步行者在都市環境之移動與空間選擇行為，是規劃者長久以來在進行都市空間規劃與分析之重要課題。其中，對步行者行為的了解，更是進行都市空間分析的重要課題。一般來說，步行者的活動行為與空間選擇，會受環境資訊與過去空間經驗等認知之影響，步行者在複雜的都市空間中活動時，如何在環境中進行空間選擇？在空間選擇時，步行者對空間之認知狀況，又會對其空間選擇產生什麼影響？則是一個有趣的課題。 然而，過去探討步行者空間選擇之相關研究，極少全面探討空間認知對個體空間選擇產生之影響，故本研究之目的，在探索步行者空間認知結果，如何影響其對空間選擇意願。本研究以台北市西門徒步區為例，將步行者之空間選擇分為活動點選擇、找路判斷與路徑選擇三部份，分別運用理論歸納、訪談與探索性因素分析(EFA)，建立步行者空間選擇影響因素後，再運用結構方程模式(SEM)，探討步行者對空間選擇影響因素及影響關係，將步行者“空間認知”對其空間選擇之影響關係進行呈現。 研究結果顯示，個體空間認知結果是影響個體空間選擇意願之主要影響因素，其對步行者活動點選擇之影響為間接影響，對步行者路徑選擇之影響則為直接影響，而個體空間認知結果包含了個體找路判斷認知結果與地區環境資訊認知結果兩部分。 在步行者找路判斷之影響因素方面，影響個體步行者找路判斷之因素，分別為地區環境資訊因素與個體空間熟悉因素；其中，地區環境資訊因素為主要影響因素，地區環境資訊因素並對個體找路判斷同時具有直接與間接之影響。 此外，在影響途徑方面，步行者於活動點選擇時，空間認知結果會透過影響地區環境對步行者之吸引，間接影響個體逛選偏好，而個體逛選偏好則對個體活動點空間選擇之意願具有直接影響。而在步行者路徑選擇時，空間認知結果會直接影響步行者之路徑選擇意願，而地區活動吸引亦會透過直接影響個體空間認知結果，間接影響步行者路徑選擇意願。 關鍵字：步行者、空間認知、空間選擇、找路 / Pedestrian movement has been analyzed by urban geographers and environmental psychologists from the mid-1960’. Particularly, spatial choice is an important part of the spatial movement to researchers. Researchers in the past studies have assumed that pedestrian spatial choice can be viewed as the result of utility-maximizing behavior which pedestrian have full environmental information. It was argued that this assumption may not reflect the real behavior. Because pedestrians do not behave with full information. The lacks of full information open the way for cognitive behavioral approach to understand the spatial decision-making process of pedestrians. The aim of this article is to explore how the spatial cognition affects an individual pedestrian’s space choice behavior. The empirical study is conducted in the Shi-Men pedestrian mall in Taipei city and the structural equation model (SEM) is used to obtain the relationship between individual spatial cognition and space choice behavior. The results are shown as follows: First, spatial cognition affects the pedestrian route choice directly but affects stops choice indirectly. The pedestrian spatial cognitive outcome directly affects the environmental attractiveness of stops, the environmental attractiveness of stops directly affects the individual shopping preference and the individual shopping preference directly affects the choice desire of stops when pedestrians choice stops in pedestrian mall. The environmental active attractiveness of stops directly affects the spatial cognition and the spatial cognitive outcome directly affects the desire of space on pedestrians route choice. Spatial cognition includes two parts: one is wayfinding cognition and the other is local environmental information cognition. Second, the factors of pedestrian wayfinding include the degree of difficulty of wayfinding, the local environmental information and the individual familiarity of space. Among these factors, the local environmental information is the main factor of pedestrian wayfinding. Pedestrian wayfinding is influenced by direct factor as well as indirect factor of the local environmental information. Keywords：Pedestrian, Spatial choice, Spatial cognition, Wayfinding.

