Thesis (Ph.D.) -- La Trobe University, 2005. / Submitted in total fulfilment of the requirements for the degree of Doctor of Philosophy to the Zoology Dept., School of Life Sciences, Faculty of Science, Technology and Engineering. Research. Includes bibliographical references (leaves 133-152). Also available via the World Wide Web.

O objetivo deste estudo foi determinar a composição da ictiofauna de riachos da sub-bacia do Alto Jacuí, região centro-norte do Rio Grande do Sul, bem como descrever e compreender as relações que permeiam a biologia alimentar de espécies de Siluriformes e de Characidae nestes mesmos riachos. As coletas foram realizadas bimensalmente de julho de 2012 a junho de 2013, através da técnica de pesca elétrica em 10 riachos. Um total de 13.247 indivíduos foram coletados, pertencentes a 42 espécies, 10 famílias e seis ordens, das quais as que mais contribuíram foram Characiformes (36%), Cichliformes (24%) e Siluriformes (14%), com 15, dez e seis espécies respectivamente. Nós reportamos o primeiro registro de Phalloceros spiloura para o sistema da Laguna dos Patos e cinco espécies ficaram ao nível de gênero devido ao fato de se tratarem de novas espécies e estão sendo revisadas e descritas. O conteúdo estomacal de 1.948 indivíduos, pertencentes a três espécies (Heptapterus sp., Rhamdia quelen e Trichomycterus poikilos) foram analisados pelo método volumétrico, sendo 59 itens alimentares identificados. Em geral, Heptapterus sp. consumiu uma alta proporção de Aegla sp., restos de plantas terrestres e Megaloptera; R. quelen consumiu peixe e Oligochaeta, seguido de Aegla sp.; enquanto a dieta de T. poikilos foi baseada em Simuliidae, Ephemeroptera e Trichoptera. A segregação alimentar das espécies, bem como do fator espécie mais classe de tamanho, foram observadas. Os valores de amplitude de nicho foram altos para todas as espécies, e somente foram baixos para os maiores tamanhos de R. quelen e Heptapterus sp. No geral, as espécies mostraram baixos valores de sobreposição de nicho, e houve uma grande frequência de sobreposição alta e intermediária para as menores classes de tamanho. O modelo nulo confirmou a partição de nicho entre as espécies e uma partilha de recursos quando analisado espécie mais classe de tamanho. Posteriormente, 1.525 estômagos de 11 espécies de Characidae foram analisados. Observou-se uma diferença significativa entre a dieta dessas espécies, com plantas terrestres e aquáticas, restos de insetos terrestres e aquáticos e adultos de Hymenoptera, mostrando-se os itens mais importantes para esta diferenciação. As análises mostraram que os padrões de alimentação estão associados as características morfológicas. A posição subterminal da boca se correlacionou com a ingestão de itens bentônicos (insetos aquáticos e matéria orgânica); boca grande permitiu a ingestão de itens mais robustos como Aegla sp. e peixes; espécies com menor variação de dentes ao longo da série externa do prémaxilar consumiram elevada proporção de plantas e insetos terrestres, assim como as espécies que apresentaram menor número de dentes na maxila. O hábito insetívoro aquático foi observado nos indivíduos com números intermediários e altos de cúspides nos dentes do maxilar. Buscando compreender a coexistência de quatro espécies sintópicas de Characidae, nós calculamos a sobreposição de nicho das espécies por período amostral e para avaliar a importância destes itens alimentares como fontes de nutrientes nós usamos análises de isótopos de carbono (13C) e nitrogênio (14N) para estimar a assimilação relativa. Todos os Characidae consumiram grandes proporções de plantas e insetos aquáticos. As espécies apresentaram baixa sopreposição na maioria dos períodos amostrados. Também, houve uma alta correspondência entre a análise do conteúdo estomacal e o resultado da análise isotópica. Astyanax xiru e Astyanax procerus assimilaram grandes frações de invertebrados terrestres e plantas aquáticas. Bryconamericus iheringi assimilou principalmente invertebrados aquáticos e secundariamente algas, enquanto Bryconamericus sp. assimilou principalmente invertebrados aquáticos e terrestres. Assim, nós demonstramos que as espécies congenéricas têm dietas mais similares, mas que estas quatro espécies de Characidar coexistem por partição de recursos, e que a assimilação de nutrientes dessas espécies coincide com o consumo relativo dos itens alimentares ingeridos. / The objective of this study was to determine the composition of fishes of Alto Jacuí subbasin streams, north-central region of Rio Grande do Sul, as well as describe and understand the relationships that permeate the feeding biology of species of Siluriformes and Characidae in these same streams. Samples were collected bimonthly from July 2012 to June 2013, through electrofishing technique in 10 streams. A total of 13,247 individuals were collected, belonging to 42 species, 10 families and six orders, of which the most contributed, were Characiformes (36%), Cichliformes (24%) and Siluriformes (14%), with 15, ten and six species, respectively. We report the first record of Phalloceros spiloura to the Laguna dos Patos system and five species were to genus level due to the fact they refer to new species and are being reviewed and described. The stomach contents of 1,948 individuals belonging to three species (Heptapterus sp., Rhamdia quelen and Trichomycterus poikilos) were analyzed by volumetric method, with 59 identified food items. Overall, Heptapterus sp. consumed a high proportion of Aegla sp., remains of terrestrial plants and Megaloptera; R. quelen consumed fish and Oligochaeta, followed by Aegla sp.; while T. poikilos diet was based on Simuliidae, Ephemeroptera and Trichoptera. Food segregation of species as well as species plus size class factors, were observed. The niche breadth values were high for all species, and only were low for the larger sizes of R. quelen and Heptapterus sp. Overall, the species showed low niche overlap values, and there was a high frequency overlay high and intermediate for smaller size classes. The null model confirmed the niche partitioning among species and a sharing of resources when analyzed species plus size class. Subsequently, 1,525 stomachs were analyzed representing 11 species of Characidae. There was a significant difference between the diet of these species, with terrestrial and aquatic plants, terrestrial and aquatic insect remains and adult Hymenoptera, being the most important items for this differentiation. The analysis showed that feeding patterns associated with many morphological features. Sub-terminal mouth positions correlated with the ingestion of benthic items (e.g., aquatic insects and organic matter); large mouths permited ingestion of robust items such as macrocrustaceans (e.g., Aegla) and fish; species that have small teeth variation along the outer row of the premaxilla consumed high proportion of plants and terrestrial insects as well as those species that showed a lower number of teeth on the maxilla. Aquatic insectivory was observed in those individuals with intermediary and high number of cusps in the maxilla teeth. To understand the coexistence of four syntopic Characidae species we calculated niche overlap of species for each sampling period, and to evaluate the importance of these food items as nutrient sources, we used carbon (13C) and nitrogen (14N) isotopes analyses to estimate relative assimilation. All characids consumed large proportions of plant material and aquatic insects. The species showed low dietary overlap throughout most of the sampling period. There was high correspondence between the gut content analyses and isotope mixing model estimates. Astyanax xiru and Astyanax procerus assimilated large fractions of terrestrial invertebrates and aquatic plants. Bryconamericus iheringi assimilated primarily aquatic invertebrates and secondarily algae, whereas Bryconamericus sp. assimilated primarily aquatic and terrestrial invertebrates. Therefore, we demonstrate that the congeneric species have more similar diets, but that these four Characidae species coexist by resource partitioning, and that they assimilate nutrients agree with the relative consumption of food items.

O objetivo deste estudo foi determinar a composição da ictiofauna de riachos da sub-bacia do Alto Jacuí, região centro-norte do Rio Grande do Sul, bem como descrever e compreender as relações que permeiam a biologia alimentar de espécies de Siluriformes e de Characidae nestes mesmos riachos. As coletas foram realizadas bimensalmente de julho de 2012 a junho de 2013, através da técnica de pesca elétrica em 10 riachos. Um total de 13.247 indivíduos foram coletados, pertencentes a 42 espécies, 10 famílias e seis ordens, das quais as que mais contribuíram foram Characiformes (36%), Cichliformes (24%) e Siluriformes (14%), com 15, dez e seis espécies respectivamente. Nós reportamos o primeiro registro de Phalloceros spiloura para o sistema da Laguna dos Patos e cinco espécies ficaram ao nível de gênero devido ao fato de se tratarem de novas espécies e estão sendo revisadas e descritas. O conteúdo estomacal de 1.948 indivíduos, pertencentes a três espécies (Heptapterus sp., Rhamdia quelen e Trichomycterus poikilos) foram analisados pelo método volumétrico, sendo 59 itens alimentares identificados. Em geral, Heptapterus sp. consumiu uma alta proporção de Aegla sp., restos de plantas terrestres e Megaloptera; R. quelen consumiu peixe e Oligochaeta, seguido de Aegla sp.; enquanto a dieta de T. poikilos foi baseada em Simuliidae, Ephemeroptera e Trichoptera. A segregação alimentar das espécies, bem como do fator espécie mais classe de tamanho, foram observadas. Os valores de amplitude de nicho foram altos para todas as espécies, e somente foram baixos para os maiores tamanhos de R. quelen e Heptapterus sp. No geral, as espécies mostraram baixos valores de sobreposição de nicho, e houve uma grande frequência de sobreposição alta e intermediária para as menores classes de tamanho. O modelo nulo confirmou a partição de nicho entre as espécies e uma partilha de recursos quando analisado espécie mais classe de tamanho. Posteriormente, 1.525 estômagos de 11 espécies de Characidae foram analisados. Observou-se uma diferença significativa entre a dieta dessas espécies, com plantas terrestres e aquáticas, restos de insetos terrestres e aquáticos e adultos de Hymenoptera, mostrando-se os itens mais importantes para esta diferenciação. As análises mostraram que os padrões de alimentação estão associados as características morfológicas. A posição subterminal da boca se correlacionou com a ingestão de itens bentônicos (insetos aquáticos e matéria orgânica); boca grande permitiu a ingestão de itens mais robustos como Aegla sp. e peixes; espécies com menor variação de dentes ao longo da série externa do prémaxilar consumiram elevada proporção de plantas e insetos terrestres, assim como as espécies que apresentaram menor número de dentes na maxila. O hábito insetívoro aquático foi observado nos indivíduos com números intermediários e altos de cúspides nos dentes do maxilar. Buscando compreender a coexistência de quatro espécies sintópicas de Characidae, nós calculamos a sobreposição de nicho das espécies por período amostral e para avaliar a importância destes itens alimentares como fontes de nutrientes nós usamos análises de isótopos de carbono (13C) e nitrogênio (14N) para estimar a assimilação relativa. Todos os Characidae consumiram grandes proporções de plantas e insetos aquáticos. As espécies apresentaram baixa sopreposição na maioria dos períodos amostrados. Também, houve uma alta correspondência entre a análise do conteúdo estomacal e o resultado da análise isotópica. Astyanax xiru e Astyanax procerus assimilaram grandes frações de invertebrados terrestres e plantas aquáticas. Bryconamericus iheringi assimilou principalmente invertebrados aquáticos e secundariamente algas, enquanto Bryconamericus sp. assimilou principalmente invertebrados aquáticos e terrestres. Assim, nós demonstramos que as espécies congenéricas têm dietas mais similares, mas que estas quatro espécies de Characidar coexistem por partição de recursos, e que a assimilação de nutrientes dessas espécies coincide com o consumo relativo dos itens alimentares ingeridos. / The objective of this study was to determine the composition of fishes of Alto Jacuí subbasin streams, north-central region of Rio Grande do Sul, as well as describe and understand the relationships that permeate the feeding biology of species of Siluriformes and Characidae in these same streams. Samples were collected bimonthly from July 2012 to June 2013, through electrofishing technique in 10 streams. A total of 13,247 individuals were collected, belonging to 42 species, 10 families and six orders, of which the most contributed, were Characiformes (36%), Cichliformes (24%) and Siluriformes (14%), with 15, ten and six species, respectively. We report the first record of Phalloceros spiloura to the Laguna dos Patos system and five species were to genus level due to the fact they refer to new species and are being reviewed and described. The stomach contents of 1,948 individuals belonging to three species (Heptapterus sp., Rhamdia quelen and Trichomycterus poikilos) were analyzed by volumetric method, with 59 identified food items. Overall, Heptapterus sp. consumed a high proportion of Aegla sp., remains of terrestrial plants and Megaloptera; R. quelen consumed fish and Oligochaeta, followed by Aegla sp.; while T. poikilos diet was based on Simuliidae, Ephemeroptera and Trichoptera. Food segregation of species as well as species plus size class factors, were observed. The niche breadth values were high for all species, and only were low for the larger sizes of R. quelen and Heptapterus sp. Overall, the species showed low niche overlap values, and there was a high frequency overlay high and intermediate for smaller size classes. The null model confirmed the niche partitioning among species and a sharing of resources when analyzed species plus size class. Subsequently, 1,525 stomachs were analyzed representing 11 species of Characidae. There was a significant difference between the diet of these species, with terrestrial and aquatic plants, terrestrial and aquatic insect remains and adult Hymenoptera, being the most important items for this differentiation. The analysis showed that feeding patterns associated with many morphological features. Sub-terminal mouth positions correlated with the ingestion of benthic items (e.g., aquatic insects and organic matter); large mouths permited ingestion of robust items such as macrocrustaceans (e.g., Aegla) and fish; species that have small teeth variation along the outer row of the premaxilla consumed high proportion of plants and terrestrial insects as well as those species that showed a lower number of teeth on the maxilla. Aquatic insectivory was observed in those individuals with intermediary and high number of cusps in the maxilla teeth. To understand the coexistence of four syntopic Characidae species we calculated niche overlap of species for each sampling period, and to evaluate the importance of these food items as nutrient sources, we used carbon (13C) and nitrogen (14N) isotopes analyses to estimate relative assimilation. All characids consumed large proportions of plant material and aquatic insects. The species showed low dietary overlap throughout most of the sampling period. There was high correspondence between the gut content analyses and isotope mixing model estimates. Astyanax xiru and Astyanax procerus assimilated large fractions of terrestrial invertebrates and aquatic plants. Bryconamericus iheringi assimilated primarily aquatic invertebrates and secondarily algae, whereas Bryconamericus sp. assimilated primarily aquatic and terrestrial invertebrates. Therefore, we demonstrate that the congeneric species have more similar diets, but that these four Characidae species coexist by resource partitioning, and that they assimilate nutrients agree with the relative consumption of food items.

