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Incremento diamétrico arbóreo em diferentes grupos funcionais e produção de serapilheira em duas florestas tropicais brasileiras / Tree diametric increment in different functional groups and litterfall production in two Brazilian tropical forestsMaurício Lamano Ferreira 08 August 2014 (has links)
A alocação de C em biomassa aérea é uma importante via do fluxo de elemento em ecossistemas terrestres. Uma forma de monitorar este processo é por meio de bandas dendrométricas que subsidiam informações relevantes sobre o perfil de incremento diamétrico arbóreo ao longo do tempo. Porém, este monitoramento se torna mais eficientes quando realizados sob a ótica de grupos funcionais, uma vez que florestas tropicais apresentam alta diversidade biológica. No entanto, deve-se considerar que a perda de carbono para o ambiente, seja em termos de respiração autotrófica, mortalidade ou produção de serapilheira, também é um atributo importante na dinâmica de florestas tropicais. Desta forma, torna-se crucial o monitoramento da entrada e saída de C em ecossistemas terrestres tropicais. O objetivo deste trabalho foi avaliar o perfil de incremento diamétrico arbóreo (IDA) e a produção de serapilheira em duas florestas tropicais brasileiras. Primeiramente instalaram-se 850 bandas dendrométricas em árvores de uma comunidade da floresta Amazônica (FAMZ). Na floresta Atlântica (FAT) foram instaladas 1532 bandas em árvores localizadas em três fitofisionomias ao longo de um gradiente altitudinal (100, 400 e 1000 metros de altitude). Para a estimativa da serapilheira 64 coletores foram espalhados aleatoriamente numa área de 20 ha da FAMZ e 40 coletores por nível de altitude foram espalhados na FAT. Na floresta Atlântica as árvores localizadas no meio do gradiente (400 m) foram as que apresentaram maior incremento diamétrico arbóreo anual, seguido pela comunidade a 100 m e por fim a comunidade localizada na maior altitude. As árvores apresentaram redução de incremento diamétrico arbóreo durante a estação de menor índice pluviométrico e menor temperatura média do ar. Na floresta Amazônica as árvores amostradas apresentaram menor taxa de incremento arbóreo durante a estação seca, que também é a mais quente. Em geral, temperatura e precipitação foram as variáveis ambientais que apresentaram grande influência no perfil de incremento diamétrico arbóreo, sendo que na FAT a temperatura atuou positivamente na taxa de incremento diamétrico arbóreo dos indivíduos amostrados a 400 e 1000 metros de altitude e na FAMZ esta variável interferiu negativamente no desenvolvimento das árvores. Em ambas as florestas a precipitação foi correlacionada positivamente com o perfil de crescimento das árvores. A análise de ordenação mostrou que em ambas as florestas houve um forte agrupamento das árvores dentro dos subgrupos funcionais, destacando assim sua a eficiência na análise de respostas ambientais em ecossistemas tropicais. A produção de serapilheira foi 20% maior na FAMZ do que na FAT. A principal fração representada na serapilheira total foi a folha. Novamente a temperatura foi uma variável ambiental importante no aporte do material decíduo. Estes resultados trazem importantes contribuições sobre o uso de grupos funcionais em estudos que visem a alocação de biomassa em florestas tropcais, além de trazer uma comparação entre a dinâmica de crescimento arbóreo nas duas maiores florestas tropicais brasileiras / Allocation of C in aboveground biomass is an importantpathway of the element flow in terrestrial ecosystems. One way to monitor this process is through dendrometric bands that support relevant information about tree diametric increment over time. However, this monitoring becomes more efficient when performed in functional groups, since tropical forests have high biodiversity levels. However, one must consider that the loss of carbon to the atmosphere, either in terms of autotrophic respiration, mortality or litter production, is also an important attribute in the dynamics of tropical forests. Thus, it is crucial to monitor the incoming and outgoing of C in tropical terrestrial ecosystems. The goal of this study was to evaluate the tree diametric increment rate and litterfall in two Brazilian tropical forests. Firstly,850 dendrometric bands ware installed on trees of a community in the Amazon rainforest (FAMZ). In the Atlantic forest (FAT) 1532 bandswere installedon trees located in three vegetation types along an altitudinal gradient (100, 400 and 1000 ma.s.l.). For litterfall, 64 collectors were placed randomly over an area of 20 ha in FAMZ and 40 collectorswere scattered per level altitude in the FAT. In the Atlantic forest trees located in the middle of the gradient (400 m a. s. l.) showed the greatest annual diametric increase, followed by trees located at 100 m (a.s.l.) and finally trees located at 1000 m (a.s.l.). During the dry season and lower mean airtemperature trees showed slow growth. In the Amazon forest the sampled trees had slower growth rate during the dry season, which was the hottest season. In general, temperature and precipitation were the environmental variables that had great influence on the tree diametric growth. The air temperature acted positively on the growth rate of individuals sampled at 400 and 1000 meters altitude in the FAT and in the FAMZ this variable showed negative effect on tree growth. In both forests precipitation was positively correlated with the tree growth rate. Ordination analysis showed that in both forests there was a strong grouping of trees within the functional subgroups, thus highlighting its efficiency in the analysis of environmental responses in tropical ecosystems. Litterfall production was 20% higher in FAMZ than in the FAT. The main fraction of total litterfall was leave. Again the temperature was an important environmental variable in the production of the deciduous material. These results provide important contributions on the use of functional groups in studies seeking at biomass allocation in tropcais forests, besides bringing a comparison between the dynamics of tree growth in the two largest Brazilian rainforests
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Diversidade e desenvolvimento de girinos (Amphibia, Anura) em corpos d’água associados a fragmentos de mata no nordeste do Brasil / Tadpole (Amphibia, Anura) diversity in water bodies associated to forest fragments in Sao Luis Island, MA, BrazilOliveira, Daniel Brito de 12 July 2010 (has links)
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Previous issue date: 2010-07-12 / Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES) / Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPQ) / Urbanization brings many consequences such as fragmentation and degradation of the remaining forest fragments. Degradation in these fragments can negatively affect the forest species, but favor their use by open area species. Thus we aimed to investigate if tadpoles of open area species are developing, together with those of forest species, in water bodies located in forest fragments in Sao Luis Island, MA, Brazil. For this we collected tadpoles in 32 pools with varied characteristics divided into three different types of environment (open area, outer edge of the forest, and forest interior), during the years 2008 and 2009. We recorded 20 species in five families (Bufonidae, Hylidae, Leptodactylidae, Leiuperidae, Microhylidae). The environmental variables accounted for much of the variation in the abundance of tadpoles, and the percentage of vegetation in the pool, the vegetation height in the forest edge, the distance to the forest, and the depth of the pool showed significative effects. Tadpoles of open area species occured in water bodies located in the forest interior. The increased presence of generalist species, and the reproductive success of open area species in the forests indicate that these areas are impacted. / A urbanização traz diversas conseqüências como a fragmentação e a degradação dos remanescentes florestais. A degradação nesses fragmentos pode afetar negativamente as espécies de mata, mas favorecer a sua utilização por espécies de áreas abertas. Assim, objetivamos investigar se girinos de espécies de áreas abertas estão se desenvolvendo, junto com os de espécies de mata, nos corpos d’água localizados em fragmentos de mata na ilha de São Luis, MA, Brasil. Para isso coletamos girinos em 32 poças de características variadas distribuídas em três tipos de ambiente (área aberta, borda externa de mata e interior de mata), durante os anos de 2008 e 2009. Registramos 20 espécies distribuídas em 5 famílias (Bufonidae, Hylidae, Leptodactylidae, Leiuperidae, Microhylidae). As variáveis ambientais que responderam por grande parte da variação nas abundâncias de girinos foram: porcentagem de vegetação no corpo d’água, a altura da vegetação na borda da mata, a distancia para a mata e a profundidade do corpo d’água. Os girinos de espécies de área aberta ocorreram nos corpos d’água localizados na mata. Além disso a maior presença de espécies generalistas e o sucesso reprodutivo das espécies de área aberta no interior das matas indicaram que estas áreas estão impactadas.