The hippocampal formation has been implicated in spatial formation for many decades. The hippocampus proper has received the most attention but other regions of the hippocampal formation contribute largely to spatial cognition. This thesis concentrated on one such region, the postsubiculum. The postsubiculum is considered important because it contains head direction cells and because it thought to be a major input to the hippocampus, via the entorhinal cortex. This thesis aims to test the functional role of the rat postsubiculum under two types of situation: one where the rat must rely on idiothetic cues for navigation, and another where the rat has visual cues present and can rely on these for orientation. The thesis also investigates hippocampal place cells and their stability over time after short exposures to novel environments. Chapter 3 of this thesis aimed to test whether the postsubiculum is necessary for path integration during a homing task. Rats were trained on a homing task on a circular platform maze. Once the task was acquired, rats were given lesions of the postsubiculum or sham lesions and then re-tested on the path integration task. The homing performance of rats with lesions of the postsubiculum was as good as that of the sham rats. A series of manipulations suggests that the rats were homing by path integration, confirmed by probe tests. The rats were then tested on a forced-choice delayed alternation T-maze task that revealed a significant impairment in alternation with delays of 5, 30, and 60 seconds. This suggests that the postsubiculum is not necessary for path integration in a homing task but is necessary for avoiding previously visited locations as is necessary in an alternation task. The experiments in Chapters 4 and 5 of this thesis aimed to investigate the effects of postsubiculum pharmacological inactivation on hippocampal CA1 place cells when rats were introduced to a novel environment with visual cues. A necessary first step was to assess place cells without any manipulation of the postsubiculum (Chapter 4) and then use information gained from this in the design of experiments in Chapter 5. Rats chronically implanted with recording electrodes in the CA1 region of the hippocampus were exposed to novel cue-rich environments whilst place fields were recorded. Following delays of 3, 6, or 24 hours, the same cells were recorded again in the same environment but with the cues rotated by 90°. Pixel-by-pixel correlations of the place fields show that stability of the place fields was significantly lower at 24 hours than at 3 hours. Stability after 6 hours was not significantly different from 3 hours. In the third set of experiments, rats were implanted with drug infusion cannulae in the postsubiculum and recording electrodes in CA1. Following infusions of either the AMPA receptor antagonist CXQX, the NMDA receptor antagonist D-AP5 or a control infusion of ACSF, place field stability was assessed as rats were exposed to a cylindrical environment with a single polarising cue card for 3 x 10 minute sessions and then again 6 hours later. There were no differences in place field correlations between the 3 drug conditions, although there was evidence of larger changes in spatial information content between cells in the CNQX and AP5 drug condition, but not the ACSF condition. The results suggest that, under the present testing conditions, place fields stability did not depend upon AMPA receptor-mediated transmission nor did it depend on NMDA receptor-mediated synaptic plasticity.

The thesis focuses on how different languages influence spatial cognition of their speakers, i. e., whether and how the differences in spatial language (linguistic representation of perspective, location, spatial scenes etc.) affect the non-linguistic spatial reasoning (orientation, spatial memory etc.). This issue has for a long time been a part of the studies of the relation between language and thouhgt under the flag of so called Sapir-Whorf hypothesis/Hypothesis of linguistic relativity. In the first half of the theoretical part of the thesis, I attempt to summarize the history of the concept of linguistic relativity since 1950s and to revise some critical claims about linguistic relativity by re-reading Whorf's works (chapter 2). The second half of the theoretical part (chapter 3) focuses in particular on the research of the interrelations between spatial thought and language. In section 3.1, I make a brief note on the notion of space in terms of cognitive linguistics. Section 3.2 provides an selective overview of the previous research of the crosslinguic spatial-cognitive diversity. Sections 3.3 and 3.4 connect the theoretical and the empirical part of the thesis. The research itself is presented in the chapter 4. It experimentally tests the hypothesis, that the language-specific...

