>Magister Scientiae - MSc / The research was driven by a need to increase the knowledge base concerning wetland ecological responses, as well as to identify and evaluate the factors driving the functioning of the Franschhoek Trust Wetland. An ecohydrological study was undertaken in which vegetation cover, depth to groundwater, water and soil chemistry were monitored at 14 sites along three transects for a 12 month period. The parameters used include temperature, pH, electrical conductivity (EC), sodium, potassium, magnesium, calcium, iron, chloride, bicarbonate, sulphate, total nitrogen, ammonia, nitrate, nitrite and phosphorus. T-tests and Principal Component Analysis (PCA) were used to analyze trends and to express the relationship between abiotic factors and vegetation. Results reflect the strong influence of hydrology, microtopography and nutrient availability in structuring vegetation composition in the wetland. The wetland has been classified as a palustrine valley bottom with channel wetland, which is predominantly groundwater-fed (phreatrotropic), but receives surface water inputs as well. Small scale gradients of microtopography allow for differences in flooding frequency and duration resulting in hydrologically distinct sites which differ chemically. Three zones were distinguished in the wetland. Hollows or low sites were characterized by intermittent flooding and drying and higher nutrient concentrations in soil and groundwater. High sites which were rarely or never flooded exhibited higher groundwater temperature and ammonia as well as iron in soils and groundwater. The inundated sites remained flooded throughout the year and were characterized by high nitrate and nitrite in soil as well as high EC, magnesium, bicarbonate, sulphate and phosphorus in groundwater. The limited availability of nitrogen in the wetland favoured plant types Typha capensis, Paspalum urvillei and Juncus .kraussii which are able to either fix nitrogen or store nitrogen during more favorable conditions. The main chemical concentration changes take place between summer and winter. The Principal Component Analyses suggest that sodium, chloride, potassium, ammonia and phosphorus are the dominant ions determining the chemistry of groundwater. Increased abstraction from the table mountain aquifer to supplement human demand may put the wetland at risk of degradation. Intensified agriculture and other land use in the area are likely to increase pollution loads into the wetland causing shifts in nutrient availability and vegetation composition. Continued and long term monitoring is essential to ensure effective management of the wetland and is highly recommended. Closer partnerships between wetland managers and scientists as well as community awareness and involvement through a volunteer monitoring programme should be encouraged

Indiana University-Purdue University Indianapolis (IUPUI) / Drylands are the largest terrestrial biome on the planet, and the critically important systems that produce approximately 40% of global net primary productivity to support nearly 2.5 billion of global population. Climate change, increasing populations and resulting anthropogenic effects are all expected to impact dryland regions over the coming decades. Considering that approximately 90% of the more than 2 billion people living in drylands are geographically located within developing countries, improved understanding of these systems is an international imperative. Although considerable progress has been made in recent years in understanding climate change impacts on hydrological cycles, there are still a large number of knowledge gaps in the field of dryland ecohydrology. These knowledge gaps largely hinder our capability to better understand and predict how climate change will affect the hydrological cycles and consequently the soil-vegetation interactions in drylands. The present study used recent technical advances in remote sensing and stable isotopes, and filled some important knowledge gaps in the understanding of the dryland systems. My study presents a novel application of the combined use of customized chambers and a laser-based isotope analyzer to directly quantify isotopic signatures of transpiration (T), evaporation (E) and evapotranspiration (ET) in situ and examine ET partitioning over a field of forage sorghum under extreme environmental conditions. We have developed a useful framework of using satellite data and trend analysis to facilitate the understanding of temporal and spatial rainfall variations in the areas of Africa where the in situ observations are scarce. By using a meta-analysis approach, we have also illustrated that higher concentrations of atmospheric CO2 induce plant water saving and the consequent available soil water increases are a likely driver of the observed greening phenomena. We have further demonstrated that Leuning’s modified Ball-Berry model and RuBP limited optimization model can generally provide a good estimate of stomatal conductance response to CO2 enrichment under different environmental conditions. All these findings provide important insights into dryland water-soil-vegetation interactions.

