This work gives an overview of the composition of the diet of bats in the eastern Mediterranean and the Middle East, discusses the structure of bat communities in different bioregions of the area in terms of hunting strategies and resource partitioning and last but not least, how echolocation and morphological characteristics of bats affect the composition of the diet. Although some samples were relatively small therefore it is necessary to look soberly at their informative value, and thus their importance is undeniable. The study in some cases provides completely the first information regarding the composition of diet in several species. Asellia arabica mainly catches Coleoptera (Scarabaeidae), Triaenops persicus hunts Lepidoptera, but Heteroptera, Coleoptera (Scarabaeidae) and Orthoptera too, while Triaenops parvus is a specialist in hunting Lepidoptera. Rhinopoma muscatellum hunts mainly Formicoidea, followed by Coleoptera (Melolonthinae) and Heteroptera, Rhinopoma hadramauticum hunts Formicoidea. Lepidoptera of different size, Heteroptera and Coleoptera are the prey of Chaerephon nigeriae. Pipistrellus hanaki has a wide niche breadth as other species of the genus Pipistrellus. In this study Brachycera, Auchenorrhyncha and Coleoptera primarily occurred in its diet. Not yet described species of...

This work gives an overview of the composition of the diet of bats in the eastern Mediterranean and the Middle East, discusses the structure of bat communities in different bioregions of the area in terms of hunting strategies and resource partitioning and last but not least, how echolocation and morphological characteristics of bats affect the composition of the diet. Although some samples were relatively small therefore it is necessary to look soberly at their informative value, and thus their importance is undeniable. The study in some cases provides completely the first information regarding the composition of diet in several species. Asellia arabica mainly catches Coleoptera (Scarabaeidae), Triaenops persicus hunts Lepidoptera, but Heteroptera, Coleoptera (Scarabaeidae) and Orthoptera too, while Triaenops parvus is a specialist in hunting Lepidoptera. Rhinopoma muscatellum hunts mainly Formicoidea, followed by Coleoptera (Melolonthinae) and Heteroptera, Rhinopoma hadramauticum hunts Formicoidea. Lepidoptera of different size, Heteroptera and Coleoptera are the prey of Chaerephon nigeriae. Pipistrellus hanaki has a wide niche breadth as other species of the genus Pipistrellus. In this study Brachycera, Auchenorrhyncha and Coleoptera primarily occurred in its diet. Not yet described species of...

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The federally threatened northern spotted owl (Strix occidentalis caurina) is the focus of intensive conservation efforts that have led to much forested land being reserved as habitat for the owl and associated wildlife species throughout the Pacific Northwest of the United States. Recently, however, a relatively new threat to spotted owls has emerged in the form of an invasive competitor: the congeneric barred owl (Strix varia). As barred owls have rapidly expanded their populations into the entire range of the northern spotted owl, mounting evidence indicates that they are displacing, hybridizing with, and even killing spotted owls. The barred owl invasion into western North America has made an already complex conservation issue even more contentious, and a lack of information on the ecological relationships between the 2 species has hampered conservation efforts. During 2007–2009 I investigated spatial relationships, habitat selection, diets, survival, and reproduction of sympatric spotted owls and barred owls in western Oregon, USA. My overall objective was to determine the potential for and possible consequences of competition for space, habitat, and food between the 2 species. My study included 29 spotted owls and 28 barred owls that were radio-marked in 36 neighboring territories and monitored over a 24-month tracking period. Based on repeated surveys of both species, the number of territories occupied by pairs of barred owls in the 745 km² study area (82) greatly outnumbered those occupied by pairs of spotted owls (15). Estimates of mean size of home-ranges and core-use areas of spotted owls (1,843 ha and 305 ha, respectively) were 2–4 times larger than those of barred owls (581 ha and 188 ha, respectively). Individual spotted and barred owls in adjacent territories often had overlapping home ranges, but inter-specific space sharing was largely restricted to broader foraging areas in the home range with minimal spatial overlap among core-use areas. I used an information-theoretic approach to rank discrete choice models representing alternative hypotheses about the influence of forest conditions and interspecific interactions on species-specific patterns of nighttime habitat selection. Spotted owls spent a disproportionate amount of time foraging on steep slopes in ravines dominated by old (>120 yrs old) conifer trees. Barred owls used available forest types more evenly than spotted owls, and were most strongly associated with patches of large hardwood and conifer trees that occupied relatively flat areas along streams. Spotted and barred owls differed in the relative use of old conifer forest (higher for spotted owls) and slope conditions (steeper slopes for spotted owls). I found no evidence that the 2 species differed in their use of young, mature, and riparian-hardwood forest types, and both species avoided forest-nonforest edges. The best resource selection function for spotted owls indicated that the relative probability of