En Amérique du Nord, la superficie des monocultures utilisatrices d'intrants tels que des fertilisants et des pesticides couvre aujourd'hui 85% des surfaces agricoles et autant de pâturages ont disparu en 20 ans afin de répondre aux besoins de l'homme. Ces changements d'usage des terres ont profondément transformé le paysage et altéré la biodiversité des agro-écosystèmes. Parmi les espèces d'oiseaux champêtres, les insectivores aériens tel l'Hirondelle bicolore, Tachycineta bicolor, ici étudiée, présente un fort taux de déclin des effectifs, reflétant probablement une dégradation des agro-écosystèmes. Les mécanismes biologiques à l'origine de ce déclin sont encore méconnus ainsi que les patrons de dispersion chez les passereaux migrateurs. Le présent travail vise à étudier les effets environnementaux sur les traits individuels (survie et succès reproducteur) et la dispersion de reproduction (probabilité de disperser et taux d'occupation des sites de reproduction) chez l'Hirondelle bicolore. Pour ce faire, 2200 reproducteurs et 8000 oisillons ont été bagués entre 2004 et 2013 et suivis en reproduction pendant 10 ans sur une aire d'étude au Sud du Québec, laquelle est composée de 40 sites et couvre une mosaïque de paysages agricoles hétérogènes. Le développement d'un nouveau modèle de capture-marquage-recapture, flexible d'utilisation, a permis de réduire les biais d'estimation des probabilités de survie et de dispersion de l'espèce. Cette approche a permis de tester l'effet de plusieurs variables en lien avec la qualité de l'habitat, l'information publique et les caractéristiques individuelles sur la variabilité des paramètres de dispersion, de survie et de succès reproducteur au niveau individuel. Les milieux cultivés intensivement associés à la présence du Moineau domestique, Passer domesticus, un compétiteur pour les cavités de reproduction, diminuent jusqu'à 19% la survie des mâles. Les femelles sont quant à elles affectées par le coût de la reproduction, qui est d'autant plus grand en milieu intensif qu'il y a présence de moineaux et une disponibilité moindre en ressources alimentaires. Pour autant, la décision de disperser n'est pas affectée par les conditions environnementales que ce soit par la présence de compétiteurs ou la qualité du milieu, et ne dépasse pas les 4% chez les mâles. Les femelles dispersant jusqu'à 14 fois plus s'appuient sur leur expérience personnelle pour décider de disperser. Un échec de reproduction augmente jusqu'à 7 fois la probabilité de disperser et pour la première fois chez une espèce à courte durée de vie, nous montrons que la probabilité de disperser est augmentée l'année suivant une première dispersion. Ces patrons de dispersion sont stables dans le temps. La dispersion apparait donc comme un processus coûteux, comparé à la fidélité au site, qui apparaît comme un phénotype minoritaire dans la population. Elle répond à des conditions défavorables de reproduction. Enfin, ce travail montre l'utilisation de l'information publique (densité en partenaires, productivité en jeunes l'année précédente) dans la décision de s'établir sur un site et ce, une fois que la décision de disperser est amorcée. La présence de moineaux et la proportion de cultures intensives dans un rayon proche participent également à la sélection d'un site. D'après mes résultats, le milieu intensif contribuerait au déclin de l'Hirondelle bicolore. / In North America, monoculture areas using high inputs of fertilizers and pesticides cover 85% of agricultural lands, and as many pastures disappeared in the past 20 years to satisfy human food needs. These land-use changes have deeply transformed landscapes and altered the biodiversity of agro-ecosystems. Among farmland birds, aerial insectivores such as Tree swallow, Tachycineta bicolor, studied here show a high rate of decline in their abundance, partly reflecting the degradation of agro-ecosystems where they breed. Biological mechanisms leading to the decline and dispersal patterns in migratory passerines are still poorly known. The present work quantifies the environmental effects on Tree swallow individual traits (survival, reproductive success) and breeding dispersal (probability to disperse and occupation rates of breeding sites). Between 2004 and 2013, 2200 breeders and 8000 chicks were ringed and monitored during 10 breeding seasons on a study area composed of 40 sites and covering a mosaic of heterogeneous agricultural landscapes in southern Québec. The development of a new, flexible capture-mark-recapture model has reduced estimate bias of survival and dispersal probabilities of Tree swallows. This approach allowed to assess the effect of several variables linked to habitat