Mink (Mustela vison) are an important species because they occur at a high trophic level, they are considered a potential indicator species for environmental contaminants, and they are a popular target species among fur trappers. Despite the importance of mink, their ecology in North America is not well understood. I sampled 59 of 60 Hudson River tributaries with 2 scent stations randomly established along each tributary for 2 10-day monitoring periods to evaluate presence/absence of mink via remote camera photographs and tracks. When sampling was completed, I quantified microhabitat characteristics at 84 randomly selected scent stations. Statistical tests for microhabitat did not indicate a difference among variables selected for analysis between scent stations where mink visits were detected and those where mink visits were not detected. I also analyzed all scent stations for macrohabitat characteristics related to human disturbance, using 150 m circular buffers in ArcGIS 9.2. Although the percent cover of human disturbance was 7.0% greater at scent stations where mink were not detected, the difference was not significant and may therefore indicate that mink may have a tolerance for moderate human disturbances. I also evaluated the performances of 2 types of remote cameras (Moultrie Game Cameras: MGC I–40 and MGC 200) in detecting mink visits at scent stations as well as comparing remote camera detections of mink visits with observations of tracks. The MGC I–40 cameras detected a significantly greater number of mink visits (n = 50) compared to the MGC 200 cameras (n = 3). Detection of mink at scent stations was also significantly greater using the remote cameras compared with observations of mink tracks. Mink were also live–trapped and implanted with subcutaneous radiotransmitters. There were 13 mink captures (0.31 mink/100 trap–nights) with radiotransmitters being implanted in 12 (11 males and 1 female) mink. Overall 166 den sites were located with a mean of 15.9 den sites/mink. Microhabitat analysis of 33 used den sites and 33 unused potentially available den sites (UPADs) indicated that shoreline cover was significantly greater at used den sites. Macrohabitat analysis using 150 m circular buffers surrounding 76 used den sites and 76 unused potentially available sites (UPASs) indicated that human disturbance was 3.3% greater at the UPASs than at used den sites, but this difference was not significant. This may further suggest that mink may have a tolerance for moderate human disturbances. Den site structures most often used by mink included brushpiles or logjams (21.1%) and bank burrows (17.5%). The use of den site structures appeared to be largely based on shoreline cover and availability. The mean linear home range of male mink was 6.6 km (SE = 0.6, range 3.2 – 8.4 km), which was similar to the female's home range of 6.5 km. Ten mink home ranges encompassed portions of both the Hudson River and its tributaries accounting for a mean of 2.8 and 4.4 km, respectively. Mean daily movement distances of mink along shorelines was 659 m (SE = 42, range 0 – 3,087 m) and was significantly greater along tributaries than along the Hudson River.

Movement of organisms is a fundamental component of many ecological processes, and should be subject to strong selective pressures. Spatial selection is the process by which individuals choose the locations to acquire necessary resources or avoid risk, and the relative importance of different factors on spatial selection may change depending on the scale being analyzed. Under the framework of optimality, an individual should attempt to structure their spatial selection economically to maximize fitness. I studied black bear (Ursus americanus) space use, habitat selection, and movement under the optimality paradigm in three populations (Michigan, Missouri, and Mississippi) at four different scales: regional, annual home range, seasonal home range, and denning sites. At the regional scale, I found that black bears displayed scale-dependent land cover selection for movement, selecting forested areas at coarser scales and avoiding anthropogenic disturbance at finer scales, and that large contiguous forests and riparian corridors most facilitate connectivity among protected areas. At the annual and seasonal home range scales, I found black bears display scale-dependent optimizing strategies. Individuals locating their annual ranges to maximize access to areas of high vegetation productivity, together with the high productivity of ranges of all sizes, suggests an energy maximizing strategy, while the negative relationship between range size and both fragmentation and forest proportion suggests area minimizing. More limiting factors act at larger scales, which suggests productivity is the strongest limiting factor and energy maximizing is the dominant strategy while plasticity allows for seasonal area minimizing. At the den site scale, I found that both female and male black bears appeared to minimize anthropogenic risk during denning; however female black bears have a flexible response to anthropogenic disturbance, attempting to minimize it when alone or with older offspring, yet having increased tolerance when infanticide is greater after cubs are born and following den emergence. By quantifying black bear space use and selection across multiple scales, diverse areas, over time, and among and within individuals, I revealed consistent scale-dependent responses to environmental and biological factors while highlighting the intrinsic plasticity of this flexible omnivore.