a location being selected was reduced if the location was within or in close proximity to a core-use area of a barred owl. I used pellet analysis and measures of food niche overlap to examine the potential for dietary competition between spatially associated pairs of spotted owls and barred owls. I identified 1,223 prey items from 15 territories occupied by pairs of spotted owls and 4,299 prey items from 24 territories occupied by pairs of barred owls. Diets of both species were dominated by nocturnal mammals, but diets of barred owls included many terrestrial, aquatic, and diurnal prey species that were rare or absent in diets of spotted owls. Northern flying squirrels (Glaucomys sabrinus), woodrats (Neotoma fuscipes, N. cinerea), and lagomorphs (Lepus americanus, Sylvilagus bachmani) were particularly important prey for both owl species, accounting for 81% and 49% of total dietary biomass for spotted owls and barred owls, respectively. Dietary overlap between pairs of spotted and barred owls in adjacent territories ranged from 28–70% (mean = 42%) In addition to overlap in resource use, I also identified strong associations between the presence of barred owls and the behavior of spotted owls, as shown by changes in space-use, habitat selection, and reproductive output of spotted owls exposed to different levels of spatial overlap with barred owls in adjacent territories. Barred owls in my study area displayed both numeric and demographic superiority over spotted owls; the annual survival probability of radio-marked spotted owls from known-fate analyses (0.81, SE = 0.05) was lower than that of barred owls (0.92, SE = 0.04), and barred owls produced over 6 times as many young over a 3-year period as spotted owls. Survival of both species was positively associated with an increasing proportion of old (>120 yrs old) conifer forest within the home range, which suggested that availability of old forest was a potential limiting factor in the competitive relationship between the 2 species. When viewed collectively, my results support the hypothesis that interference competition with a high density of barred owls for territorial space can act to constrain the availability of critical resources required for successful recruitment and reproduction of spotted owls. My findings have broad implications for the conservation of spotted owls, as they suggest that spatial heterogeneity in survival and reproduction may arise not only because of differences among territories in the quality of forest habitat, but also because of the spatial distribution of an invasive competitor. / Graduation date: 2012 / This pdf will not be made available until April 12th, 2012.

Chionochloa, a genus of snow tussock grasses native to New Zealand, exhibits pronounced mast seeding. Chionochloa suffers very high levels of pre-dispersal flower and seed predation by three main insects: Eucalyptodiplosis chionochloae, a cecidomyiid midge, which is formally described here; Megacraspedus calamogonus, a gelechiid moth and Diplotoxa similis, a chloropid fly. Seven species of parasitoids that attack these seed predators were discovered. Four species parasitize M. calamogonus (one tachinid fly and three hymenopteran wasps), one parasitizes D. similis (Hymenoptera: Eulophidae) and two parasitize E. chionochloae, (a pteromalid wasp Gastrancistrus sp. and a platygastrid wasp Zelostemma chionochloae, which is given a formal description here). The abundance, predation levels by each of the insect species, and interactions between all the organisms in the community were studied across three elevations at Mount Hutt over three summer seasons. M. calamogonus was most abundant at 450 m altitude during all three seasons. D. similis was most common at 1070 m altitude, while its predation levels peaked in low flowering seasons and decreased in high seasons. E. chionochloae was abundant in all three altitudes and increased its predation levels with increasing flowering intensity. E. chionochloae was confirmed to use prolonged diapause of at least three years. Prolonged diapause was also confirmed in its two parasitoids. Chionochloa plants were manipulated with various treatments to test the effect on diapause in E. chionochloae and its two parasitoids. Treatments included plant warming, root pruning, gibberellic acid sprayed on the plants and combinations of these treatments. All three insects changed their emergence in response to some treatments and therefore it was suggested that combined with risk-spreading diapause, they may use some predicting to emerge from prolonged diapause. E. chionochloae control their diapause following some of the cues that Chionochloa use for flowering, while Z. chionochloae and Gastrancistrus in some cases follow their host’s cues and in others use similar cues as Chionochloa plants. Emergence or diapause predictions differed across elevations and plant species in all three insect seed/flower predators. E. chionochloae had female-biased sex ratios in different populations even after prolonged diapause. There was week evidence that both parasitoid species are female-biased in the first emergence year and male-biased after more than one year in diapause. Therefore it was suggested that diapause is not more costly for females of E. chionochloae and its parasitoid than for males. Females of all three species were not found to be better predictors (i.e, more likely to respond to treatments by not entering extended diapause) than males. The complex interactions of all the organisms in this web are thought to be sensitive to climate, and it was suggested that the global climate change may alter this sensitive system.