L’adaptation des fonctions motrices permet l’optimisation des interactions avec l’environnement et ses modifications. Une des grandes questions posées à ce sujet concerne la spécificité des modifications implémentées. Dans la littérature traditionnelle sur l’adaptation visuo-manuelle au port de prismes, on retrouve une généralisation de l’adaptation à des positions spatiales non-apprises, mais un très faible transfert de l’adaptation aux autres effecteurs moteurs. Par contraste, les résultats thérapeutiques acquis chez le patient négligent depuis 12 ans suggèrent que l’adaptation visuo-manuelle peut produire des effets à tous les niveaux affectés par cette pathologie. Cette opposition apparente pose la question de la validité du modèle pathologique pour explorer l’adaptation sensori-motrice, et une façon d’y répondre est d’explorer les effets de l’adaptation sur les fonctions perturbées par la négligence chez le sujet normal. Ces trois volets de la littérature apportent des éclairages complémentaires sur la question de la généralisation des adaptations. Par la mise en évidence d’une généralisation des effets consécutifs de l’adaptation prismatique, notamment à un niveau transmodal, non impliqué dans la procédure d’adaptation per se, ce travail de thèse apporte des éléments pertinents en terme de niveau d’action et d’organisation des réseaux impliqués, laissant suggérer un effet de restructuration sur des représentations spatiales de haut niveau, permettant d’élargir l’orientation des stratégies de réhabilitation, par la mise en évidence d’une activation dynamique de fonctions et de réseaux liés à l’intégration multi-sensorielle, nécessaire aux représentations spatiales / Adaptation of motor functions allows optimization of interactions with environment and its alterations. One major question concerns specificity of implemented modifications. Classical data about visuo-manual adaptation to prisms reveal generalization of adaptation to non learned spatial locations, but a very poor transfer to others motor effectors. By contrast, therapeutic results obtained in neglect patients since 12 years suggest that visuo-manual adaptation could produce effects at various levels affected by neglect. This apparent opposition raises the question of validity of pathologic model to explore sensori-motor adaptation, and one way to answer is to explore effects of prism adaptation on disturbed functions by neglect in normal subject. These three sections of review bring out complementary lightings about question of adaptations generalization. By underlying generalization of after-effects of prism adaptation, in particular at a transmodal level, non implicated in adaptative procedure per se, these results bring some relevant arguments in terms of level of action and implicated networks organization, suggesting a restructuring effect on high level spatial representations, allowing to enlarge orientation of rehabilitative strategies. These results bring out a dynamic activation of functions and networks linked to multisensory integration, appropriate to spatial representations

Um zu verstehen, wie das Gehirn von Säugetieren funktioniert, untersuchen wir wie neuronale Aktivität einerseits zu Kognition beträgt und andererseits komplexe Verhaltensweisen ermöglicht. Im Fokus dieser Doktorarbeit stehen dabei zwei Regionen der Großhirnrinde der Ratte: der parahippocampale Cortex und der motorische Cortex. Im ersten Teil haben wir neuronale Schaltkreise im parahippocampalen Cortex und in den oberen Schichten des enthorhinalen Cortex untersucht, während Ratten ihre Umgebung räumlich erkunden. Diese beiden Regionen tragen wesentlich zum Orientierungssinn bei. Dabei haben wir herausgefunden, dass anatomische Identität und Einbindung in den Microschaltkreis einerseits räumliche neuronale Signale, wie zum Beispiel der Aktivität von grid cells, border cells und head-direction cells, bestimmen. Andererseits tragen diese beiden Eigenschaften auch zur temporalen Präzision neuronaler Signale bei, wie zum Beispiel in Form von spike bursts, theta Modulation und phase precession. Im zweiten Teil dieser Doktorarbeit untersuchen wir die Aktivität von Neuronen im Vibrissen Motorcortex während komplexer Bewegungsabläufe der Schnurrhaare, die dem natürlichen Repertoire der Ratte entstammen: eigeninitiierte Bewegungen in freier Luft, Berührung von Artgenossen zur sozialen Interaktion und das Abtasten von Objekten. Dabei haben wir herausgefunden, dass neuronale Aktivität im Motorcortex während der Bewegung der Schnurrhaare unterdrückt ist, dass elektrische Microstimulation zum Rückzug der Schnurrhaare führt und, dass pharmakologische Blockade Bewegung der Schnurrhaare fördert. Um diese überraschende Beobachtung in einen breiteren Kontext zu integrieren, endet dieser Teil mit einer Bewertung der Literatur zu der bewegungsunterdrückenden Wirkung von Motorcortex Aktivität bei Nagetieren, Primaten und Menschen. / In order to understand how the mammalian brain works, we must investigate how neural activity contributes to