Abstract The grey wolf is the most widely distributed of all land mammals and is a habitat generalist that inhabits all the vegetation types of the Northern Hemisphere. Wolves also breed well and have the potential to rapidly expand to new areas. In Finland, the wolf is a game species, and as a result of Finland’s membership of the European Union, the wolf population is subject to very limited hunting due to the obligation to protect the species. Mainly for this reason, the wolf population in Finland has increased significantly in recent years. In particular, the birth rate has developed favourably and the number of litters increased from just four in 1996 to 20 in 2005. It also seems at present that the wolf population in Finland is no longer following the fluctuations in wolf numbers in Russian Karelia. The general aim of this dissertation is to provide applicable knowledge for wolf management and conservation purposes, and especially to examine the effects of human-modified landscapes on wolf population expansion in Finland. Various aspects of habitat selection were investigated in four sub-studies to gain a thorough insight into the space use and habitat needs of wolves. Species data came from a long-term wolf population study that included location information from 85 radio- and GPS-GSM-collared wolves from 1998 onwards, as well as track location data based on about 30 000 annual observations recorded with geographical coordinates by a local network of experts on large carnivores. I found that adaptability makes it possible for the wolf to live in the multiple-use, semi-wild forests of Finland and that no restrictions are imposed by the landscape on wolf population growth and expansion. In general, the results of my dissertation provide evidence that wolves tend to avoid the presence of human influence when establishing a territory and also when selecting their den site. However, as wolf numbers increase, conflict situations will more frequently occur between wolves and humans, although the risk of depredation events, for example among sheep farms, varies between farms in Finland. That is, there are some environmental and farm level factors that are associated with wolf depredation. The breeding wolf population in Finland has gradually expanded and the first litters have recently been born in western Finland after an absence of more than 100 years. The geographical distance to the Scandinavian population is shorter from these new western territories than from the population’s core area in eastern Finland. This could potentially increase the likelihood of dispersal from Finland to Scandinavia. / Tiivistelmä Susi on habitaattigeneralisti, jonka levinneisyys kattaa kaikki pohjoisen pallonpuoliskon kasvillisuustyypit. Sudet myös lisääntyvät hyvin ja voivat näin ollen levittäytyä nopeasti uusille alueille. Suomessa susi on riistalaji, jonka metsästys on rajoitettua Euroopan unionin jäsenyyden myötä. Lähinnä tästä syystä Suomen susikanta on kasvanut voimakkaasti 1990-luvun ja 2000-luvun alun aikana. Erityisesti syntyvyys on kasvanut: pentueiden määrä lisääntyi neljästä 20:een vuosien 1996 ja 2005 välillä. Näyttäisi siltä, ettei Suomen nykyinen susipopulaatio enää seuraa Venäjän Karjalassa tapahtuvia kannanvaihteluita. Väitöskirjan tavoitteena on ollut tuottaa susikannan hoidossa ja suojelussa käytettävissä olevaa tietoa ja erityisesti tutkia susikannan levittäytymistä ihmisen muokkaamassa ympäristössä Suomessa. Neljässä osatutkimuksessa on käsitelty suden habitaatinvalintaa eri näkökulmista, jotta saataisiin kokonaiskuva suden tilankäytöstä Suomessa. Tutkimusaineisto rakentui pitkäaikaisesta seuranta-aineistosta, jota on kerätty vuodesta 1989 lähtien. Vuosien 1989 ja 2010 välillä 85 sutta on pannoitettu radio- tai GPS-GSM –pannalla. Tutkimuksessa käytettiin hyväksi myös noin 30 000 vuosittain tehtyä jälkihavaintoa. Tutkimustulosten perusteella susi selviää hyvin Suomen monikäyttömetsissä, eivätkä maisematyypit itsestään rajoita susipopulaation kasvua ja levittäytymistä. Toisaalta tulokset osoittavat myös sen, että sudet välttelevät ihmistoimintoja valitessaan reviiri- tai pesäpaikkoja. Kuitenkin susien lukumäärän kasvaessa ristiriitatilanteet susien ja ihmisten välillä tulevat lisääntymään, vaikka susivahinkoriskin suuruus vaihteleekin Suomessa huomattavasti. Esimerkiksi lammastilojen susivahinkoriski riippuu sekä ympäristötekijöistä, että tilan sisäisistä tekijöistä. Kaiken kaikkiaan lisääntyvän susipopulaation esiintymisalue on Suomessa vähitellen kasvanut leviten itärajan tuntumasta kohti länttä. Tämän myötä maantieteellinen etäisyys Skandinavian susipopulaatioon on pienentynyt viimeisten vuosien aikana. Tämä voi edesauttaa susiyksilöiden siirtymistä Suomesta Skandinaviaan.