No description available.

Fragile states are characterized by institutions which do not have the political will or ability to reduce poverty in the interests of their citizen, to establish basic social security, to promote a successful development process, and to guarantee security and human rights. The regional disintegration processes after the period of imperialism and the fall of the iron curtain have created many new states, which still are politically unstable and unable for a sustainable development. In the literature such states are describes as "weak", "failing or failed", "collapsed", "conflict or post-conflict" - dependant on the extent of the particular state failure. Several indicators try to describe such states and partly allow for projections of the future development. Then the role of taxation is discussed in detail before recommendations for the development cooperation are presented. Obviously taxation plays a key role for the democratization process in fragile states.

In this study, the hydrology and physical, chemical and biological variables of a shallow lake system including the Lakes Mogan and Eymir between 1997-2005 were evaluated. In Lake Eymir, a biomanipulation study was conducted between August, 1998 &ndash / December, 1999. Upon biomanipulation, Lake Eymir shifted to clearwater state with submerged vegetation domination during 2000-2003. However, in 2004, the lake shifted back to algae-dominated turbid state since the buffer mechanisms provided by submerged plants were absent. In the summer of 2005, fish kills were observed due to algal bloom. However, due to increasing hydraulic residence time in the lake, internal processes became more important for nutrients. Lake Mogan faces seasonal and interannual water level fluctuations. During the low water levels experienced in 2001 and 2005, which coincided with the high hydraulic residence times, the in-lake phosphorus amount was controlled by internal processes rather than external loading. Moreover, results revealed that hydrology and submerged plants were important in the ecology of Lake Mogan. Furthermore, the relationship between the phytoplankton, zooplankton and the environment in Lakes Eymir and Mogan, which was predicted via Canonical Corresponding Analysis, revealed that nutrients and water transparency were both important for plankton communities. Both the top-down and bottom up effects were valid in Lake Eymir, while only the bottom-up effect and submerged plants were important for Lake Mogan. Finally, the present study provided a good example for the submerged plant dominated clearwater state triggered by biomanipulation, and the impact of hydrology on the ecology of shallow lakes.

Prescribed fire is a common tool used to restore native diversity, control invasive species, and reduce fuel loads. However, fire alone can be insufficient to restore pre-settlement vegetation; other factors such as differences in native and invasive species pools, deer herbivory, seed availability, and the spatial pattern of the fire can influence vegetation trajectories and restoration outcomes. Central Texas is a mosaic of savanna and mixed woodlands co-dominated by Quercus buckleyi (Texas red oak) and Juniperus ashei (Ashe juniper). In a savanna, I studied the joint effects of initial species composition (native-dominated or invasive-dominated) and disturbance (high-intensity fire, clipping, or control) on the ability of native species to establish, survive, and resist invasion by Bothriochloa ischaemum, an invasive C4 grass (ch. 1). Native savanna patches were resistant to invasion following high-intensity fire; fire can be used to selectively control B. ischaemum. In central Texas savanna and woodlands, under fire suppression and overabundant white-tailed deer, Quercus spp. are failing to regenerate, while J. ashei is increasing in abundance. To better understand vegetation trajectories following J. ashei removal in savanna, I studied the soil seedbank along a chronosequence of J. ashei invasion (ch. 5). In woodland, I studied the joint effects of prescribed fire and deer (ch. 2), clearing of J. ashei followed by high-intensity slash-pile burns (ch. 3), and wildfires (ch. 4) on the abundance and size of J. ashei and of hardwoods. Hardwoods resprouted vigorously after fire; J. ashei individuals of all sizes were killed by fire and slow to re-colonize. These management interventions failed to increase Q. buckleyi seedling abundance. It is likely that deer control is necessary to allow fire to have positive effects on the regeneration of oaks in this region, and wherever deer are over-abundant. However, deer can indirectly benefit hardwoods by reducing competition with palatable forbs (ch. 3). In general, these results show that fire suppression in central Texas oak-dominated woodlands is causing a shift not to more mesic-adapted species, as observed in the eastern US, but to J. ashei, which is at least as xeric-adapted as oak, a process I termed 'juniperization'. / text