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Estimation of biomass for calculating carbon storage and CO2 sequestration using remote sensing technology in Yok Don National Park, Central Highlands of Vietnam / Ước lượng sinh khối cho tính toán lượng tích trữ các bon và hấp thụ CO2 ở Vườn Quốc gia Yok Đôn, Tây Nguyên Việt Nam, bằng cách sử dụng công nghệ viễn thámNguyen, Viet Luong 15 November 2012 (has links) (PDF)
Global warming and climate change are closely related to the amount of CO2 in the air. Forest ecosystem plays very important role in the global carbon cycle; CO2 from the atmosphere is taken up by vegetation and stored as plant biomass. Therefore, quantifying biomass and carbon sequestration in tropical forests has a significant concern within the United Nations Framework Convention on Climate Change (UNFCC), Kyoto Protocol and Reducing Emission from Deforestation and Forest Degradation (REDD) program for the purpose of the improvement of national carbon accounting as well as for addressing the potential areas for carbon credits, basis for payment for environmental services. The aim of research is to estimate biomass and carbon stocks in tropical forests using remote sensing data for dry forest of central highlands of Vietnam. This result showed that from satellite images of the SPOT, satellites could build the land cover map, carbon map and biomass map of Yok Don National Park, Central Highlands of Vietnam. Through which also the biomass (above ground biomass and below ground biomass) of each type of forest can be calculated. For instance the biomass of the dry forest (Dry Dipterocarp Forest) is 153.49 tones x ha-1, biomass of rich forest is 343.35 tones x ha-1, biomass of medium forest is 210.34 tones x ha-1 and biomass of poor forest & scrub are 33.56 tones x ha-1. / Sự ấm lên toàn cầu và biến đổi khí hậu có liên quan chặt chẽ với tổng lượng CO2 trong không khí. Hệ sinh thái rừng có vai trò rất quan trọng trong chu trình các bon toàn cầu; khí CO2 trong khí quyển được hấp thụ bởi thảm thực vật dưới dạng sinh khối. Vì vậy, việc xác định sinh khối và carbon tích trữ trong rừng nhiệt đới đã có được sự quan tâm đáng kể trong Công ước của Liên hiệp quốc về biến đổi khí hậu (UNFCC), Nghị định thư Kyoto và Chương trình giảm phát thải từ phá rừng và suy thoái rừng (REDD) gần đây, nhằm cho mục đích cải thiện việc tính toán lượng các bon tích trữ cũng như giải quyết các vấn đề tiềm năng cho tín dụng các bon, làm cơ sở cho việc thanh toán cho các dịch vụ môi trường. Mục đích của nghiên cứu này là ước lượng sinh khối và các bon lưu trữ trong các khu rừng nhiệt đới bằng cách sử dụng dữ liệu viễn thám, mà ở nghiên cứu này là cho rừng khộp Tây Nguyên của Việt Nam. Kết quả cho thấy rằng, từ ảnh vệ tinh SPOT có thể xây dựng bản đồ lớp phủ thực vật, bản đồ các bon và bản độ sinh khối của Vườn quốc gia Yok Đôn, Tây Nguyên Việt Nam. Qua đó đã tính toán được sinh khối (bao gồm cả trên mặt đất và dưới mặt đất) như: đối với sinh khối của rừng khô cây họ dầu (Dry Dipterocarp Forest) là 153,59 tấn/ha, sinh khối rừng giàu là 343,35 tấn/ha, sinh khối rừng trung bình là 210,34 tấn/ha và sinh khối rừng nghèo&cây bụi là 33,56 tấn/ha.