quality, public information and individual characteristics on dispersal, survival and reproductive success parameters of individuals. Within intensively cultivated landscapes associated to the presence of a nest site competitor, the House sparrow, Passer domesticus, male survival is up to 19% lower. Females are affected by the cost of reproduction, especially in intensive landscapes where House sparrows and found and where food resources and nest site appear limited. Dispersal probability is not affected by environmental conditions either through the presence of House sparrows or habitat quality, and is restricted to 4% in males. Females, which disperse up to 14 times more than males, base their decision on their personal experience. Breeding failure increases up to 7 times the probability to disperse and for the first time in a short-lived species, I show that dispersal likelihood increases if the individual has dispersed in the previous year. These dispersal patterns are stable in time. Dispersal appears as a costly process compared to site fidelity and is a minority phenotype in the population. It appears an answer to unfavorable condition for reproduction. Finally, this work shows the use of public information (partners density, fledglings productivity the previous year) on the decision to settle on a site and this, after the dispersal decision has been initiated. The presence of House sparrows and the proportion of intensive areas near nestboxes are used as information to select a breeding site. Based on my findings, agricultural intensification likely plays a role in the decline of Tree swallow populations.

The Blue Penguin (Eudyptula minor) is assumed to be declining over much of its range, largely due to introduced predators. Anecdotal evidence suggests that one of the areas of declining population is the West Coast of the South Island. The purpose of this study was to determine the reasons for the assumed decline of blue penguins in South Westland. This was done by studying breeding ecology at several blue penguin colonies to assess the importance of breeding success and adult mortality on the penguin population. Three blue penguin colonies in South Westland, at Five Mile and Three Mile beaches south of Okarito, and at the Wanganui River mouth near Harihari, were monitored throughout the 2008/09 breeding season. During each burrow visit the number of eggs and chicks were recorded as well as the date of laying, hatching or fledging. Five colonies of blue penguin were also monitored in Buller over the same breeding season in a study conducted by the West Coast Blue Penguin Trust, a community trust based on the West Coast. The results of both studies were compared to determine the effect of predator control on breeding parameters, such as breeding success. Of 137 eggs laid in South Westland, 108 chicks survived until fledging, giving an overall breeding success of 78.8%. In Buller, 64 chicks survived to fledging from 101 eggs laid, resulting in an overall breeding success of 63.4%. Breeding success was significantly higher at penguin colonies in South Westland, compared to the Buller colonies. There was no evidence that predator control had an effect on breeding success in South Westland or Buller. The mean number of chicks fledged per pair that produced eggs was 1.55 in South Westland and 1.16 in Buller. The overall proportion of occupied breeding burrows compared to the total number of suitable burrows at the South Westland sites was 73.8% (n = 103). At the Buller sites, only 60.3 % (n = 151) of the total number of burrows was occupied. Road kills are a major threat to blue penguins in Buller due to the proximity of colonies to the state highway. Fortunately, incidences of road death in South Westland are rare and due to the distance from roads, do not pose a significant threat to South Westland blue penguins. Further study of blue penguin colonies in South Westland is needed to learn more about annual variation in breeding productivity and to determine if breeding success is consistently high over an extended time period. If this is the case, then the cause of blue penguin decline on the West Coast is unlikely to be due to problems with breeding as the breeding success during this study is one of the highest recorded for blue penguins. Although there was no apparent effect of predator control on breeding productivity during this study there is evidence from other locations that predators, in particular stoats, have contributed to the decline of blue penguin populations. More research into the impact of predators on penguins over a longer period of time is needed on the West Coast before a change is made to how predators are managed.

