various taxonomic levels the family. Some of these studies demonstrated that many of the traditional assemblages do not correspond to monophyletic groups, and they have proposed novel intrafamilial arrangements for the family. In this context, the present study presents a phylogenetic hypothesis for the Accipitridae based on a study of the cranial and post-cranial skeletal material from a large taxonomic representation of the group. Some 433 specimens were consulted, from Brazilian and North-American collections, and representing 113 species and 59 accipitrid genera, with an additional four used as outgroup taxa belonging to the order Accipitriformes. From the comparative study of the material, 161 osteological characters were coded, of which 116 were binary and 45 non-ordered multistate and attributed to 117 terminal taxa. The data matrix was submitted to parsimony analysis resulting in 36 equally most parsimonious trees. The strict consensus topology was well-resolved and largely congruent with the phylogenetic hypotheses for Accipitridae based on molecular data, with 11 of the 14 subfamilies proposed by those studies recovered in the present study, all with significant support. Based on the topology obtained, and known phylogenies of the group, a taxonomic rearrangement for the family Accipitridae was proposed comprising 10 subfamilies, two of which each include three tribes: Gypaetinae (Gypaetini, Polyboroidini and Pernini), Macheiramphinae, Aegypiinae, Circaetinae, Elaninae, Harpiinae, Aquilinae, Haliaeetinae, Milvinae and Buteoninae (Buteonini, Circini and Accipitrini). Additional recommendations were made for updating the nomenclature of some genera and species in the Accipitridae.

As garças e os socós, aves da família Ardeidae, são pernaltas com tamanho médio ou grande, possuem hábitos solitários ou gregários e estão distribuídas por todos os continentes exceto a Antártica. Tradicionalmente, a família é colocada dentro da ordem Ciconiiformes, juntamente com Scopidae (ave-cabeça-de-martelo), Threskiornithidae (íbis), Balaenicipitidae (bico-de-sapato) e Ciconiidae (cegonhas). Entretanto, vários autores propõem a separação das garças e socós em ordem à parte, no caso, Ardeiformes. Outro problema relacionado à taxonomia da família Ardeidae refere-se à divergência dos autores quanto à validade ou à abrangência de alguns gêneros. O presente estudo tem como objetivo novos exames morfológicos, especialmente do esqueleto, visando a obtenção de dados que possam prover soluções para a taxonomia dessa família. Foram utilizados 66 esqueletos completos de Ardeidae, representando 22 espécies distribuídas em 13 gêneros. Os dados indicam que Syrigma sibilatrix e Pilherodius pileatus possuem caracteres suficientes para serem mantidas em gêneros monotípicos, dentro da subfamília Ardeinae. Egretta Alba possui caracteres que suportam a sua inclusão no gênero Ardea, como Ardea Alba. Os gêneros Butorides e Bubulcus também possuem características suficientes que são válidas e monotípicas. Em relação a Cochlearius fica claro ser um autêntico ardeídeo, embora possua estrutura singular no crânio, e deve ser tratado como uma garça noturna (Nycticoracinae). O gênero Nyctanassa é outro gênero válido, pois Nyctanassa violacea difere de Nycticorax por importantes detalhes osteológicos. A classificação de Martinez-Villalta & Motis (1992) é mais coerente com os resultados obtidos, apesar de algumas discordâncias, na qual a família é dividida em quatro subfamílias: Ardeinae, Nycticoracinae, Tigrisomatinae e Botaurinae / Herons and bitterns are medium to large sized birds of the family Ardeidae. With long bills, neck and legs they are adapted for wading and feeding on fishes and other kinds of small animals that occur near water. They live in all regions of the world except Antarctica. Traditionally, the family is included in the order Ciconiiformes, together the Scopidae (hammerheads), Threskiornithidae (ibises), Balaenicipitidae (whalebills) and Ciconiidae (storks). However, some authors have proposed a new order, Ardeiformes, for these birds. Another taxonomic problem in the family Ardeidae concerns the validity and the limits of some genera. This study presents a new approach to the morphology, mainly of the skeleton, of this group looking for new data that may provide insights for the taxonomy of this family. Sixty six complete skeletons of Ardeidae, representing 13 genera and 22 species were studied. Our studies indicate that the species Syrigma sibilatrix and Pilherodius pileatus have sufficient characters to be maintained in their monotypic genera within the subfamily Ardeinae. Egretta Alba has several characters that support its inclusion in the genus Ardea, as Ardea Alba. Butorides and Bubulcus also have particular features that are valid and monotypical. For Cochlearius, it is very clear that it is an Ardeidae, and despite its special bill and skull structure, it must be included in the night-heron group (Nycticoracinae). The genus Nyctanassa is another valid genus and differs from species of the genus Nycticorax in several osteological characters. The classification of the Ardeidae by Martinez-Villalta & Motis (1992) is the most consistent with this study, in spite of some differences, and that the family Ardeidae should be divided in four subfamilies: Ardeinae, Nycticoracinae, Tigrisomatinae and Botaurinae

