Appendices removed due to copyright restrictions Appendix 1: Beausoleil, N J, Stafford, KJ, Mellor, DJ. 2005. Sheep show more aversion to a dog than to a human in an arena test. Applied Animal Behaviour Science, 91 : 2 1 9-232. Appendix 2: Beausoleil, NJ, Stafford, KJ, MelIor, DJ. Does direct human eye contact function as a warning cue for domestic sheep, Ovis aries? Journal of Comparative Psychology, J 20 (3) : 269-279. Appendix 3:Beausoleil, NJ, Stafford, KJ, MelIor, DJ. 2004. Can we use change in core body temperature to evaluate stress in sheep? Proceedings of the New Zealand Society of Animal Production, 64: 72-76. Appendix 4: Beausoleil, N.J, Mellor, DJ, Stafford, KJ 2004. Methods for marking New Zealand wildlife: amphibians, reptiles and marine mammals. Department of Conservation, Wellington, New Zealand 147p. ISBN 0-478-2263 1-4. Appendix 5: Mellor, DJ, Beausoleil, NJ, Stafford, KJ 2004. Marking amphibians, reptiles and marine mammals: Animal welfare, practicalities and public perceptions in New Zealand. Miscellaneous Publication, Department of Conservation, Wellington, New Zealand 55p. ISBN 0-478-22563-6. / Both humans and dogs are integral in sheep production systems; however, which is more aversive to sheep, or indeed, whether either causes significant stress, has not been shown experimentally. The aim of this thesis was to examine some behavioural and physiological responses of domestic sheep to the presence of humans or dogs. An arena test was used to measure the relative aversion of sheep to the presence of a human or dog, as well as to elucidate differences in the responses of flocks at the University of Western Australia (UWA) which were putatively selected for differences in fearfulness. A Y maze preference test was used to 'ask' sheep whether they preferred a human shaking a rattle or a barking dog. In both tests, adrenocortical responses were measured concurrently to support the interpretation of behaviour. The presence of a human or dog in the arena elicited significantly more avoidance and vigilance behaviour and less exploration than did the presence of a control object. However, the dog elicited significantly more of this fear-related behaviour, and significantly larger adrenocortical responses than did the human. Sheep also expressed a clear preference for a human shaking a rattle over a barking dog in the Y maze test and exhibited larger adrenocortical responses to the dog than to the human in the Y maze facility. The UWA flocks differed in their expression of locomotor and vocal activity; MA sheep were more active/vocal than the other flocks, not only in the presence of the human but also with the box or dog. MA sheep expressed less avoidance and vigilance and more exploration than the other flocks in the presence of the human and exhibited significantly lower plasma cortisol concentrations than LA sheep after exposure to the human (10-min sample). However, there were no inter-flock differences in fear-related behaviour or adrenocortical responses when the flocks were presented with the box or dog. The results do not support the notion that the UWA flocks have been selected for differences in a consistent predisposition to react fearfully. The adrenocortical responses measured in these studies were only moderate in magnitude and duration, with peak plasma cortisol concentrations 2-3 times higher than pre-treatment values, and all concentrations returning to pre-treatment levels within one hour of the start of treatment. If these observations are confirmed in practical situations, the presence of humans and dogs during routine handling should cause little concern on the basis of animal welfare. However, limiting the presence of dogs in certain situations (e.g. before slaughter) may reduce stress in domestic sheep. Significant methodological developments in this research include the use of multivariate statistical techniques to analyze arena behaviour, the concurrent measurement of adrenocortical and behavioural responses in the arena and Y maze tests, and the explicit testing of the effects of individual lateral biases on choice behaviour in a Y maze test. Future studies should measure sheep stress responses to the presence of humans and dogs in practical situations.