At Cape Gantheaume, Kangaroo Island (South Australia), adult male, lactating female and juvenile New Zealand (NZ) and Australian fur seals regularly return to the same colony, creating the potential for intra- and inter-specific foraging competition in nearby waters. I hypothesised that these demographic groups would exhibit distinct foraging strategies, which reduce competition and facilitate their coexistence. I analysed the diet of adult male, adult female and juvenile NZ fur seals and adult male Australian fur seals and studied the diving behaviour of adult male and lactating female NZ fur seals and the at-sea movements of juvenile, adult male and lactating female NZ fur seals. Female diet reflected that of a generalist predator, influenced by prey availability and their dependant pups� fasting abilities. In contrast, adult male NZ and Australian fur seals used larger and more energy-rich prey, most likely because they could more efficiently access and handle such prey. Juvenile fur seals primarily utilised small lantern fish, which occur south of the shelf break, in pelagic waters. Juveniles undertook the longest foraging trips and adult males conducted more lengthy trips than lactating females, which perform relatively brief trips in order to regularly nurse their pups. Unlike lactating females, some adult males appeared to rest underwater by performing dives that were characterised by a period of passive drifting through the water column. The large body sizes of adult males and lactating females facilitated the use of both benthic and pelagic habitats, but adult males dived deeper and for longer than lactating females, facilitating vertical separation of their foraging habitats. Spatial overlap in foraging habitats among the age/sex groups was minimal, because lactating females typically utilised continental shelf waters and males used deeper water over the shelf break, beyond female foraging grounds. Furthermore, juveniles used pelagic waters, up to 1000 km south of the regions used by lactating females and adult males. The age and sex groups in this study employed dramatically different strategies to maximise their survival and reproductive success. Their prey and foraging habitats are likely to be shaped by body size differences, which determine their different physiological constraints and metabolic requirements. I suggest that these physiological constraints and the lactation constraints on females are the primary factors that reduce competition, thereby facilitating niche partitioning.

When species compete for resources, in a stable homogeneous environment, there are two possible outcomes. The first is that one species will out-compete the other and exclude it from the environment. This is known as the competitive exclusion principle. The second is that both species will manage to coexist. Coexistence can only occur if the species’ niches are differentiated such that interspecific competition is minimised, or eliminated. This outcome is known as resource partitioning. Two closely related Australian skink species of the Egernia genus, Egernia multiscutata and Egernia whitii, are abundant and sympatric on Wedge Island in South Australia’s Spencer Gulf. The species are morphologically very similar and appear to have very similar life histories and habitat requirements. Ostensibly, they would compete for limiting resources in this environment. This thesis is the first investigation into resource partitioning in this previously unstudied model organism. I report the results of multi-faceted investigations into the coexistence of the skinks, E. multiscutata and E. whitii on Wedge Island and the evidence for, and mechanisms of, any facultative resource partitioning between them. Study methods involved a transect survey of most of Wedge Island to determine the species’ distributions and any evidence for resource partitioning; a morphological comparison to investigate any potential competitive advantages of either species; a habitat choice experiment to establish retreat-site preferences in the absence of interspecific interference; and, a series of staged dyadic encounter experiments to investigate interspecific competitive interactions. Resource partitioning was evidenced by differential distributions of the species among substrates containing the elements required for permanent refuge shelters. This partitioning was not mediated by avoidance of particular substrates but by the presence of the opponent species, combined with attraction to suitable substrates. Asymmetries in some morphological characters were found to confer a potential competitive advantage to E. multiscutata in agonistic encounters with E. whitii. Both species were found to have the same refuge site preferences when interference competition was experimentally removed. This result was not concordant with observed resource partitioning in the field and suggests that the habitat choices of both species are modified by the presence of the opponent species. Analyses of staged dyadic encounter experiments showed that E. multiscutata was more likely to gain greater access to a contested habitat resource and more likely to exclude E. whitii from the resource than vice-versa. Nevertheless, the outcome of competitive interactions was not completely deterministic and there was some tolerance of co-habitation. E. multiscutata’s competitive advantage was attributable largely to its greater mass and head dimensions relative to snout to vent length. However, differential behavioural responses to the threat of larger opponent size also played an important part in resource partitioning between the species.