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Road Impact on Deforestation and Jaguar Habitat Loss in the Mayan ForestConde Ovando, Dalia Amor 25 July 2008 (has links)
<p>The construction of roads, either as an economic tool or as necessity for the implementation of other infrastructure projects is increasing in the tropical forest worldwide. However, roads are one of the main deforestation drivers in the tropics. In this study we analyzed the impact of road investments on both deforestation and jaguar habitat loss, in the Mayan Forest. As well we used these results to forecast the impact of two road investments planned in the region. Our results show that roads are the single deforestation driver in low developed areas, whether many other drivers play and important role in high developed areas. In the short term, the impact of a road in a low developed area is lower than in a road in a high developed area, which could be the result of the lag effect between road construction and forest colonization. This is consistent since roads resulted to be a significant deforestation driver for at least two decades. Roads significantly affect jaguar's habitat selection; however males showed a higher tolerance than females. From 1980 to 2000 female jaguars lost 36% of their habitat wile males lost 22%. Our forecasting of the impact of the proposed road, shows that it will promote the deforestation of approximately 16,851 has, and the jaguar habitat loss of 146,929, during the first decade; meanwhile the alternative route will have and impact of 2519 hectares and the habitat loss of 899 hectares.</p> / Dissertation
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Why do Birds Migrate? The Role of Food, Habitat, Predation, and CompetitionBoyle, Alice January 2006 (has links)
The ultimate causes of bird migration are largely unknown despite more than a century of research. By studying partially migratory short-distance tropical migrants and by employing comparative methods, some difficulties in testing hypotheses for evolution of migration can be overcome. Using comparative methods I tested the evolutionary precursor hypothesis, a major hypothesis for why migration evolved in some lineages and not in others. The results of this study conflicted with many assumptions and predictions of the evolutionary precursor hypothesis. Most importantly, migratory behavior was not related to diet and habitat in simple ways. The interaction between diet and habitat, as well as consistent associations between flocking behavior and migration suggested that food variability is poorly captured by the surrogates embodied in the evolutionary precursor hypothesis. I then employed comparative methods to studying tropical altitudinal migration. Comparisons of diets and fruit preferences between species pairs showed that migrants are more frugivorous, eat a broader diversity of fruits, and have diets that more strongly resemble their preferences than do residents. Although providing evidence that food limitation plays a role in altitudinal migration, these results do not support the hypothesis that interspecific competition explains variation in migratory behavior. Next, I provided the first test of a predation-based hypothesis to explain altitudinal migration. Migrants breed at higher elevations than where they spend their non-breeding season. Thus, birds may migrate uphill to escape high nest predation risk at lower elevations. Results from this experimental study are largely consistent with this hypothesis, but anomalies between predicted and observed patterns suggest that either migration of lowland birds occurs in response to other factors, or that anthropogenic change has altered the tradeoffs involved in migratory decisions. Finally, I focus on a single migrant species and evaluate (a) two food-based hypotheses to explain the destination of migration movements, and (b) mechanisms underlying intra-specific differences in migratory strategy. Food can explain why Corapipo altera migrate uphill, but not why they migrate downhill. My data on sex bias and body condition leads to a new hypothesis explaining the complete annual cycle of this tropical migrant bird.