cognition and generates complex behavioral output. In this thesis I present work, which focuses on two regions of the cerebral cortex of rats: parahippocampal cortex and motor cortex. In the first part of the thesis we investigate neural circuits in the parasubiculum and the superficial medial enthorhinal cortex, two structures that play a key role in spatial cognition. Briefly, we find that the in these regions, anatomical identity and microcircuit embedding is a major determinant of both spatial discharge patterns (such as the discharge patterns of grid cells, border cells and head-direction cells) and temporal coding features (such as spike bursts, theta-modulation and phase precession). In the second part of the thesis we investigate the activity of neurons in vibrissa motor cortex during complex motor behaviors, which play a vital role in rat ecology: self-initiated bouts of exploratory whisking in air, whisking to touch conspecifics during social interactions and whisking to palpate objects. Briefly, we find that neural activity decreases during whisking behaviors, that microstimulation leads to whisker retraction and that pharmacological blockade increases whisker movement. Thus, our observations collectively suggest that a primary role of vibrissa motor cortex activity is to suppress whisking behaviors. The second part of the thesis concludes with a literature review of motor suppressive effects of motor cortical activity across rodents, primates and humans to put this unexpected finding in a broader context.

Les gens produisent des mouvements spontanés pendant des tâches de raisonnement spatial. Ces mouvements aident-ils à la performance de la tâche? Nous avons étudié le rôle des mouvements spontanés dans l'orientation spatiale en utilisant des tâches de prise de perspective spatiale (PPS) dans lesquelles les participants devaient imaginer un point de vue différent de leur point de vue actuel. Nous nous sommes concentrés sur des perspectives exigeant des rotations mentales de soi - particulièrement difficiles, elles sont grandement facilitées par un mouvement actif, même en l'absence de vision. La contribution motrice à la performance de la tâche pourrait résulter d'un mécanisme prédictif, qui anticipe les conséquences d'une action avant son exécution, comme un modèle d'anticipation interne (Wolpert & Flanagan, 2001), inhibant par la suite l’exécution du mouvement si nécessaire. Les mouvements observés peuvent être des traces visibles de ce processus. En utilisant un système de capture de mouvements, nous avons montré que les rotations de la tête sont géométriquement liées à la PPS : leur direction et amplitude étaient liées à la direction et à l'angle entre les perspectives réelles et imaginée (Exp. 1). Chez les contrôleurs aériens, qualifiés ou apprentis, seule la direction de la rotation de la tête était liée à la PPS, reflétant probablement l'expertise spatiale ainsi que le rôle crucial de la direction dans la rotation mentale (Exp. 2). Dans un environnement virtuel, les rotations de tête spontanées étaient liées à une performance accrue. Cependant, les rotations volontaires, qui imitent celles qui sont produites spontanément, ne facilitent pas la performance de navigation (Exp. 3), mais l'empêchent lorsqu’elles sont contraires à la direction de la rotation virtuelle. Nos résultats suggèrent une contribution motrice spécifique à l'orientation spatiale, compatible avec la prédiction motrice. / People produce spontaneous movements during spatial reasoning tasks. Do they relate to task performance? We investigated the role of spontaneous movements in spatial orientation using spatial perspective-taking (SPT) tasks where participants adopted imaginary perspectives. We focused on imaginary perspectives requiring mental rotations of the self as they are particularly difficult and greatly facilitated by active movement in the absence of vision. Motor contribution to task performance could result from a predictive mechanism, which anticipates the consequences of an action before its execution, such as an internal forward model (Wolpert & Flanagan, 2001), further inhibiting full rotations of the head. Observed movements may be visible traces of this process. Using motion capture, we showed that head movements are geometrically related to SPT: both the direction and amplitude of head rotations were related to the direction and angle between the actual and imagined perspectives (Exp. 1). In air traffic controllers and apprentices, only the direction of head rotation was related to SPT, probably reflecting spatial expertise and its crucial role in mental rotation (Exp. 2). In a virtual environment, spontaneous head rotations were related to increased performance. However voluntary rotations, emulating the spontaneously produced ones, did not facilitate navigation performance (Exp. 3), but hindered it when inconsistent with the direction of virtual rotation. Overall, our findings suggest a specific motor contribution to spatial orientation consistent with motor prediction.