Maintaining flows and quality of water resources is critical to support ecosystem services and consumptive needs. Understanding impacts of changes in climate and land use on ecohydrologic processes in a watershed is vital to sustaining water resources for multiple uses. This study completes a continental and regional scale assessment using statistical and simulation modeling to investigate ecohydrologic impacts within watershed systems. Watersheds across the continental United States have diverse hydrogeomorphic characters, mean temperatures, soil moistures, precipitation and evaporation patterns that influence runoff processes. Changes in climate affect runoff by impacting available soil moisture, evaporation, precipitation and vegetative patterns. A one percent increase in annual soil moisture may cause a five percent increase in runoff in watersheds across the continent. Low soil moisture and high temperatures influence runoff patterns in specific regions. Spring runoff is increased by the influence Spring soil moisture, Winter and Spring evaporation, and Winter and Spring evaporation. Spring runoff is decreased by increases in Winter and Spring temperatures and increases in the vegetation index. Winter runoff is affected by maximum vegetative index, temperature, soil moisture, evaporation and precipitation. Contributing factors to runoff are influenced by geomorphic and seasonal variations requiring strategies that are site-specific and use system-wide information. Regional scale watershed analysis investigates the influence of landscape metrics on temporal streamflow processes in multiple gauged watersheds in Massachusetts, U.S.A. Time of concentration, recession coefficient, base flow index, and peak flow are hydrologic metrics used to relate to landscape metrics derived using FRAGSTAT software. Peak flow increases with increasing perimeter-area fractal dimensions, and Contagion index and decreases as Landscape Shape Index increases. There was an increasing trend in the fractal dimension over time indicative of more complex shape of patches in watershed. Base flow index and recession coefficient fluctuated from low to high decreasing recently. This could be indicative of open space legislation, conservation efforts and reforestation within the state in the last ten years. Coastal systems provide valuable ecosystem services and are vulnerable to impacts of changes in climate and continental land use patterns. Effects of land use and climate change on runoff, suspended sediments, total nitrogen and total phosphorus are simulated for coastal watersheds around the Boston Bay ecosystem. The SWAT (Soil and Water Assessment Tool) model, a continuous-time, semi distributed, process-based model, is used to simulate the watershed ecohydrologic process affecting coastal bodies. Urbanization in watersheds increased runoff by as much as 80% from the baseline. Land use change poses a major threat to water quality impacts affecting coastal ecosystems. Total nitrogen increased average of 53.8% with conservative changes in climate and land use. Total phosphorus increased an average of 57.3% with conservative changes in land use and climate change. Climate change alone causes up to 40% increase in runoff and when combined with a 3.25% increase in urban development runoff increased an average of 114%. Coastal ecosystems are impacted by nutrient runoff from watersheds. Continued urbanization and changes in climate will increase total nitrogen, total phosphorus and suspended sediments in coastal ecosystems. Continental scale runoff is affected by soil moisture and vegetative cover. Cover crops, low tillage farm practices and natural vegetation contribute to less runoff. Developing policies that encourage protection of soil structure could minimize runoff and aid in maintaining sustainable water resources. Best Management Practices and Low impact development at the national level with continued stormwater legislation directed towards sustainable land use policy will improve water quantity and quality. Fragmentation observed in Massachusetts increases the number of urban parcels and decreases the size of forested areas. Faster runoff patterns are observed but recent land management may be changing this runoff pattern. Municipal and state zoning ordinance to preserve open space and large forest patches will restrict urban growth to specific regions of a watershed. This could improve quantities of water available to ecosystems. Increases in total nitrogen, phosphorus and suspended sediments to coastal ecosystems can be minimized with use of riparian buffers and Best Management Practices within coastal watersheds. Urbanization and climate change threatens coastal ecosystems and national policy to preserve and restrict development of coastal areas will preserve coastal ecosystem services.