Orientadores: Sergio Furtado dos Reis, Glauco Machado / Tese (doutorado) - Universidade Estadual de Campinas, Instituto de Biologia / Made available in DSpace on 2018-08-09T02:04:21Z (GMT). No. of bitstreams: 1 Araujo_MarcioSilva_D.pdf: 2407731 bytes, checksum: 8775de8c335558a4381e1293aad0f17e (MD5) Previous issue date: 2007 / Resumo: A teoria ecológica clássica, em especial a teoria de nicho, foi construída sob a suposição de que os indivíduos de uma população são equivalentes em termos da utilização de recursos. Entretanto, é sabido que os indivíduos de uma população podem variar no uso de recursos e que essa variação pode ter importantes implicações ecológicas e evolutivas. Essa variação interindividual pode dar origem a morfotipos discretos (¿polimorfismo de recursos¿) ou ser contínua (¿especialização individual¿). O presente estudo teve como objetivo investigar a variação interindividual no uso de recursos em quatro populações de rãs do Cerrado brasileiro (Leptodactylus sp., L. fuscus, Eleutherodactylus cf. juipoca e Proceratophrys sp.), uma população de vespas-caçadoras de uma área de Mata Atlântica (Trypoxylon albonigrum) e uma população do peixe lacustre Gasterosteus aculeatus da Columbia Britânica, Canada. Houve evidência de variação interindividual em todas as populações estudadas, indicando que esse fenômeno não é exclusivo de comunidades temperadas de baixa diversidade. Houve uma associação entre a amplitude dos nichos populacionais e o grau de variação interindividual, indicando que os nichos individuais permanecem estreitos apesar da expansão do nicho populacional. Esse padrão é consistente com a presença de trade-offs funcionais associados ao uso dos recursos. A base dos trade-offs permanece desconhecida no caso das rãs e das vespas, mas é provavelmente comportamental. No caso de G. aculeatus, os trade-offs têm base morfológica, mas são mediados pelo comportamento. Além disso, foi identificado um padrão de partição de recursos inédito nesses peixes, em que os indivíduos formam microguildas que representam subdivisões dos recursos litorâneos e pelágicos. São propostos dois novos métodos para a investigação da variação intrapopulacional no uso de recursos, um deles baseado no uso de isótopos estáveis de carbono (d13C) e o outro na teoria de redes complexas / Abstract: Ecological theory, and specially niche theory, was built on the assumption that individuals are equivalent in terms of resource use. However, the individuals in a population may vary in their resources, and this interindividual variation may have important ecological and evolutionary implications. Such variation may give rise to discrete morphological groups (¿resource olymorphism¿) or it may be more continuous (¿individual specialization¿). In the present study, we investigated interindividual variation in resource use in four populations of frogs inhabiting the Brazilian Cerrado (Leptodactylus sp., L. fuscus, Eleutherodactylus cf. juipoca e Proceratophrys sp.), one population of hunting-wasp of the Atlantic Rainforest (Trypoxylon albonigrum), and one population of sticklebacks (Gasterosteus aculeatus) from British Columbia, Canada. We found evidence of interindividual diet variation in all studied populations, indicating that such variation is not restricted to temperate, depauperate comunities. There was an association between niche width and the degree of interindividual variation, indicating that individual niches remain constrained as the population niche expands. This pattern is consistent with the presence of functional trade-offs associated with resource use. In the case of the frogs and the wasps, the nature of the trade-offs remains unknown, but are likely to be behavioral. In the sticklebacks, the trade-offs have a morphological basis, but are mediated by behavior. We found that individual sticklebacks partition resources within littoral and within pelagic prey, which represents a finer pattern of resource partitioning than the traditional ¿littoral-pelagic¿ dichotomy. Two new methods for the quantification of interindividual diet variation are proposed, one based on carbon stable isotopes (d13C) and another based on complex-network theory / Doutorado / Ecologia / Doutor em Ecologia