The concept of proximity is an important aspect of human reasoning. Despite the diversity of applications that require proximity measures, the most intuitive notion is that of spatial nearness. The aim of this thesis is to investigate the underpinnings of the notion of nearness, propose suitable formalisations and apply them to the processing of GIS data. More particularly, this work offers a framework for spatial proximity that supports the development of more intuitive tools for users of geographic data processing applications. Many of the existing spatial reasoning formalisms do not account for proximity at all while others stipulate it by using natural language expressions as symbolic values. Some approaches suggest the association of spatial relations with fuzzy membership grades to be calculated for locations in a map using Euclidean distance. However, distance is not the only factor that influences nearness perception. Hence, previous work suggests that nearness should be defined from a more basic notion of influence area. I argue that this approach is flawed, and that nearness should rather be defined from a new, richer notion of impact area that takes both the nature of an object and the surrounding environment into account. A suitable notion of nearness considers the impact areas of both objects whose degree of nearness is assessed. This is opposed to the common approach of only taking one of both objects, seen as a reference to assess the nearness of the other to it, into consideration. Cognitive findings are incorporated to make the framework more relevant to the users of Geographic Information Systems (GIS) with respect to their own spatial cognition. GIS users bring a wealth of knowledge about physical space, particularly geographic space, into the processing of GIS data. This is taken into account by introducing the notion of context. Context represents either an expert in the context field or information from the context field as collated by an expert. In order to evaluate and to show the practical implications of the framework, experiments are conducted on a GIS dataset incorporating expert knowledge from the Touristic Road Travel domain.

The concept of proximity is an important aspect of human reasoning. Despite the diversity of applications that require proximity measures, the most intuitive notion is that of spatial nearness. The aim of this thesis is to investigate the underpinnings of the notion of nearness, propose suitable formalisations and apply them to the processing of GIS data. More particularly, this work offers a framework for spatial proximity that supports the development of more intuitive tools for users of geographic data processing applications. Many of the existing spatial reasoning formalisms do not account for proximity at all while others stipulate it by using natural language expressions as symbolic values. Some approaches suggest the association of spatial relations with fuzzy membership grades to be calculated for locations in a map using Euclidean distance. However, distance is not the only factor that influences nearness perception. Hence, previous work suggests that nearness should be defined from a more basic notion of influence area. I argue that this approach is flawed, and that nearness should rather be defined from a new, richer notion of impact area that takes both the nature of an object and the surrounding environment into account. A suitable notion of nearness considers the impact areas of both objects whose degree of nearness is assessed. This is opposed to the common approach of only taking one of both objects, seen as a reference to assess the nearness of the other to it, into consideration. Cognitive findings are incorporated to make the framework more relevant to the users of Geographic Information Systems (GIS) with respect to their own spatial cognition. GIS users bring a wealth of knowledge about physical space, particularly geographic space, into the processing of GIS data. This is taken into account by introducing the notion of context. Context represents either an expert in the context field or information from the context field as collated by an expert. In order to evaluate and to show the practical implications of the framework, experiments are conducted on a GIS dataset incorporating expert knowledge from the Touristic Road Travel domain.