Northern peatlands have acted as persistent sinks of CO2 throughout the Holocene largely owing to their ability to maintain shallow water table depths that limit decomposition rates and supports the growth of keystone vegetation including Sphagnum mosses. There is concern, however, that the future success and ecosystem function of these northern peat deposits may be at risk to climate change, where temperatures and evaporation rates are predicted to increase substantially in the next century. While numerous studies have examined the hydrology and carbon dynamics in large expansive peatland systems where a water table (WT) is ever-present, relatively little research has been done on small scale peat-accumulating systems where their vulnerability remains unknown. One region where a broad spectrum in the scale of peat accumulation is present is in the bedrock depressions of Canadian Shield rock barrens, which are of special importance as many peat deposits here provide habitat to species at risk including the Blanding’s Turtle and the Massassauga Rattlesnake. This study examines the controls that govern water storage dynamics and moss water availability in 18 different peat-accumulating depressions that vary in size, catchment area, and sediment composition. The magnitude of WT variability was often several times greater in shallower bedrock depressions (<50 cm deep) as compared to deeper ‘bogs’ (>60 cm deep). The magnitude of depression WT variability appeared to be closely linked to the WT depth (WTD), the relative proportions of different sediment types within the depression, and the depth dependant specific yield (Sy) of each sediment type. Sites which contained large fractions of Polytrichum moss or mineral soil – which were more common in shallow depressions ¬¬– had the greatest WT variability due to the lower porosity and Sy of this sediment as compared to Sphagnum peat. Sphagnum dominated ‘vernal pools’ (30-50 cm deep) had a WT variability two to three times greater than Sphagnum dominated bogs at WTDs > 20-25 cm, which may be related to exceptionally high ash concentrations near the base of vernal pools which reduced peat porosity and Sy as compared to more organic-rich peat. As compared to bogs, pits (<15 cm deep) and vernal pools had greater rates of WT decline during drying intervals, deeper average WTDs when a WT was present, and extended periods of WT absence during the summer months. As such, moss growing in pits and vernal pools generally had lower near-surface water availability as compared to bogs, though the importance of depression depth in determining the timing of moss stress is also dependant on the hydrophysical properties (Kunsat and moisture retention) of the moss species in question. WT dynamics and moss water availability were generally weakly correlated to depression catchment size, although during wetter periods of the year the rate of WT recession was moderated in pits and vernal pools which had an upslope depression that could provide sustained water inputs for multiple days after rainfall. The results of this study suggest that depression depth may be a first order control in determining peatland vulnerability to future regime shifts induced by external forcings or disturbances. Furthermore, this study suggests that systematic differences may exist between the hydrophysical properties of peat in shallow vs. large bedrock depressions, potentially resulting from contrasts in fire frequency/severity, and/or the degree of humification/compression among geological settings. / Thesis / Master of Science (MSc) / Canada is home to one of the largest reservoirs of organic carbon stored on land in the world, in unique ecosystems called peatlands. Peatlands are formed in wetland environments where a thick layer of organic matter has accumulated over time due to the average rate of vegetation growth on the surface of peatlands exceeding the rate of decomposition of the underlying organic matter. This net accumulation of organic matter over time has caused peatlands to act as a long term sink of carbon dioxide, which is a greenhouse gas that is a primary driver of global warming. The ability of peatlands to have slow decomposition rates and support the growth of key peatland vegetation, most notably various species of ‘peat moss’, is highly dependent upon their ability to keep their water table (i.e. the surface below which pore spaces in the organic matter are saturated with water) close to their growing surface. There is concern, however, that a warmer and dryer climate in the future could cause deeper water table positions in peatlands, thereby increasing decomposition rates, decreasing the growth rate of peat moss, and potentially turning peatlands into a net source of carbon dioxide. Most peatland studies to date, however, have focused on water storage/movement and carbon exchange in large, deep peatland systems, whereas relatively little research has been conducted on smaller peatlands. As such, the vulnerability of these smaller peatlands to future climate change remains uncertain. One region where peatlands exist over a wide range of different sizes and landscape positions is in bedrock depressions of the Canadian Shield, which are of special interest as they also provide habitat for species at risk including the Blanding’s Turtle and the Massassauga Rattlesnake. This study looked at how the water table positions and water availability to different species of peat moss compared over the growing season between 18 peatlands of different sizes and landscape position (i.e. peatlands with a relatively ‘small’ and ‘large’ area upslope of them). This study finds that deeper peatlands (with organic matter layers > 60 cm deep) usually had a shallower water table over the summer months than shallower peatlands (< 50 cm deep), primarily due to differences in the properties of the organic matter underlying their growing surfaces. Furthermore, each of the 12 studied peatlands < 50 cm deep lost their water table for a considerable amount of time during the summer (when their water table position dropped below the underlying bedrock of the depression), whereas each of the six peatlands > 60 cm deep had a water table present for the entire growing season. Surprisingly, a peatland’s position on the landscape seemed to have a relatively minor effect on determining the depth/presence of its water table. As deeper peatlands usually had a water table that was closer to the growing surface and was always present, more moisture was available to the peat moss growing at their surface than for peat moss in shallower depressions, though this moisture availability also depended upon the growth form of the different species of peat moss (some species of peat moss were better at accessing subsurface water than others). Through its impact on water table positions and moisture availability for peat moss, peatland depth is likely a primary control governing peatland vulnerability climate change, with shallower peatlands being more vulnerable to warmer and dryer conditions in the future.

No description available.