Thesis (Ph.D.)--University of Aberdeen, 2005. / Includes bibliographical references. Available in PDF format via the World Wide Web.

Pushing wireless data traffic onto small cells is important for alleviating congestion in the over-loaded macrocellular network. However, the ultimate potential of such load balancing and its effect on overall system performance is not well understood. With the ongoing deployment of multiple classes of access points (APs) with each class differing in transmit power, employed frequency band, and backhaul capacity, the network is evolving into a complex and “organic” heterogeneous network or HetNet. Resorting to system-level simulations for design insights is increasingly prohibitive with such growing network complexity. The goal of this dissertation is to develop realistic yet tractable frameworks to model and analyze load balancing dynamics while incorporating the heterogeneous nature of these networks. First, this dissertation introduces and analyzes a class of user-AP association strategies, called stationary association, and the resulting association cells for HetNets modeled as stationary point processes. A “Feller-paradox”-like relationship is established between the area of the association cell containing the origin and that of a typical association cell. This chapter also provides a foundation for subsequent chapters, as association strategies directly dictate the load distribution across the network. Second, this dissertation proposes a baseline model to characterize downlink rate and signal-to-interference-plus-noise-ratio (SINR) in an M-band K-tier HetNet with a general weighted path loss based association. Each class of APs is modeled as an independent Poisson point process (PPP) and may differ in deployment density, transmit power, bandwidth (resource), and path loss exponent. It is shown that the optimum fraction of traffic offloaded to maximize SINR coverage is not in general the same as the one that maximizes rate coverage. One of the main outcomes is demonstrating the aggressive of- floading required for out-of-band small cells (like WiFi) as compared to those for in-band (like picocells). To achieve aggressive load balancing, the offloaded users often have much lower downlink SINR than they would on the macrocell, particularly in co-channel small cells. This SINR degradation can be partially alleviated through interference avoidance, for example time or frequency resource partitioning, whereby the macrocell turns off in some fraction of such resources. As the third contribution, this dissertation proposes a tractable framework to analyze joint load balancing and resource partitioning in co-channel HetNets. Fourth, this dissertation investigates the impact of uplink load balancing. Power control and spatial interference correlation complicate the mathixematical analysis for the uplink as compared to the downlink. A novel generative model is proposed to characterize the uplink rate distribution as a function of the association and power control parameters, and used to show the optimal amount of channel inversion increases with the path loss variance in the network. In contrast to the downlink, minimum path loss association is shown to be optimal for uplink rate coverage. Fifth, this dissertation develops a model for characterizing rate distribution in self-backhauled millimeter wave (mmWave) cellular networks and thus generalizes the earlier multi-band offloading framework to the co-existence of current ultra high frequency (UHF) HetNets and mmWave networks. MmWave cellular systems will require high gain directional antennas and dense AP deployments. The analysis shows that in sharp contrast to the interferencelimited nature of UHF cellular networks, mmWave networks are usually noiselimited. As a desirable side effect, high gain antennas yield interference isolation, providing an opportunity to incorporate self-backhauling. For load balancing, the large bandwidth at mmWave makes offloading users, with reliable mmWave links, optimal for rate. / text

Meyer’s Parrot Poicephalus meyeri is the smallest of the nine Poicephalus parrots, forming the P. meyeri superspecies complex with five congeners. Their distributional range far exceeds any other African parrot, extending throughout subtropical Africa. Meyer’s Parrots had previously not been studied in the wild, and therefore, gathering high-quality empirical data on their behavioural ecology became a research and conservation priority. The primary aim of the study was to correlate environmental (e.g. rainfall, habitat availability, resource characteristics, food resource abundance and temperature) and social (e.g. inter- and intra-specific competition, predation, and human disturbance) variables with aspects of their ecology (e.g. flight activity, food item preferences, breeding activity, and group dynamics) to evaluate the