Le sens de verticalité constitue un référentiel spatial fondamental pour percevoir et agir dans l'espace. Il permet notamment de déterminer la position de notre corps par rapport à la direction de la gravité et participe à l’organisation du contrôle postural chez l’homme. L'évaluation du sens de verticalité pour une meilleure compréhension des troubles posturaux et spatiaux de patients ayant présentés un accident vasculaire cérébral (AVC) devient pratique courante. Cependant, l’absence de standardisation et de connaissances des qualités psychométriques des mesures de la perception de la verticale limite considérablement l’utilisation clinique de ces outils et l’élaboration d’essais thérapeutiques pour le suivi des troubles posturaux en pathologie neurologique. L’AVC constitue également le modèle lésionnel humain privilégié pour l’étude du sens de verticalité. L'identification des aires cérébrales sous-tendant les processus d’intégration multisensorielle du sens de verticalité reposent majoritairement sur de techniques modernes d’analyse lésionnelle. Cependant, les connaissances des bases cérébrales de la construction et de la mise à jour du sens de verticalité sont encore partielles et appellent à des travaux complémentaires. Nos travaux de recherche visaient à améliorer notre compréhension du sens de verticalité par l’étude de ces altérations en pathologie vasculaire cérébrale, selon une double approche méthodologique et physiopathologique. Nous avons montré que l’évaluation de la perception visuelle de la verticale dans les suites d’un AVC requiert une installation précise des patients (maintien droit du tronc et de la tête chez les patients présentant des troubles posturaux) et doit être basée sur un minimum de 10 essais pour assurer une fidélité inter-essais élevée. Dans ces conditions d’évaluation, l’orientation de la verticale visuelle présente une excellente fidélité inter- et intra-évaluateur, garantissant la fiabilité de cette mesure pour la pratique et la recherche cliniques. Concernant la perception posturale de la verticale, nous proposons une procédure simplifiée et standardisée du test de la roue pour l’évaluation des biais contralésionnels de la verticale posturale après AVC hémisphérique. Enfin, nous identifions au moyen d’une analyse statistique lésionnelle un réseau d’aires corticales et sous-corticales impliqué dans la perception visuelle et posturale de la verticale. Le cœur de ces régions polymodales du sens de verticalité est centré sur le cortex operculo-insulaire et le thalamus postérolatéral, avec une nette prédominance hémisphérique droite et un chevauchement des aires nodales du cortex vestibulaire. Nos résultats nous permettent de mieux comprendre les bases cérébrales du sens de verticalité et constituent des guides importants pour l'utilisation clinique de la mesure du sens de verticalité.Mots clés: Sens de verticalité, AVC, méthodologie d’évaluation, bases neurales, cortex vestibulaire / The sense of verticality is a major spatial referential for perception and action in space. It allows determining our body position relatively to the gravity and it contributes to the organization of the postural control in humans. The assessment of the sense of verticality for a better understanding of postural and spatial disorders in stroke patients becomes common practice. However, the lack of standardization and psychometrical studies of the verticality perception assessment limits considerably the clinical integration of these tools and the development of therapeutic clinical trials for the follow-up of postural disorders in neurological diseases. Stroke is also the primary model for studying the sense of verticality. The identification of neural bases underlying the multisensory integration processes of the sense of verticality is based essentially on modern techniques of lesion analysis. However, our knowledge of the neural bases of the construction and updating of the sense of verticality are partial and require further studies. Our research aimed to improve our understanding of the sense of verticality through study of these disorders after stroke, according to a methodological and physiopathological dual approach. We showed that the assessment of the visual vertical perception after stroke requires a specific postural setting (the trunk and the head maintained upright in stroke patients with postural disorders) and should be based on 10 trials to achieve a high inter-trials reliability. Under these conditions, visual vertical orientation has excellent inter- and intrarater reliability, ensuring the reliability of this measure for both clinical practice and research. For the postural perception of the vertical, we proposed a simplified and standardized procedure with the wheel test to assess contralesional postural vertical biases after hemispheric stroke. Finally, by means of a lesional statistical analysis, we identified a cortical and subcortical areas network of the visual and postural vertical perception. The polymodal regions of the sense of verticality are centered on the operculo-insular cortex and the posterolateral thalamus, with a right hemisphere predominance and a partial overlap with the core regions of the vestibular cortex. Our results provide a better understanding of the neural bases of the sense of verticality in humans and constitute guidelines for the clinical use of the verticality perception measures.Keywords: Sense of verticality, stroke, assessment methodology, neural bases, vestibular cortex.

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