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Interspecific and Size-dependent Variation in Carbon Concentration and Wood Chemical Traits of Tropical TreesMartin, Adam 17 December 2012 (has links)
Tropical forests play a major role in global carbon (C) dynamics and maintain some of the highest biological complexity on Earth; however, little is known about how variation in wood chemical traits contributes to tropical forest structure and function. This research examines inter- and intraspecific patterns in wood chemical traits in order to understand 1) the role wood chemical traits play in tropical forest C dynamics, and 2) the adaptive significance of wood chemical traits in tropical trees. I found wood C concentration varies widely among co-occurring tropical tree species, with average C concentration (47.4 ± 0.33% w/w (S.E.)) being significantly lower than values assumed in prominent forest C accounting protocols. Failing to account for this variation leads to overestimates of ~3.3 – 5.3% in tropical forest C accounting, an error that compounds significantly at larger spatial scales. I also show that oven drying samples prior to elemental analysis underestimates wood C concentration by 2.5 ± 0.17%, due to the loss of the “volatile C fraction”. Counter to expectations, I found wood C concentration is not
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phylogenetically conserved nor correlated to species demography or life history traits. Wood chemical traits showed consistent size-dependent patterns: wood C (in 16 of 24 species) and lignin (in 15 of 16 species) was higher in saplings vs. conspecific canopy trees. These patterns, complimented by phylogenetic analyses, suggest saplings require wood chemical traits that confer greater pathogen defense. When analyzed across a continuous size spectrum, I found wood C concentration (and leaf structural traits) increases linearly, while wood starch concentration (and leaf traits associated with C gain) shows “hump-shaped” patterns with peak values closely preceding reproductive onset; the latter result suggests C may limit growth in larger trees. Overall, my dissertation provides one of the first comprehensive examinations of wood chemical trait variation in tropical trees. In doing so it provides novel, timely, and critical insights into how wood chemical traits contribute to tropical forest structure and function.
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Interspecific and Size-dependent Variation in Carbon Concentration and Wood Chemical Traits of Tropical TreesMartin, Adam 17 December 2012 (has links)
Tropical forests play a major role in global carbon (C) dynamics and maintain some of the highest biological complexity on Earth; however, little is known about how variation in wood chemical traits contributes to tropical forest structure and function. This research examines inter- and intraspecific patterns in wood chemical traits in order to understand 1) the role wood chemical traits play in tropical forest C dynamics, and 2) the adaptive significance of wood chemical traits in tropical trees. I found wood C concentration varies widely among co-occurring tropical tree species, with average C concentration (47.4 ± 0.33% w/w (S.E.)) being significantly lower than values assumed in prominent forest C accounting protocols. Failing to account for this variation leads to overestimates of ~3.3 – 5.3% in tropical forest C accounting, an error that compounds significantly at larger spatial scales. I also show that oven drying samples prior to elemental analysis underestimates wood C concentration by 2.5 ± 0.17%, due to the loss of the “volatile C fraction”. Counter to expectations, I found wood C concentration is not
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phylogenetically conserved nor correlated to species demography or life history traits. Wood chemical traits showed consistent size-dependent patterns: wood C (in 16 of 24 species) and lignin (in 15 of 16 species) was higher in saplings vs. conspecific canopy trees. These patterns, complimented by phylogenetic analyses, suggest saplings require wood chemical traits that confer greater pathogen defense. When analyzed across a continuous size spectrum, I found wood C concentration (and leaf structural traits) increases linearly, while wood starch concentration (and leaf traits associated with C gain) shows “hump-shaped” patterns with peak values closely preceding reproductive onset; the latter result suggests C may limit growth in larger trees. Overall, my dissertation provides one of the first comprehensive examinations of wood chemical trait variation in tropical trees. In doing so it provides novel, timely, and critical insights into how wood chemical traits contribute to tropical forest structure and function.