Coral reefs are suffering unprecedented declines in coral cover and species diversity. These changes are often associated with  substantial shifts in community structure to new dominant organisms. Ultimately, these “phase shifts” can be persistent and very difficult to return from. Building insurance against degradation and decreasing the likelihood of reefs undergoing shifts to undesirable states will require sustainable management practices that uphold coral reef resilience. This thesis consists of five papers that contribute new knowledge useful for managing the resilience of coral reefs, and other marine ecosystems. Paper I shows how the morphology of natural substrate (dead coral colonies) can significantly influence coral recruitment patterns. Paper II focuses on larval lipid levels, a key determinant of coral dispersal potential, in a common Caribbean coral (Favia fragum). It shows that i) lipid levels exhibit a significant, non-linear reduction throughout the larval release period of F. fragum and ii) exposure to a common pollutant (copper) could potentially lead to a more rapid lipid consumption in the larvae. Paper III presents a broader analysis of the different undesirable states a coral reef can shift to as a consequence of reef degradation. It concludes that different states are caused by different driving factors and that management must explicitly acknowledge this. Paper IV proposes a suite of resilience indicators that can help managers assess when a coral-dominated reef might be moving towards a shift to an undesirable state. These indicators capture key-processes occuring on different temporal and spatial scales and signal resilience loss early enough for managers to take appropriate measures. Finally, Paper V reviews the feedback loops that reinforce the undesirable states of five important marine ecosystems and suggests certain strategies that can ease the restoration back to healthier conditions. Managing these critical feedbacks will recquire monitoring the processes underpinning these feedbacks, breaking already established feedbacks loops through large-scale management trials and acknowledging transdisciplinary solutions that move management beyond the discipline of ecology / At the time of the doctoral defense, the following papers were unpublished and had a status as follows: Paper 2: Submitted. Paper 5: In progress.