<p>This scientific paper is about the changes in the skeleton caused by the diseases osteoarthritis and osteoporosis. Six males and one female from Banken 1, S:ta Gertrud  and S:t Hans in Visby were chosen for an osteological analysis. All individuals are adults and dates back to the middle ages. One male suffered from both osteoporosis and osteoarthritis and one female and three males suffered from osteoarthritis. One male might have been in the beginning faze of osteoarthritis, and one male shows no sign of any of the diseases. In addition to osteoarthritis and osteoporosis, some of the individuals suffer from other pathological conditions. Some of the bones from most of the individuals were x-rayed at Visby hospital, to see if medical technique could show some additional information to the osteological analysis. The x-rays were interpreted by doctor Staffan Jennerholm from Visby hospital, but other doctors have also participated. The x-rays confirmed results from the osteological analysis in most cases, although it showed new information in several cases. Some bones from three individuals were taken to Roland Alvarssons` Doctor Practice in Visby to measure the bone density, to check if any of the individuals had osteoporosis. The result confirmed that one male had osteoporosis, as expected from the osteological analysis.</p> / Noen steder i oppgaven står det "osteoartrithis", men det skal stå "osteoarthritis"

The Mycenaean cemetery at Kallithea Laganidia is the first comprehensive study of a cemetery sample from the periphery of the Mycenaean world. Previous studies have focused primarily on remains from palace centers. Even though it is known that the Mycenaeans populated Achaea , very little is known about this more rural population. Archaeologically and bioarchaeologically the region of Achaea has been neglected by formal and organized research, and as a result almost nothing is known about the population. This project has three aims. First, to provide new demographic data about sex, age, health, and the culture of these Mycenaeans. Secondly, via osteological analysis, to examine the hypothesis that the social stratification indicated by the associated grave goods in the tombs is reflected in the spatial orientation of each tomb and the health status of the individuals buried in the graves. Finally, to address the issue of “orphaned” archaeological collections, excavated in rescue operations, which then languish in storage for years or decades. The Kallithea Laganidia cemetery was in use from LHIIIA to LHIIIC and consists of one tholos and 23 chamber tombs. The tholos is a monumental high status tomb, and was in use both before and after the construction and use of the chamber tombs. Five of the chamber tombs were selected as a representative sample of the cemetery for this thesis. The tombs contained both men and women, and adults and children were represented among the tombs, indicating that they should provide a reasonable cross section of the population that buried their dead at Kallithea Laganidia. This osteological data showed and confirmed that the status differences seen in the grave goods from the tombs are also reflected in the tombs according to spatial distribution. The varying quality of burial offerings among the tombs of Kallithea Laganidia suggest that the tombs closer to the tholos contain burials of the socially elite, and the tombs farther away from the tholos contain burials of lower social classes. The pathology data collected, and more specifically the dental pathology data, do reflect social stratification among the sample’s five tombs, particularly when looking at antemortem tooth loss and severe dental wear. In addition, there are indications of status or behaviour differences between the sexes. Kallithean women seem to have been exposed to infection during life more often than men. Women have higher rates of infectious disease, and indications of more antemortem cranial trauma than men. Also, the presence of men, women, and children among secondary burials within these tombs suggested that there is a familial or linear tie within each tomb. The Kallithea Laganidia cemetery has the potential to yield new and informative data about the Achaean Mycenaean population. From this small sampling of 38 burials from five tombs, already the demography and paleopathology of this peripheral group is beginning to be deciphered.