Les changements climatiques ont un impact certain sur les écosystèmes marins. Un déplacement vers le sud des principaux systèmes de front servant de sites de nourrissage pour de nombreuses espèces de prédateurs supérieurs est susceptible de se produire dans les zones subantarctiques. Les « central place foragers », tels que les pinnipèdes, sont susceptibles de faire face à une augmentation de la distance entre leurs lieux d'alimentation et leurs colonies d'élevage. Nous avons étudié l’impact des changements climatiques sur le succès d’élevage et la dynamique de population des otaries à fourrure (Arctocephalus gazella) des Îles Kerguelen par le biais du développement du modèle individu centré, MarCPFS (Marine Central Place Foragers Simulator) qui montre que la survie du couple femelle-jeune est particulièrement sensible à la répartition des proies (abondance et structure), à la capacité de mémorisation des meilleurs sites de nourrissages trouvés par la femelle pendant la période d'élevage, à la taille des femelles et à la distance qu'il faut parcourir pour trouver la ressource. Les résultats suggèrent qu’au cours des trois prochaines décennies un déplacement vers le sud supérieur à 2km an 1 pourrait compromettre la survie et la durabilité des populations. Un couplage avec un modèle de simulation de la dynamique océanique et de la ressource (SEAPODYM) a permis des projections jusqu’à la fin de ce siècle basées sur le scénarios RCP8.5 du GIEC (2014), confirmant les tendances mises à jour précédemment. Ces travaux montrent que les perspectives de maintien des populations sont pessimistes y compris en envisageant une adaptation de la taille des individus. / Climate change has certain impact on the marine ecosystems. A southward shift in productive frontal systems serving as the main foraging sites for many top predator species is likely to occur in the Subantarctic areas. Central place foragers, as seabirds and pinnipeds, are thus likely to cope with an increase in the distance between foraging locations and their land-based breeding colonies. We studied the impact of climate change on the breeding success and population dynamics of Antarctic fur seals (Arctocephalus gazella) at Kerguelen Islands by means of an individual based model we developped, MarCPFS (Marine Central Place Foragers Simulator) which showed that the survival of the female-pup pair is particularly sensitive to the distribution of preys (abundance and structure), to the memorization abilities of the best resource sites found by the female during the rearing period, to the female size and to the foraging distance which it is necessary to cover at each trip. The results suggest that during the next three decades a southward shift greater than 2 km year-1 could compromise the survival and the sustainability of the populations. A coupling with a model of simulation of the oceanic dynamics and the resource (SEAPODYM) allowed projections till the end of this century based on scenarios RCP8.5 of the IPCC (2014), confirming the trends previously obtained. Globally, these works lead us to conclude with pessimistic perspectives about the sustainability of populations even when considering an evolution through time towards individuals of greater sizes.

Nests are key structures for the reproduction of majority of avian species and as such they should be subject to natural selection. Six hypotheses have been suggested to explain variance in avian nest size. In my master thesis I evaluate their validity in the Great Reed Warbler (Acrocephalus arundinaceus). First two hypotheses describe responses of nest size to predation and brood parasitism. These two selection pressures may lead to the reduction of nest size, but no evidence of their impact on nest dimensions was obtained. However, I found a significant but negative relationship between the probability of nest predation and soft nest height. No such relationship was found between the probability of brood parasitism and nest size characteristics. The incidence of brood parasitism was affected only by nest visibility from the nearest cuckoo perch site and distance from open water. More visible nests suffered heavier parasitism while those located deeper in reed beds were better protected from cuckoo parasitism. Another four hypotheses describe selection pressures which favour large nests or some of their functional parts. The thermoregulatory hypothesis, the sexual display hypothesis and the nest support hypothesis did not explain nest size variation. I found support for the clutch size hypothesis,...