Conselho Nacional de Desenvolvimento Científico e Tecnológico / Alguns recifes de coral da Baía de Todos os Santos passaram a ser dominado pelo zoantídeo Epizoanthus gabrieli em 2003. Fenômeno resultante da degradação dos recife de coral que atinge 20% desses ecossistemas e ameaça outros 35% no mundo. Apesar de sua importância, apenas a mudança na comunidade para o domínio de macroalgas foi suficientemente estudado. Assim, torna-se urgente estudos sobres alterações envolvendo a dominância de outros organismos. Estes fornecem subsídios para produção de modelos funcionais que podem ajudar na tomada de decisão para o manejo destes ecossistemas tanto na prevenção destas alterações quanto na recuperação de suas comunidades. Os objetivos deste trabalho são (i) verificar se este fenômeno constitui uma mudança de fase, a partir da redução da abundância de corais e persistência da alta cobertura de E. gabrieli por pelo menos cinco anos, (ii) avaliar os efeitos da competição entre este zoantídeo e corais adultos e recrutas com experimentos manipulativos e (iii) investigar os efeitos desta dominância na assembleia de peixes recifais. Os resultados confirmaram a existência de uma mudança de fase, sugerindo que a abundância de E. gabrieli aumentou em 2003 ou antes e que até 2007 houve uma redução da cobertura de coral, condição que se mantêm pelo menos até 2013. As três espécies de corais testadas mostram-se muito sensíveis ao contato com E. gabrieli, com necrose em 78% das colônias e ocupação do esqueleto dos corais em 35% dos casos em um período de 118 dias. Além disso, um modelo feito a partir dos dados de proporção de colônias de corais em contato com este zoantídeo e a cobertura de E. gabrieli sugere que quando o zoantídeo atinge 6% de cobertura, 50% das colônias de corais entram em contato com o mesmo. Estes dados são fortes evidências de que a redução da cobertura de coral observada entre 2003 e 2007 foi causada por competição entre estes organismos. Não foi observado efeito negativo no recrutamento do coral em substrato artificial livre em recifes dominados por E. gabrieli, nem nas proximidades das suas colônias. Isso sugere que uma suposta redução da cobertura deste zoantídeo deve ser acompanhada pelo aumento da taxa de recrutamento de corais e que a competição com a inibição do recrutamento não suporta um efeito de histerese. Foi constatado que esta mudança de fase reduz a riqueza de espécies de peixes recifais, apresentando dez espécies a menos que os recifes normais, e que favorecem os invertívoros moveis em detrimento dos carnívoros e invertívoros sésseis. Contudo não se observou diferença na abundância de peixes. / Some coral reefs of Todos os Santos Bay have undergone a drastic change in the benthic community. In 2003, they began to be dominated by the zoanthid Epizoanthus gabrieli. This phenomenon is a consequence of coral reef degradation that now reaches approximately 20% of coral reefs worldwide and and threatens another 35%. The most dramatic consequences of this degradation are the phase shift phenomena which are characterized by reduction of coral abundance and increase of organisms that do not build reefs. Despite their importance, only a phase shift to the domain of macroalgae has been sufficiently studied. Given this situation, it is urgent to create functional models that demonstrate the ecological processes that occur in the installation of alternative states that persist over time. These models provide important information that can help in decision making on management measures both for prevention and recovery of these ecosystems. The aims of this work are (i) to verify if this phenomenon is a phase shift, (i.e. the reduction of the abundance of coral and persistence of high coverage E. gabrieli for more than five years of monitoring), (ii) evaluate the effects of competition between this zoanthid and recruits and adult coral with manipulative experiments and (iii) study the effects of dominance on the fish assemblage. The result confirmed the existence of a phase shift, suggesting that the abundance of E. gabrieli increased in 2003 or before and in 2007 there was a reduction in coral cover, and that this condition remained until at least 2013. The three coral species tested show great sensitivity to contact with E. gabrieli, with necrosis in 78% of the colonies and with overgrow in 35% of cases in a period of 118 days. In addition, a model made from the data of a proportion of coral colonies in contact with this zoanthid and coverage of E. gabrieli suggests that when the zoanthid reaches 6% coverage, 50% of coral colonies come in contact with it. These data are strong evidence that the reduction of coral cover observed between 2003 and 2007 was caused by competition between these organisms. There was no negative effect on the recruitment of coral on artificial substrate free in reefs dominated by E. gabrieli or near their colonies. This suggests that a presumed reduction of coverage of this zoanthid should be accompanied by an increase in coral recruitment rates and the competition by inhibition of recruitment does not support an effect of hysteresis. It was found that this phase shift reduces species richness of reef fishes, by ten species unless reefs normal, and that favor mobile invertivores to the detriment of carnivores and sessile invertivores. However there was no difference in the abundance of fish.