Comprendre les relations entre écohydrologie et structure fonctionnelle des communautés végétales est une problématique croissante en écologie. Cette thèse a pour objectif d'analyser les effets d'un gradient de ressource édaphique sur la structure et le fonctionnement de parcours herbacés dans le sud de la France, en intégrant les niveaux de l'écosystème à la plante. Par modélisation du bilan hydrique, les différences d'utilisation de l'eau au niveau de l'écosystème ont été quantifiées pour cinq années contrastées révélatrices de la variabilité des propriétés des sols et du fonctionnement végétal. Utilisation de l'eau et productivité des plantes sont proportionnelles à la réserve utile des sols et à son taux de remplissage, suggérant un équilibre fonctionnel entre les communautés végétales et la disponibilité de l'eau locale. Par une approche ‘traits', nous avons ensuite montré que le ratio entre surface foliaire et surface racinaire totales de la communauté est le facteur déterminant une relation 'allométrique' entre évapotranspiration potentielle et capacité de prélèvements hydriques. La coordination entre les compartiments racinaire et aérien de la végétation dépend d'une suite d'adaptations architecturales et morphologiques. Des modifications d'abondances relatives spécifiques et le turn-over d'espèces génèrent une variabilité de morphologie et de fonctionnement des communautés permettant leur adaptation à des ressources limitées. Au niveau plante, une expérimentation de transplantation a permis de quantifier l'importance relative de la disponibilité hydrique et des interactions entre plantes sur la productivité individuelle de trois espèces-cibles. L'importance de la compétition s'est révélée plus élevée sous fort niveau de ressource et dépendait plus des relations hiérarchiques entre traits que de la biomasse. La compétition entre plantes affecte les axes de niche fonctionnelle des espèces de façon indépendante, avec une convergence de la ‘teneur en matière sèche des feuilles' pour des faibles valeurs mais une plus forte divergence fonctionnelle pour la ‘hauteur des plantes'. Cette approche fonctionnelle a permis d'éclairer le rôle de la diversité végétale dans la réponse des systèmes écologiques à des ressources hydriques variables. De plus, ces résultats sur les propriétés écohydrologiques des communautés naturelles peuvent contribuer à la conception d'agro-écosytèmes complexes mieux adaptés aux sécheresses intenses prévues dans le contexte actuel de changement climatique. / Understanding the relationships between ecohydrology and functional structure of plant communities is a timely issue in plant ecology. From ecosystem to species levels, this works investigated the effects of a resource availability gradient on the structure and functioning of Mediterranean rangelands in Southern France. Using a water-balance modeling approach, differences in water-use at the ecosystem level were quantified across five contrasting years, accounting for the variability in both soil characteristics and vegetation functioning. Plant water-use and productivity scaled with soil water storage capacity and its filling rate, suggesting a functional equilibrium between plant communities and local water availability. Using a trait-based approach, we then showed that the ratio of total leaf area of the community on total root area determined an allometric relationship between potential evapotranspiration and water uptake capacity. The “coordination” between above- and belowground compartment of the vegetation was achieved by a suite of architectural and morphological adaptations. Modifications of species relative abundances and species turn-over explained morphological and functioning adaptations to cope with limiting resources. Finally at species level, a removal experiment allowed to quantify the relative importance of water limitations and effects of plant-plant interaction on individual plant productivity of three target species of graminoïds. The importance of competition increased with soil resource availability and depended more on trait-trait hierarchical distances among species than standing biomass. Competition among plants affects the axes of the functional niche of species independently, leading to the convergence of leaf dry matter content towards low values but to greater functional divergence for plant height. Overall, this functional approach provided an integrative understanding of the role of plant diversity in the response of ecological systems to changes in water availability. Furthermore, these results on ecohydrological properties of natural communities can contribute to the design of complex agro-ecosystem better adapted to intense droughts predicted under climate change.