Ο Μαυροπετρίτης (Falco eleonorae Géné, 1839) είναι ένα μεσαίου μεγέθους μεταναστευτικό γεράκι το οποίο επισκέπτεται τη χώρα μας τους καλοκαιρινούς μήνες για να αναπαραχθεί. Οι αναπαραγωγικές αποικίες εντοπίζονται σε νησιά και ακτές της Μεσογείου, της Μακαρονησίας και της βορειοδυτικής Αφρικής, ενώ τους υπόλοιπους μήνες το είδος απαντά στη Μαδαγασκάρη και γειτονικές περιοχές. Στο παρελθόν έχει πραγματοποιηθεί ένας μεγάλος αριθμός μελετών σχετικά με την περίοδο αναπαραγωγής, ενώ για τον υπόλοιπο κύκλο ζωής του είδους οι γνώσεις μας μέχρι τις αρχές του 21ου αιώνα ήταν περιορισμένες. Στην παρούσα διατριβή μελετώνται τα βασικότερα στάδια του κύκλου ζωής του Μαυροπετρίτη, επικεντρώνοντας στην περίοδο αναπαραγωγής, μετανάστευσης και διαχείμασης ατόμων που αναπαράγονται στην Ελλάδα. Συγκεκριμένα, διερευνάται (α) η αναπαραγωγική επιτυχία του είδους και η επίδραση περιβαλλοντικών παραγόντων στην έκβαση της αναπαραγωγικής επιτυχίας, (β) το πρότυπο κατανομής των αναπαραγωγικών αποικιών στο Αιγαίο και Ιόνιο πέλαγος, (γ) τα κριτήρια που χρησιμοποιούν τα αναπαραγωγικά ζευγάρια κατά την επιλογή θέσης φωλιάσματος σε ακατοίκητες νησίδες του Αιγαίου πελάγους, (δ) η περίοδος μετανάστευσης τεσσάρων γερακιών από μια αποικία του Κεντρικού Αιγαίου και οι παράγοντες που επηρεάζουν το πρότυπο μετανάστευσής τους και, (ε) η περίοδος διαχείμασης και το πρότυπο κατανομής των τεσσάρων γερακιών στην περιοχή διαχείμασης. Η εκπόνηση της παρούσας διατριβής βασίσθηκε σε μεγάλο βαθμό στα πρωτογενή δεδομένα που συλλέχθηκαν στο πλαίσιο του προγράμματος «LIFE - ΦΥΣΗ 2003 Δράσεις για την προστασία του Μαυροπετρίτη (Falco eleonorae) στην Ελλάδα (LIFE 03NAT/GR/000091)», με κύριο ανάδοχο την Ελληνική Ορνιθολογική Εταιρεία (Ε.Ο.Ε.) σε συνεργασία με το Μουσείο Φυσικής Ιστορίας Κρήτης (ΜΦΙΚ), το Υπουργείο Αγροτικής Ανάπτυξης και Τροφίμων και τη Βρετανική Ορνιθολογική Εταιρεία (RSPB) και με χρηματοδότηση της Ευρωπαϊκής Επιτροπής και συγχρηματοδότηση του ιδρύματος «Α. Γ.Λεβέντης». Το πρόγραμμα παρακολούθησης της μετανάστευσης των τεσσάρων Μαυροπετριτών πραγματοποιήθηκε στο πλαίσιο του προγράμματος «Μελέτη των πτηνών του ελληνικού θαλάσσιου χώρου» με ανάδοχο την Ελληνική Ορνιθολογική Εταιρεία και με χρηματοδότηση του ιδρύματος «Α. Γ. Λεβέντης». / Eleonora's falcon (Falco eleonorae Géné, 1839) is a medium-sized migratory raptor that visits the Greek islands during the breeding season. Its breeding colonies are distributed on islands and coasts of the Mediterranean Sea, Macaronesia and northwest Africa, while during the rest of the year the species is found in Madagascar and surrounding islets. In previous years most studies focused on the breeding period, while up to the 21st century our knowledge with regards to the rest of its life cycle remained relatively poor. In this thesis i study the main stages of Eleonora's falcon life cycle, focusing on the breeding, migratory and wintering period of individuals that breed in Greece. In particular, i investigate (a) the breeding success of the species and the effect of environmental factors on the outcome of the breeding effort, (b) the distribution pattern of the breeding colonies in the Aegean and Ionean Sea, (c) the criteria the breeding pairs use at the time of nesting site selection in uninhabited islets of the Aegean Sea, (d) the migratory period of four falcons originating from a breeding colony of the Central Aegean Sea and the factors that shape the irmigratory pattern and, (e) the wintering period and the distribution pattern of the four falcons in their wintering grounds. This thesis was based to a great extent on the data collected in the frame of the project "LIFE-Nature 2003 Conservation Measures for Falco eleonorae in Greece (LIFE 03NAT/GR/000091)" undertaken by the Hellenic Ornithological Society (H.O.S.) in collaboration with the Natural History Museum of Crete (NHMC), the Ministry of Rural Development and Food and the Royal Society for the Protection of Birds (RSPB), which was funded by the European Commission and cofunded by the A.G. Leventis Foundation. The migration tracking project was funded by the A.G. Leventis Foundation through project "Survey and Conservation of Seabirds in Greece".