degree of specialization in resource use (e.g. trophic, nesting and habitat niche metrics). African deforestation rates are the highest in the world, resulting in twelve out of the eighteen Meyer’s Parrot range states undergoing drastic loss of forest cover over the last 25 years. Most commentary on the population status of Meyer’s Parrots and other Poicephalus parrots pre-dates this period of rapid deforestation In addition, over 75 000 wild-caught Meyer’s Parrots and almost 1 million wild-caught Poicephalus parrots have been recorded in international trade since 1975. Empirical data from this study was used to identify ecological weaknesses (e.g. niche specialization or low breeding turnover) for evaluation within the context of deforestation in the African subtropics. Baseline data on the breeding biology and nest cavity requirements of Meyer’s Parrots was also necessary to assess the viability of applying the conservative sustained-harvest model to African parrots. A unifying goal of this study was to advance our knowledge of the ecology of African parrots and other Psittaciformes by assessing the validity of current hypotheses put forward in the literature. The Meyer’s Parrot Project was initiated in January 2004 on Vundumtiki Island in the north-eastern part of the Okavango Delta, Botswana. Due to high flood waters between March and July 2004, road transects were postponed till August 2004. Transects were conducted at Vundumtiki from August 2004 to July 2005 and February 2007 to August 2007, and at Mombo from August 2005 to January 2006. During 480 road transects over 24 months, food item preferences closely tracked fruiting and flowering phenology, resulting in significant positive correlations between Levins’ niche breadth, rainfall and food resource availability. Meyer’s Parrot can, therefore, be considered opportunistic generalists predispersal seed predator that tracks resource availability across a wide suite of potential food items, including 71 different food items from 37 tree species in 16 families. Meyer’s Parrots were, however, found to be habitat specialists preferring established galleries of riverine forest and associated Acacia-Combretum marginal woodland. These strong habitat associations facilitate their wide distribution throughout the Kavango Basin, Linyanti Swamps, down the Zambezi valley, up along the Rift Valley system in associations with the great lakes, through the Upper Nile and the Sudd, and west as far as Lake Chad through the Sahel. Seventy-five nest cavities were measured during this study, including 28 nest cavities utilized by Meyer’s Parrots within the 430ha sample area at Vundumtiki. Over 1700 hours of intensive nest observations at six nest cavities was undertaken. Meyer’s Parrots formed socially monogamous pair-bonds maintained over at least four breeding seasons. Breeding pairs established breeding territories up to an estimated 160ha within which there were 1–6 nest cavities. Eggs hatched asynchronously, yet nestlings fledged synchronously with similar body size and condition. There was evidence to support the incidence of extra-pair copulations, however, mitochondrial DNA sequence data are required to confirm the incidence of extra-pair fertilizations. Meyer’s Parrots had no preferences in regard to nest tree species beyond the incidence of suitable nest cavities, which are selected and further excavated to accommodate their non-random nest cavity preferences. There was a significant non-nesting Meyer’s Parrot population during the breeding season, likely due to this longlived cavity-nester delaying nesting until a suitable breeding territory becomes available. Meyer’s Parrots utilize communal roosts during summer and disperse from them according to the Foraging Dispersal Hypothesis. Due to the requirement to roost during the middle of the day to avoid heat stress, Meyer’s Parrots have bimodal flight and feeding activity patterns. The highest probability of locating Meyer’s Parrots is between 08h30 and 11h00 during summer when both adults are feeding on the seeds of fleshy-fruits in riverine forest communities. Due to the paucity of data on the current distribution and population status of Meyer’s Parrots and other African parrots, a continent-wide survey of all African parrots represents a conservation priority. Current deforestation rates in several Meyer’s Parrot range, their specialist habitat associations, and lack of evidence to support adaptability to a changing landscape mosaic necessitate the re-classification of Meyer’s Parrots as data deficient or nearthreatened. Based on low breeding population due to limited breeding opportunities, the CITES Appendix II wild-caught bird trade should also be halted until the sustainability of this trade has been evaluated and the relevant information made available. / Thesis (Ph.D.)-University of KwaZulu-Natal, Pietermaritzburg, 2008.