During the 1979 breeding season four pairs of northern harriers, or marsh hawks (Circus cyaneus) and four pairs of short-eared owls (Asio flammeus) were studied in Cache Valley, Utah. The study was concerned solely with diurnal resource utilization, and did not examine the owls' nocturnal activities. The home range of each harrier pair overlapped substantially with that of an owl pair. Percent habitat overlap for hawk-owl pairs varied from 39 percent to 72 percent. Observations were made to determine if differences existed in their utilization of habitat and food resources, or in their daily and seasonal activity patterns. Both species utilized mainly wet old field and pasture habitat types for their hunting efforts. In general wet old fields were utilized more than expected based on their availability, while pasture, bare ground, and harvested field habitats were used less than expected. Pairs of hawks and owls sharing common habitats generally showed differences in preferred hunting habitats. An analysis of variance showed that hawks and owls were making strikes in different habitat types and to some extent in different parts of the habitat. Harriers and owls nested in different habitat types. Breeding seasons of the two species overlapped almost totally, but interspecific differences were detected in time-activity budgets. Overall, the owls were more sedentary than the hawks. Both species spent approximately 10 percent of the day in hunting-related activites, but timing of hunting varied from pair to pair. Overlapping pairs generally differed in their daily distribution of hunting time. The analysis of variance showed that there was a significant difference in the timing of strikes made by harriers and owls. Both species were feeding primarily on small mammals in the study area, and food resources were probably not a limiting factor for either population. Though northern harriers and short-eared owls appear to have a high degree of niche overlap, this study showed that where eight individual pairs of the two species came into contact they differed in time-activity budgets and habitat utilization. Coexistence between these two species may be enhanced by the fact that they both feed on an abundant prey resource. By subtle habitat and time budget preferences, reinforced through interspecific aggression, they can avoid competition.