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Radiação solar global e fotossinteticamente ativa (PAR) acima e abaixo do dossel de floresta de mata atlântica no estado de Alagoas / Global solar and photosynthetically active radiation (PAR) above and below the canopy of atlantic Forest in Alagoas stateAndrade, Antonio Marcos Delfino de 18 March 2011 (has links)
Conselho Nacional de Desenvolvimento Científico e Tecnológico / A Mata Atlântica é um dos mais ricos ecossistemas em termos de diversidade biológica do mundo. Ela é representada por vários tipos de vegetações, além de ser caracterizada pelo alto grau de endemismo. Atualmente, esta floresta, se encontra bastante fragmentada, restando apenas 7,5%. Apesar disso, ainda mantém níveis extremamente altos de biodiversidade e endemismo. Contudo, este bioma, provavelmente, é o mais devastado e seriamente ameaçado do planeta. Apesar da importância do estudo sobre Mata Atlântica, pouco se conhece dos processos físicos, químicos e biológicos existentes acima e abaixo do dossel. O estudo da radiação solar é importante para entender os diversos processos físicos, químicos e biológicos que ocorrem na biosfera, em particular em floresta. Ainda no ambiente de floresta a radiação solar se destaca entre outros fatores, sendo fundamental nos processos de fotossíntese, aquecimento do ar e da superfície e evapotranspiração. O processo de fotossíntese ocorre apenas em parte do espectro da Radiação Solar, compreendido entre os comprimentos de onda eletromagnética de 0,4 a 0,7 μm e denominada de Radiação Fotossinteticamente Ativa (PAR). Com base no exposto, o objetivo é caracterizar a evolução temporal da Radiação Solar Global (Rg) e Fotossinteticamente Ativa acima e abaixo de um fragmento remanescente de Mata Atlântica no Estado de Alagoas. O estudo foi realizado, no período de outubro/2009 a setembro/2010, com a obtenção dos dados (Rg, PAR, PAR refletida e precipitação) provenientes de uma torre micrometeorológica (10º17 36 S, 36º17 24 W) de 26 metros instalada numa área de floresta de Mata Atlântica. A área de estudo está localizada na Reserva Particular do Patrimônio Natural (RPPN) Lula Lobo I inserida na fazenda Capiatã A, pertencente à Usina Coruripe Açúcar e Álcool S/A, no município de Coruripe, Alagoas. Verificou-se que a radiação solar incidente (Radiação Solar Global Externa (Rg_Ext) e Radiação Fotossinteticamente Ativa na parte Externa (PAR_Ext)) seguem a sazonalidade imposta pelo movimento aparente do Sol. Na época seca observou-se os maiores registros de Rg_Ext e PAR_Ext, com respectivas médias superiores a 900 W m-2 e 600 W m-2. Enquanto que, na época chuvosa foram registradas as menores médias, com Rg_Ext abaixo de 500 W m-2 e PAR_Ext inferior a 300 W m-2. Entretanto, uma pequena fração da radiação solar atinge o solo da floresta, ou seja, menos de 8% incide sobre a superfície. A radiação solar que incide no chão da floresta tem grande dependência do ângulo zenital (Z) e da estrutura do dossel. Notou-se também grande influência do Z na Rg_Ext e PAR_Ext, onde se verificou que quanto maior Z, menores são os valores destas componentes, em razão do maior caminho ótico que os raios solares tem que percorrer para atingir à superfície.