This thesis deals with the importance of boar tusks to the people at the Middle Neolithic settlement of Ajvide, Eksta parish on Gotland. The thesis discusses various issues concerning the Ajvide swine and their tusks, such as the question of domestication and the various functions ascribed to tusks. In order to cast light upon these questions the result of a study of all the tusks found on Ajvide thus far as well as their respective grave context is presented. The study also includes an osteological analysis of the tusks. Furthermore, ethnoarchaeological comparisons are made with Papua New Guinea and Vanuatu; contemporary societies presumed to share common traits with Middle Neolithic societies on Gotland and where swine and their tusks likewise assume a prominent position.

The purpose of this research is to better understand the health of the Roebuck Iroquois population through the study of the growth of its children. Four avenues of osteological analysis are employed to investigate this topic: the comparison of craniofacial measurements to dental age, comparison of craniofacial measurements with one another, assessment of fluctuating asymmetry, and comparisons to other populations. The Roebuck subadults demonstrated a pattern of craniofacial growth consistent with the pattern for a normal, healthy child. Growth spurts in the craniofacial complex were more difficult to observe and interpret than spurs in the long bones. Fluctuating asymmetry was not found in any of the bilateral measurements, indicating that any stress experienced by the Roebuck subadults during development was not great enough to have a detectable impact on cranial symmetry. Roebuck appeared to be similar in size and growth to two other populations with similar subsistence strategies and diets.

The purpose of this research is to better understand the health of the Roebuck Iroquois population through the study of the growth of its children. Four avenues of osteological analysis are employed to investigate this topic: the comparison of craniofacial measurements to dental age, comparison of craniofacial measurements with one another, assessment of fluctuating asymmetry, and comparisons to other populations. The Roebuck subadults demonstrated a pattern of craniofacial growth consistent with the pattern for a normal, healthy child. Growth spurts in the craniofacial complex were more difficult to observe and interpret than spurs in the long bones. Fluctuating asymmetry was not found in any of the bilateral measurements, indicating that any stress experienced by the Roebuck subadults during development was not great enough to have a detectable impact on cranial symmetry. Roebuck appeared to be similar in size and growth to two other populations with similar subsistence strategies and diets.

A trauma frequency analysis was undertaken on a skeletal sample (n = 75) from the skeletal collections of the medieval Augustinian Priory of St. Mary Merton and the post-medieval lower cemetery of St. Bride’s Church. Fourty-four individuals exhibited trauma on one or more cranial elements. Cranial bones were arranged in different groupings for analysis: inclusive samples of 100% complete, at least 75% complete, at least 25% complete, entire sample; and independent samples of 100% complete, 75 - <100% complete, 25 - <75% complete, and <25% complete. Crania were categorized as 100% complete and incomplete. Four frequencies were calculated (frequency of lesions, of individuals with lesions, of individuals with multiple lesions, and the number of lesions per injured individuals) for each category and cranial element. The results illustrate a general trend towards a decrease in frequency as more fragmentary material is included, illustrating that including the more fragmentary material may bias the results towards underestimating trauma frequencies. However, Fisher’s exact tests do not show statistically significant differences between frequencies in the independent samples analysis, except for individuals with lesions on the right nasal bone. Further research into the effect of fragmentation and poor preservation in skeletal research, cranial trauma research in particular, is required. / Graduate