Conselho Nacional de Desenvolvimento Científico e Tecnológico / Alguns recifes de coral da Baía de Todos os Santos passaram a ser dominado pelo zoantídeo Epizoanthus gabrieli em 2003. Fenômeno resultante da degradação dos recife de coral que atinge 20% desses ecossistemas e ameaça outros 35% no mundo. Apesar de sua importância, apenas a mudança na comunidade para o domínio de macroalgas foi suficientemente estudado. Assim, torna-se urgente estudos sobres alterações envolvendo a dominância de outros organismos. Estes fornecem subsídios para produção de modelos funcionais que podem ajudar na tomada de decisão para o manejo destes ecossistemas tanto na prevenção destas alterações quanto na recuperação de suas comunidades. Os objetivos deste trabalho são (i) verificar se este fenômeno constitui uma mudança de fase, a partir da redução da abundância de corais e persistência da alta cobertura de E. gabrieli por pelo menos cinco anos, (ii) avaliar os efeitos da competição entre este zoantídeo e corais adultos e recrutas com experimentos manipulativos e (iii) investigar os efeitos desta dominância na assembleia de peixes recifais. Os resultados confirmaram a existência de uma mudança de fase, sugerindo que a abundância de E. gabrieli aumentou em 2003 ou antes e que até 2007 houve uma redução da cobertura de coral, condição que se mantêm pelo menos até 2013. As três espécies de corais testadas mostram-se muito sensíveis ao contato com E. gabrieli, com necrose em 78% das colônias e ocupação do esqueleto dos corais em 35% dos casos em um período de 118 dias. Além disso, um modelo feito a partir dos dados de proporção de colônias de corais em contato com este zoantídeo e a cobertura de E. gabrieli sugere que quando o zoantídeo atinge 6% de cobertura, 50% das colônias de corais entram em contato com o mesmo. Estes dados são fortes evidências de que a redução da cobertura de coral observada entre 2003 e 2007 foi causada por competição entre estes organismos. Não foi observado efeito negativo no recrutamento do coral em substrato artificial livre em recifes dominados por E. gabrieli, nem nas proximidades das suas colônias. Isso sugere que uma suposta redução da cobertura deste zoantídeo deve ser acompanhada pelo aumento da taxa de recrutamento de corais e que a competição com a inibição do recrutamento não suporta um efeito de histerese. Foi constatado que esta mudança de fase reduz a riqueza de espécies de peixes recifais, apresentando dez espécies a menos que os recifes normais, e que favorecem os invertívoros moveis em detrimento dos carnívoros e invertívoros sésseis. Contudo não se observou diferença na abundância de peixes. / Some coral reefs of Todos os Santos Bay have undergone a drastic change in the benthic community. In 2003, they began to be dominated by the zoanthid Epizoanthus gabrieli. This phenomenon is a consequence of coral reef degradation that now reaches approximately 20% of coral reefs worldwide and and threatens another 35%. The most dramatic consequences of this degradation are the phase shift phenomena which are characterized by reduction of coral abundance and increase of organisms that do not build reefs. Despite their importance, only a phase shift to the domain of macroalgae has been sufficiently studied. Given this situation, it is urgent to create functional models that demonstrate the ecological processes that occur in the installation of alternative states that persist over time. These models provide important information that can help in decision making on management measures both for prevention and recovery of these ecosystems. The aims of this work are (i) to verify if this phenomenon is a phase shift, (i.e. the reduction of the abundance of coral and persistence of high coverage E. gabrieli for more than five years of monitoring), (ii) evaluate the effects of competition between this zoanthid and recruits and adult coral with manipulative experiments and (iii) study the effects of dominance on the fish assemblage. The result confirmed the existence of a phase shift, suggesting that the abundance of E. gabrieli increased in 2003 or before and in 2007 there was a reduction in coral cover, and that this condition remained until at least 2013. The three coral species tested show great sensitivity to contact with E. gabrieli, with necrosis in 78% of the colonies and with overgrow in 35% of cases in a period of 118 days. In addition, a model made from the data of a proportion of coral colonies in contact with this zoanthid and coverage of E. gabrieli suggests that when the zoanthid reaches 6% coverage, 50% of coral colonies come in contact with it. These data are strong evidence that the reduction of coral cover observed between 2003 and 2007 was caused by competition between these organisms. There was no negative effect on the recruitment of coral on artificial substrate free in reefs dominated by E. gabrieli or near their colonies. This suggests that a presumed reduction of coverage of this zoanthid should be accompanied by an increase in coral recruitment rates and the competition by inhibition of recruitment does not support an effect of hysteresis. It was found that this phase shift reduces species richness of reef fishes, by ten species unless reefs normal, and that favor mobile invertivores to the detriment of carnivores and sessile invertivores. However there was no difference in the abundance of fish.

The historic Everglades Ridge and Slough landscape maintained regularly spaced and elevated sawgrass ridges interspersed among exposed deeper-water sloughs; however, widespread but irregular hydrologic modification has degraded much of this landscape patterning. My study assessed the effects of hydrologic modification on vegetation community distinctness within the Ridge and Slough landscape through sampling species composition at fine-scales along a hydrologic gradient to measure the magnitude of segregation of species among patch types. The results show that vegetation community and topographic variation degradation is widespread, with distinctness differences proceeding and possibly being driven by topographic variation loss. Vegetation responses to past hydrologic regime modifications are likely affected by temporal lags; however, vegetation distinctness regeneration may also be hindered by a vegetatively homogeneous alternative stable state. Hydrologic regime restoration is critical for Ridge and Slough patterned landscape reestablishment, but management targets are complicated by vegetation response lags and possibly alternative stable states.