Studies of mating systems and social organisation have been central to understanding of the evolution of social behaviour. The European badger Meles meles is a good species in which to study these processes, as its complex social system provides an opportunity to investigate how both natural and kin selection shape the evolution of mating systems and social structure. In this thesis, I use behavioural and genetic data to describe the mating system and social organisation of a high-density badger population and examine the occurrence of cooperative breeding. I genotyped 915 (85%) badgers trapped in Wytham Woods (1987–2005), 630 of which were cubs, and assigned both parents to 331 cubs with 95% confidence. This revealed a polygynandrous mating system, with up to five mothers and five fathers per social group. Mounting behaviour was also polygynandrous and I show the strongest evidence to date for multiple-paternity litters. I demonstrate, for the first time, that groups consisted of close and distant kin: approximately one third of group members were first-order kin, and overall group members had slightly lower relatedness levels than half-siblings. Within groups, adult and yearling females had higher pairwise relatedness than males, and neighbouring groups contained relatives. These findings result from the high level (42%) of extra-group paternities, 86% of which were assigned to neighbouring males. For the first time I show that females avoided inbreeding by mating with extra-group males; however, incestuous matings did occur. Promiscuous and repeated mountings were observed, which may reduce male–male aggression and infanticide, but may also promote sperm competition, genetic diversity, and / or genetic compatibility. Just under a third of adult males and females were assigned parentage each year and I quantify, for the first time, reproductive skew within badger groups. Correlations between relatedness, group productivity, and reproductive skew were not consistent with the predictions of incomplete-control models; rather, resource availability may play a role. Older and younger badgers displayed reduced annual breeding success, with male success increasing initially with experience. The Restraint, Constraint, and Selection Hypotheses did not explain the age-related breeding pattern in females. Variance in lifetime breeding success (LBS) was greater for males. Males that only bred within or only outside of their groups had half the LBS of males that did both. Females that were assigned maternity probably bred cooperatively and allonursed non-offspring, which has not been demonstrated previously. No benefit was established, however, in terms of litter size, probability of offspring breeding, or offspring lifetime breeding success, with more mothers in a group. In conclusion, badger social groups are fostered through kinship ties. Polygynandry and repeated mounting may have evolved originally to reduce male–male aggression and infanticide by males, through paternity masking. Although plural breeding occurs, group living appears to be costly. Motivation to disperse may be reduced through high-levels of extra-group paternities, which may also reduce inbreeding. Cooperative breeding among mothers may represent a low-cost behaviour with indirect benefits due to high levels of relatedness between female group-members. Badger sociality therefore represents an early stage in the evolution of social behaviour.

No description available.