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A Performance Guide for Heitor Villa-Lobos's Quatro Canções Da Floresta Do AmazonasJanuary 2017 (has links)
abstract: This paper is a performance guide for Quatro Canções da Floresta do Amazonas [Four Songs of The Amazon Forest] by Brazil's most prolific composer, Heitor Villa-Lobos. The primary purpose of the paper is to serve as a source for the correct pronunciation of the Brazilian Portuguese language of the songs. It will begin with an overview of Heitor Villa-Lobos's life and career, showing how his compositions catalyzed the Nationalistic movement in Brazilian classical music. His inclusion of native and folk elements into classical compositions was a significant innovation, which places Villa-Lobos as one of the most important Brazilian classical composers. Furthermore, this paper will explore the issue of Brazilian Portuguese diction in depth, using the Quatro Canções da Floresta do Amazonas to aid non-native Brazilian speakers. This includes an International Phonetic Alphabet transcription of the songs, as well as a recording of the songs being read and sung by the author, a link to which can be found in the appendix. / Dissertation/Thesis / Doctoral Dissertation Music 2017
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Padrões fenológicos de espécies arbóreas em um fragmento de floresta semidecídua no estado de São Paulo: definição de padrões sazonais e comparação entre anosRubim, Paulo [UNESP] 14 June 2006 (has links) (PDF)
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rubim_p_me_rcla.pdf: 840416 bytes, checksum: 477d55c54031309c90dd351bc26b73ab (MD5) / Fundação de Amparo à Pesquisa do Estado de São Paulo (FAPESP) / Florestas tropicais sob clima com estações seca e chuvosa definidas tendem a apresentar padrões fenológicos adaptados ao regime sazonal das condições ambientais. Para descrever e entender os padrees fenológicos vegetativos e reprodutivos de uma floresta semidecídua, 90 espécies arbóreas foram monitoradas durante quatro anos tentando responder as seguintes perguntas: (i) Existe sazonalidade na fenologia da floresta estudada? O padrão encontrado se repete durante os anos de estudo? (ii) Quais fatores climáticos médios ou atuais estão relacionados a cada fenofase observada? (iii) O padrão de frutificação está relacionado ao modo de dispersão das sementes? Foi utilizada estatística circular para testar a sazonalidade dos eventos fenológicos e correlações para testar as influências do clima médio (30 anos) e atual (período de quatro anos) no padrão fenológico entre os anos (V= 48 meses) e em cada ano (jV= 12 meses). Queda e brotamento foliar apresentaram padrões fortemente sazonais. sendo que a queda foliar ocorreu durante a estação seca e o brotamento na transição seca- chuvosa durante os quatro anos de estudo. Antese não apresentou padrão significativarnente sazonal. porém apresentou maior atividade durante toda estação chuvosa. A frutificação (frutos maduros) de espécies anemocóricas foi fortemente sazonal, ocorrendo durante a estação seca, enquanto espécies dispersas por animais e autocóricas não apresentaram padrões significativarnente sazonais, frutificando ao longo do ano todo. Queda foliar. antese, frutos maduros e frutos anernocóricos apresentaram correlaçàes significativas mais fortes com as médias climáticas do que com as variações climáticas do período de estudo. A fenologia das espécies de floresta semidecídua não diferiu significativamente entre os quatro anos de estudo. sugerindo mm padrão temporal previsível... / Tropical forests growing under a seasonal climate with a marked dry and wet season tend te adapt the plant phenotogy to the euvronmental seasonaIty. Tu desenhe and uuclerstand the reproductive and vegetative phenological patterns of a tropical sernideciduous forest we monitored 90 tree species over feia years aim te answer the following questions: (i) Are the reproductive and vegetative phenological patterns seasonal? Are the observed patterns repeated over the studied years? (ii) Which climatie factor. Inean or in the period, is related to the observed phenophases? (iii) Does the fruiting patterns vary among diíferent seed dispersal modes? We applied circular statistics analyses to test for the phenology seasonality and spearman correlation to test for the influence ofrnean climate (30 years) and actual ciimate (four years study period) on the phenological patterns over the years (N = 48) and ou each year (N = 12). Leaf fali and ieaf flushing were highiy seasonal, leaf fali occurring during the dry season and Ieaf flushing in the dry lo wet season transition during the four years of study. Flowering (anthesis) did not show a signiflcantiy seasonal pattern. although more species flowered during the wet season. Fruiting (ripe fruits) of wind dispersed species were highly seasonal. occurring in the drv season. whereas ftuiting patterns of animal and self dispersed fruits were nol seasonal. occurring ali year round. Leaf fali, flowering, ripe fruiting. and wind-dispersed fruiting presented significant correlations with climatic factors, and the correlations were stronger with mean ciimatic factor than with actual climatic factors. Phenoiogical pattems did not differ among years in the sernideciduous forest. suggesting a predictable pattern. at the sarne time every year. and siapporting rnost of the seasonal patterns described in short term studies.
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