No description available.

Abstract The negative impact of invasive plant species on biodiversity and ecosystem functions, such as productivity and nutrient cycling has been deemed a global epidemic. To address this worldwide concern, information is needed on how the invasion process happens and how to control an existing invasion. The main aim of the research presented in this thesis was to develop a better understanding of the interacting role different mechanisms play in facilitating invasion and then link this understanding to the design of more effective control strategies. This aim is significant because traditional weed control strategies are not working. The estimated cost of controlling weeds in Australia is $1.4 billion per year in agricultural landscapes. Despite this substantial investment, invasive weed species are estimated to continue to cost the agricultural industry $2.2 billion per year in loss of yield. Current control strategies tend to focus on killing or removing an invasive plant species directly with the application of herbicides and/or mechanical removal. These strategies have proven ineffectual because the plant communities that assemble after management often remain dominated by the same invader or another. In this thesis, I use a combination of empirical and modelling techniques to investigate how disturbance regimes and competitive interactions between invasive plants and native plants can be manipulated to improve the efficacy of restoration efforts. To do this, I use the model scenario of the invasion of Eragrostis curvula (African lovegrass), an invasive grass species introduced into Australia in the early 1900s from South Africa. This species has now spread into every Australian state and territory (chapter 2). I specifically focus on two mechanisms: (1) disturbance, i.e. cattle grazing, and (2) competitive interactions. In chapter 3, I examine connections between dominance and competitive differences among African lovegrass and several functionally similar native grass species in a pasture community. To test the displacement hypothesis, I used a glasshouse competition trial to investigate interactions between African lovegrass and two non-persisting native grass species (Themeda australis and Bothriochloa decipiens) with manipulations of resources, neighbour density, and establishment order. To test the partitioning hypothesis, I compared in situ water use patterns among African lovegrass and two coexisting native grass species (Aristida calycina and Aristida personata) based on the assumption that water is the most limiting resource in this system. The key finding of this chapter is that competition can have important, but contingent, impacts on dominance. Competitive differences appear to partially contribute to abundance patterns after establishment, but may be relatively unimportant during the establishment phase where disturbance appears more critical. In chapter 4, I provide evidence that the identification of mechanisms that led to an invasion, while crucial for the development of effective preventative measures and understanding the invasion process, may not be necessary for the design of more effective control strategies. To examine the effects of different control strategies on African lovegrass and the resultant community, I established a large factorial field-trial with a split-plot design. I manipulated grazing, soil nutrient levels and the presence of the invader. The most common control strategy (removing the causal disturbance and killing the invasive grass), based implicitly on traditional equilibrium models, was not an effective option for restoring a desirable native community. Instead, this strategy led to the dominance of a secondary invader. The most effective control strategy was based on alternative stable states models and involved maintaining grazing, and increasing the palatability of the invader with fertilizers. The key finding of this chapter is that novel approaches for control, which consider the dynamics of the invader-dominated system, are needed. In chapter 5, I investigate the benefits of explicitly incorporating actions that manipulate disturbance (natural or imposed) into control efforts. To do this, I first developed a process model that described the dynamics of an invader whose establishment is preferentially favoured by disturbance. I then couched this model in a decision theory framework, a stochastic dynamic program, and applied a case-study of another invasive plant species, Mimosa pigra (a perennial legume shrub and pan-tropical weed). The key finding of this chapter is that strategies should not only focus on existing invader-dominated sites, but should also protect sites occupied by native species from disturbances that facilitate invasion. The research discussed in this thesis makes three key contributions to a better understanding of the invasion process and the design of more effective control strategies: 1) the search for one key mechanism is not sufficient because multiple mechanisms can interact or shift in importance to facilitate different stages of invasion, 2) a novel approach is needed to restore a more desirable native community because the dynamics of the invader-dominated system can differ from the historical native community, and 3) control efforts should be broadened in focus to include protection of the integrity of native communities from disturbances that facilitate invasion.