No description available.

No description available.

\ / Doctorat en sciences, Spécialisation biologie animale / info:eu-repo/semantics/nonPublished

No description available.

Purpose – The purpose of this thesis is to understand how specialist firms develop in a concentrated industry by analyzing how microbreweries develop and operate on the Belgian market. Then, the literature reviewed used to build the theoretical foundation of this work is centered around the Resource Partitioning Theory and Niche Marketing Theory to explain this phenomenon. Design/methodology/approach – A cross-case analysis was undertaken. Five Belgian Microbreweries were examined. The data were gathered through interviews conducted with owners of microbreweries based on a structured questionnaire, as well as through secondary data. In addition, an analysis of the Belgian beer industry was conducted, using as secondary data, such as the existing literature on the beer industry, as well annual reports from Beers Associations. Findings – In opposition to the big companies that dominates the center of the market leveraging on economies of scales, the microbreweries strived to create unique products that have a high quality in term of taste. Moreover, they focused their efforts in their region, creating value first for local consumers and not all have an ambition to sell in the whole country or to internationalize the product. Thus, the microbreweries managed to create both functional and symbolic value to the consumers. However, the authors found that the symbolic value created by the breweries had more component than cultural heritage, local and specialist identity. Then, relationship was of importance for the microbreweries as they sought personal contact with their consumer and tried to be involved in their community as well as cooperated with other breweries (coopetition).   Overall, the microbreweries practices followed what is described in niche-marketing and entrepreneurial literature. Originality/value – First, this research participates to the resource partitioning literature by contributing to the limited knowledge on how specialist firms are operating, Then, this exploratory study is focused on an understudied area that is how microbreweries runs, that is especially true on the Belgian market as it was barely explored before. Thereafter, this thesis can be used as inspiration for new researches in different countries or with different approaches and/or theories.