Appendices removed due to copyright restrictions Appendix 1: Beausoleil, N J, Stafford, KJ, Mellor, DJ. 2005. Sheep show more aversion to a dog than to a human in an arena test. Applied Animal Behaviour Science, 91 : 2 1 9-232. Appendix 2: Beausoleil, NJ, Stafford, KJ, MelIor, DJ. Does direct human eye contact function as a warning cue for domestic sheep, Ovis aries? Journal of Comparative Psychology, J 20 (3) : 269-279. Appendix 3:Beausoleil, NJ, Stafford, KJ, MelIor, DJ. 2004. Can we use change in core body temperature to evaluate stress in sheep? Proceedings of the New Zealand Society of Animal Production, 64: 72-76. Appendix 4: Beausoleil, N.J, Mellor, DJ, Stafford, KJ 2004. Methods for marking New Zealand wildlife: amphibians, reptiles and marine mammals. Department of Conservation, Wellington, New Zealand 147p. ISBN 0-478-2263 1-4. Appendix 5: Mellor, DJ, Beausoleil, NJ, Stafford, KJ 2004. Marking amphibians, reptiles and marine mammals: Animal welfare, practicalities and public perceptions in New Zealand. Miscellaneous Publication, Department of Conservation, Wellington, New Zealand 55p. ISBN 0-478-22563-6. / Both humans and dogs are integral in sheep production systems; however, which is more aversive to sheep, or indeed, whether either causes significant stress, has not been shown experimentally. The aim of this thesis was to examine some behavioural and physiological responses of domestic sheep to the presence of humans or dogs. An arena test was used to measure the relative aversion of sheep to the presence of a human or dog, as well as to elucidate differences in the responses of flocks at the University of Western Australia (UWA) which were putatively selected for differences in fearfulness. A Y maze preference test was used to 'ask' sheep whether they preferred a human shaking a rattle or a barking dog. In both tests, adrenocortical responses were measured concurrently to support the interpretation of behaviour. The presence of a human or dog in the arena elicited significantly more avoidance and vigilance behaviour and less exploration than did the presence of a control object. However, the dog elicited significantly more of this fear-related behaviour, and significantly larger adrenocortical responses than did the human. Sheep also expressed a clear preference for a human shaking a rattle over a barking dog in the Y maze test and exhibited larger adrenocortical responses to the dog than to the human in the Y maze facility. The UWA flocks differed in their expression of locomotor and vocal activity; MA sheep were more active/vocal than the other flocks, not only in the presence of the human but also with the box or dog. MA sheep expressed less avoidance and vigilance and more exploration than the other flocks in the presence of the human and exhibited significantly lower plasma cortisol concentrations than LA sheep after exposure to the human (10-min sample). However, there were no inter-flock differences in fear-related behaviour or adrenocortical responses when the flocks were presented with the box or dog. The results do not support the notion that the UWA flocks have been selected for differences in a consistent predisposition to react fearfully. The adrenocortical responses measured in these studies were only moderate in magnitude and duration, with peak plasma cortisol concentrations 2-3 times higher than pre-treatment values, and all concentrations returning to pre-treatment levels within one hour of the start of treatment. If these observations are confirmed in practical situations, the presence of humans and dogs during routine handling should cause little concern on the basis of animal welfare. However, limiting the presence of dogs in certain situations (e.g. before slaughter) may reduce stress in domestic sheep. Significant methodological developments in this research include the use of multivariate statistical techniques to analyze arena behaviour, the concurrent measurement of adrenocortical and behavioural responses in the arena and Y maze tests, and the explicit testing of the effects of individual lateral biases on choice behaviour in a Y maze test. Future studies should measure sheep stress responses to the presence of humans and dogs in practical situations.

Appendices removed due to copyright restrictions Appendix 1: Beausoleil, N J, Stafford, KJ, Mellor, DJ. 2005. Sheep show more aversion to a dog than to a human in an arena test. Applied Animal Behaviour Science, 91 : 2 1 9-232. Appendix 2: Beausoleil, NJ, Stafford, KJ, MelIor, DJ. Does direct human eye contact function as a warning cue for domestic sheep, Ovis aries? Journal of Comparative Psychology, J 20 (3) : 269-279. Appendix 3:Beausoleil, NJ, Stafford, KJ, MelIor, DJ. 2004. Can we use change in core body temperature to evaluate stress in sheep? Proceedings of the New Zealand Society of Animal Production, 64: 72-76. Appendix 4: Beausoleil, N.J, Mellor, DJ, Stafford, KJ 2004. Methods for marking New Zealand wildlife: amphibians, reptiles and marine mammals. Department of Conservation, Wellington, New Zealand 147p. ISBN 0-478-2263 1-4. Appendix 5: Mellor, DJ, Beausoleil, NJ, Stafford, KJ 2004. Marking amphibians, reptiles and marine mammals: Animal welfare, practicalities and public perceptions in New Zealand. Miscellaneous Publication, Department of Conservation, Wellington, New Zealand 55p. ISBN 0-478-22563-6. / Both humans and dogs are integral in sheep production systems; however, which is more aversive to sheep, or indeed, whether either causes significant stress, has not been shown experimentally. The aim of this thesis was to examine some behavioural and physiological responses of domestic sheep to the presence of humans or dogs. An arena test was used to measure the relative aversion of sheep to the presence of a human or dog, as well as to elucidate differences in the responses of flocks at the University of Western Australia (UWA) which were putatively selected for differences in fearfulness. A Y maze preference test was used to 'ask' sheep whether they preferred a human shaking a rattle or a barking dog. In both tests, adrenocortical responses were measured concurrently to support the interpretation of behaviour. The presence of a human or dog in the arena elicited significantly more avoidance and vigilance behaviour and less exploration than did the presence of a control object. However, the dog elicited significantly more of this fear-related behaviour, and significantly larger adrenocortical responses than did the human. Sheep also expressed a clear preference for a human shaking a rattle over a barking dog in the Y maze test and exhibited larger adrenocortical responses to the dog than to the human in the Y maze facility. The UWA flocks differed in their expression of locomotor and vocal activity; MA sheep were more active/vocal than the other flocks, not only in the presence of the human but also with the box or dog. MA sheep expressed less avoidance and vigilance and more exploration than the other flocks in the presence of the human and exhibited significantly lower plasma cortisol concentrations than LA sheep after exposure to the human (10-min sample). However, there were no inter-flock differences in fear-related behaviour or adrenocortical responses when the flocks were presented with the box or dog. The results do not support the notion that the UWA flocks have been selected for differences in a consistent predisposition to react fearfully. The adrenocortical responses measured in these studies were only moderate in magnitude and duration, with peak plasma cortisol concentrations 2-3 times higher than pre-treatment values, and all concentrations returning to pre-treatment levels within one hour of the start of treatment. If these observations are confirmed in practical situations, the presence of humans and dogs during routine handling should cause little concern on the basis of animal welfare. However, limiting the presence of dogs in certain situations (e.g. before slaughter) may reduce stress in domestic sheep. Significant methodological developments in this research include the use of multivariate statistical techniques to analyze arena behaviour, the concurrent measurement of adrenocortical and behavioural responses in the arena and Y maze tests, and the explicit testing of the effects of individual lateral biases on choice behaviour in a Y maze test. Future studies should measure sheep stress responses to the presence of humans and dogs in practical situations.