The purpose of this study was to use geometric morphometrics (GM) to investigate the relationships between non-metric traits of the human hip bone: the greater sciatic notch (GSN), the ventral arc (VA), the subpubic contour (SPC), and the ischiopubic ramus ridge (IPRR), estimated skeletal sex, and shape. Fifty-nine undocumented left hip bone specimens were visually assessed for skeletal sex using recognized standards of sex estimation for the GSN (Buikstra and Ubelaker, 1994). The VA, SPC, and IPRR were assessed according to Klales et al., (2012). The Non-metric traits were scored on a five-scale scheme. Skeletal sex was classified as either male, possible male, indeterminate sex, possible female, or female. Three-dimensional computer models were created of the hip bones using the NextEngine 3D desktop surface scanner. Thirty landmarks were selected to represent the hip bone in three-dimensional shape for GM analysis. Twenty-seven of the selected landmarks were reliable according to suggested digitizing error measurements. The apex of the auricular surface, the arcurate eminence, and the anterior gluteal line were the least precise in the test for digitizing error. Geometric morphometric analysis of the computer models were performed using MorphoJ software. Principal component analysis identified the patterns of hip bone shape within the sex categories. A Procrustes ANOVA and a Spearman's correlation tested the significance between hip bone shape and estimated skeletal sex, and between hip bone shape and non-metric trait morphology. Patterns of hip bone shape in the ischium could not be identified by sex, however sex differences were identified in ischium size. Patterns of hip bone shape in the whole hip bone, segmented ilium and segmented pubis were distinguishable by larger sex groups (males = male and possible male categories; females = female and possible female categories). Shape patterns alluded to differences between females and possible females, however, shape patterns did not distinguish males from possible males. Individuals of indeterminate sex shared similar hip bone shapes as males and were therefore included in that larger sex group. Hip bone shape was also correlated with GSN, SPC, IPRR, and VA. However, the strength of the correlation differed between non-metric traits and certain components of hip bone shape. The GSN and SPC had the strongest correlation (p=<0.01) with the whole hip bone, the ilium and the pubis at distinguishing between larger male and female sex groups. The IPRR, and GSN had the strongest correlation (p=<0.01) with the pubis at distinguishing females and possible females. The results of the study suggest that non-metric traits can discern patterns of female shape better than patterns of male shape. Further research into discerning patterns of male hip bone shape and non-metric trait variation using GM is suggested. The results of the study also suggest that patterns of pubis shape might exist among females and could be identifiable using pubis non-metric trait scores. This result lends credence to the practice of estimating sex on a five-scale gradient rather than on a male/female dichotomous division, in order to capture the morphological variation of female hip bone better. / Graduate / 0327 / 0339 / heatherisobelrobertson@gmail.com

The Mycenaean cemetery at Kallithea Laganidia is the first comprehensive study of a cemetery sample from the periphery of the Mycenaean world. Previous studies have focused primarily on remains from palace centers. Even though it is known that the Mycenaeans populated Achaea , very little is known about this more rural population. Archaeologically and bioarchaeologically the region of Achaea has been neglected by formal and organized research, and as a result almost nothing is known about the population. This project has three aims. First, to provide new demographic data about sex, age, health, and the culture of these Mycenaeans. Secondly, via osteological analysis, to examine the hypothesis that the social stratification indicated by the associated grave goods in the tombs is reflected in the spatial orientation of each tomb and the health status of the individuals buried in the graves. Finally, to address the issue of “orphaned” archaeological collections, excavated in rescue operations, which then languish in storage for years or decades. The Kallithea Laganidia cemetery was in use from LHIIIA to LHIIIC and consists of one tholos and 23 chamber tombs. The tholos is a monumental high status tomb, and was in use both before and after the construction and use of the chamber tombs. Five of the chamber tombs were selected as a representative sample of the cemetery for this thesis. The tombs contained both men and women, and adults and children were represented among the tombs, indicating that they should provide a reasonable cross section of the population that buried their dead at Kallithea Laganidia. This osteological data showed and confirmed that the status differences seen in the grave goods from the tombs are also reflected in the tombs according to spatial distribution. The varying quality of burial offerings among the tombs of Kallithea Laganidia suggest that the tombs closer to the tholos contain burials of the socially elite, and the tombs farther away from the tholos contain burials of lower social classes. The pathology data collected, and more specifically the dental pathology data, do reflect social stratification among the sample’s five tombs, particularly when looking at antemortem tooth loss and severe dental wear. In addition, there are indications of status or behaviour differences between the sexes. Kallithean women seem to have been exposed to infection during life more often than men. Women have higher rates of infectious disease, and indications of more antemortem cranial trauma than men. Also, the presence of men, women, and children among secondary burials within these tombs suggested that there is a familial or linear tie within each tomb. The Kallithea Laganidia cemetery has the potential to yield new and informative data about the Achaean Mycenaean population. From this small sampling of 38 burials from five tombs, already the demography and paleopathology of this peripheral group is beginning to be deciphered.