This thesis set out to examine the effect of maternal undernutrition and exposure to stressors between pregnancy day 50 to 100 and 100 to 147 of pregnancy on the ewe and her lamb. The long-term effects of these stressors during pregnancy on lamb growth, plasma cortisol response to a stressor, metabolism, behaviour and future reproductive success were examined. Mid-pregnancy shearing The component of mid-pregnancy shearing that causes the increase in lamb birth weight is unknown. It was hypothesised that the increase in lamb birth weight was due to the stress response of the ewe to shearing. This work examined the effect of a range of stressors at approximately day 80 of pregnancy. These stressors included yarding, crutching and sham-shearing that may be components of the shearing procedure that produce a stress response. In addition, repeated stressors between day 74 and 106 of pregnancy including isolation, sham-shearing and exogenous cortisol injection were used to examine the role of a longer-term stress response on lamb birth weight. Mid-pregnancy shearing has consistently resulted in an increase in lamb birth weight, however all the other stressors investigated had no effect. Therefore, the hormonal stress response of ewes to shearing was unlikely to be the cause of the increase in lamb birth weight. Mid-pregnancy shearing also resulted in minor changes in ewe and lamb behaviour 12 to 24 h after birth. Shearing during pregnancy had no effect on the cortisol response of ram lambs to handling or castration however differences were observed between singleton- and twin-born lambs. Ewe nutrition during pregnancy Ewes mildly undernourished in between days 70 and 107 of pregnancy that were then provided with adequate nutrition between days 108-145 of pregnancy gave birth to lambs with similar birth weights as ewes well-fed during both periods. Therefore the effects of undernutrition on lamb birth weight can be minimised if undernutrition can be limited to the earlier period (day 70-107 and 108-147 of pregnancy). Mild undernutrition in both periods (day 70 – 145 of pregnancy), resulted in lambs that were lighter and that exhibited behaviours associated with a greater ‘drive’ to maintain contact with their dam than lambs born to ewes well-fed during the same period. This suggests that even mild undernutrition in both mid- and late- pregnancy should be avoided. The effect of maternal nutrition and lamb litter size on the behaviour of female offspring was examined at 1 and 2 years of age. Litter size had no effect on ewe behaviour at 1 year of age however at 2 years of age twin-born ewes had higher maternal behaviour scores than triplet-born ewe lambs. Nutrition of the maternal grand dam during pregnancy had only a minor effect on the behaviour of female offspring at 1 year of age and no effect on maternal behaviour at 2 years of age.

This thesis set out to examine the effect of maternal undernutrition and exposure to stressors between pregnancy day 50 to 100 and 100 to 147 of pregnancy on the ewe and her lamb. The long-term effects of these stressors during pregnancy on lamb growth, plasma cortisol response to a stressor, metabolism, behaviour and future reproductive success were examined. Mid-pregnancy shearing The component of mid-pregnancy shearing that causes the increase in lamb birth weight is unknown. It was hypothesised that the increase in lamb birth weight was due to the stress response of the ewe to shearing. This work examined the effect of a range of stressors at approximately day 80 of pregnancy. These stressors included yarding, crutching and sham-shearing that may be components of the shearing procedure that produce a stress response. In addition, repeated stressors between day 74 and 106 of pregnancy including isolation, sham-shearing and exogenous cortisol injection were used to examine the role of a longer-term stress response on lamb birth weight. Mid-pregnancy shearing has consistently resulted in an increase in lamb birth weight, however all the other stressors investigated had no effect. Therefore, the hormonal stress response of ewes to shearing was unlikely to be the cause of the increase in lamb birth weight. Mid-pregnancy shearing also resulted in minor changes in ewe and lamb behaviour 12 to 24 h after birth. Shearing during pregnancy had no effect on the cortisol response of ram lambs to handling or castration however differences were observed between singleton- and twin-born lambs. Ewe nutrition during pregnancy Ewes mildly undernourished in between days 70 and 107 of pregnancy that were then provided with adequate nutrition between days 108-145 of pregnancy gave birth to lambs with similar birth weights as ewes well-fed during both periods. Therefore the effects of undernutrition on lamb birth weight can be minimised if undernutrition can be limited to the earlier period (day 70-107 and 108-147 of pregnancy). Mild undernutrition in both periods (day 70 – 145 of pregnancy), resulted in lambs that were lighter and that exhibited behaviours associated with a greater ‘drive’ to maintain contact with their dam than lambs born to ewes well-fed during the same period. This suggests that even mild undernutrition in both mid- and late- pregnancy should be avoided. The effect of maternal nutrition and lamb litter size on the behaviour of female offspring was examined at 1 and 2 years of age. Litter size had no effect on ewe behaviour at 1 year of age however at 2 years of age twin-born ewes had higher maternal behaviour scores than triplet-born ewe lambs. Nutrition of the maternal grand dam during pregnancy had only a minor effect on the behaviour of female offspring at 1 year of age and no effect on maternal behaviour at 2 years of age.

Diss. Uppsala : Sveriges lantbruksuniversitet, 2004.

Diss. (sammanfattning) Uppsala : Sveriges lantbruksuniv., 2007. / Härtill 4 uppsatser.

Thesis (M.Sc.) Uppsala : Sveriges lantbruksuniv. / Härtill 1 uppsatser.

For several centuries following their introduction into the British Isles by the Normans, rabbits were farmed on man-made warrens. The right to hunt rabbits during the medieval period was restricted to the highest strata of society and warrens, and rabbit products, carried connotations of wealth and exclusivity. During the post-medieval period, as rabbits became less expensive, their exclusivity declined and access to the species increased across a wider spread of the population. Consequently, later warrens tended to be purely commercial ventures that in places lingered as a form of animal husbandry up until the early twentieth century. Evidence of these warrens is particularly common across England and Wales and typically, although not exclusively, takes the form of pillow mounds, earthworks created to encourage rabbits to burrow. Despite their longevity and high numbers, warrens remain relatively little studied. This thesis investigates surviving warren architecture within south-west England, incorporating archaeological data into a GIS in order to identify the locational, morphological and typological trends of the region’s warrens. It also assesses associations between warrens and other classes of archaeology, notably elite residences and parks, large ecclesiastical institutions and prehistoric earthworks. Doing so allows for a better understanding of warrens’ roles within their immediate environs and of their relationships with other aspects of the human landscape. This study also addresses natural geographical aspects of the landscape in order to determine the principal factors that influenced where warrens were installed. This study investigates documentary reference to warrens as many have not survived within the landscape. Medieval chancery rolls in particular allow for the creation of a national framework of warrening so that the South West can be compared and contrasted to other regions of medieval England. Documentary references, both medieval and post-medieval, to the South West’s warrens allow for the creation of a discrete regional history that defines the context for the establishment of the region’s warren architecture. This study assesses how rabbits were interpreted by medieval society and discusses symbolism, particularly the visual role played by warrens in advertising their owners’ wealth and any possible religious concepts associated with rabbits.

This Master thesis is dedicated to pasture and the overall image of animal husbandry in Central Europe in La Téne Period and Roman Age. An important part is evaluation of osteological assemblages grazed animals, the question of the spectrum of species and creating models applied in the territory of today's Czech republic. A meaningful component of this study is comparison farmed animals at the time of La Téne and Roman period. Keywords: pasture - animal husbandry - agriculture - La Téne period - Roman period - archaeozoology