Environ 65% des eaux intérieures sont menacées modérément ou fortement par l'altération anthropique et le changement climatique. La Terre a perdu environ la moitié de ses habitats d'eau intérieure au cours des cent dernières années et le prélèvement mondial d'eau devrait augmenter de plus de 50% au cours des 20 prochaines années. Dans ce contexte, la concurrence pour les ressources en eau entre les humains et les écosystèmes est appelée à s'intensifier au détriment de la biodiversité d'eau douce. Pour cette raison, il est aujourd'hui impératif de quantifier et de sauvegarder les besoins en eau de ces écosystèmesEn tant qu'outil global d'aide à la prise de décision, plusieurs modèles d'évaluation d'impact de la consommation d'eau sur les écosystèmes ont été proposés dans le cadre de l'analyse du cycle de vie (ACV). L'un des défauts importants des modèles actuels d'ACV, qu'ils soient mécanistes ou non-mécanistes, midpoint ou endpoint, c’est la prise en compte limitée des besoins environnementaux en eau (EWR). Pour cette raison, les approches existantes pour l'évaluation des EWR ont été étudiées, afin d'évaluer les avantages potentiels d'une meilleure prise en compte des principes écohydrologiques dans l’ACV sur les écosystèmesCela a permis de définir le concept d'habitat d'eau douce en ACV et d'élaborer un cadre conceptuel pour l'application des méthodes de simulation d'habitat dans la modélisation. Un indicateur midpoint du potentiel de changement d'habitat (HCP) évaluant l'impact de la modification du débit sur les habitats des poissons et des invertébrés dans les cours d'eau a été élaboré. Le nouveau modèle a été testé sur le réseau fluvial français à l'échelle du tronçon fluvial, en caractérisant le changement d'habitat en saison sèche et humide. Les HCP ont ensuite été agrégés à l'échelle du bassin versant. Après, le nouveau modèle a été généralisé pour permettre une extension globale. Les variables d'entrée du modèle HCP généralisé ont été calculées à partir des bases de données et des modèles existants sur une résolution mensuelle. Les limites, l'incertitude et les perspectives de recherche de la nouvelle approche ont finalement été discutées. / Approximately 65% of inland waters are under moderate or high threat by anthropogenic alteration and climate change. Earth has lost around half of inland water habitats in the last hundred years and global water withdrawal is expected to increase by more than 50% within the next 20 years. In this context, competition for water resources between humans and ecosystems is set to rise at the expense of freshwater-dependent biodiversity. For this reason, nowadays it is imperative to quantify and safeguard water needs of freshwater-dependent ecosystems.As a global tool to support decision-making, in life cycle assessment (LCA) several models for life cycle impact assessment (LCIA) of water consumption on ecosystems have been proposed. One important flaw of current LCIA models, whether they are mechanistic or non-mechanistic, midpoint or endpoint-oriented, is their limited consideration of environmental water requirements (EWR). For this reason, existing approaches for EWR assessment have been investigated to evaluate potential benefits of better including ecohydrological principles in LCIA on ecosystems.This enabled the definition of the concept of freshwater habitat within the boundaries of LCIA and the development of a framework for the application of habitat simulation methods in LCIA modeling. A midpoint habitat change potential (HCP) indicator assessing the impact of flow alteration on instream habitats of fish and invertebrates was developed. The new model has been tested on the French river network at the river reach scale, characterizing habitat change in wet and dry seasons. HCPs were then aggregated at watershed scale. The new model was subsequently generalized to allow global extension. Input variables of the generalized HCP model have been calculated from existing databases and models on a monthly resolution. Limitations, uncertainty and research perspectives of the new approach are discussed.

A expansão do tecido urbano e o adensamento das cidades têm um impacto negativo sobre os recursos hídricos, tanto na quantidade quanto na qualidade das águas no ambiente, pois aumenta as cargas de poluentes no meio e altera o ciclo hidrológico natural, criando riscos à população. É possível reduzir tais riscos através do diagnóstico, planejamento e gestão adequada das áreas de risco para a proteção civil e das comunidades. A abordagem ecohidrológica, que considera a relação funcional entre hidrologia, sistemas aquáticos e sua biota na escala de bacia hidrográfica, incorporando aspectos quantitativos e qualitativos da água em uma visão de interdependência entre Ecologia e Hidrologia, permite o diagnóstico, planejamento e manejo adequado dos cursos hídricos em benefício tanto dos humanos quando dos demais seres vivos. Este trabalho busca investigar a dinâmica ecohidrológica no espaço e no tempo da Bacia do Rio Monjolinho, localizada no Município de São Carlos (SP), no contexto de gestão de riscos de desastres hidrológicos. Os processos quantitativos são investigados por meio de simulações com o SWMM (Storm Water Management Model) e os dados de simulação são utilizados, por vez, na determinação do IP (Índice de Perigo) de pontos estratégicos da bacia hidrográfica e também das áreas com maior risco de inundação. Com relação à qualidade, onze variáveis clássicas de qualidade dágua são determinadas experimentalmente em 15 pontos da bacia a fim de caracterizar a dinâmica das cargas de poluentes, permitindo desta maneira a avaliação do risco biológico na bacia. Os resultados quali-quantitativos mostram que os vales dos rios oferecem riscos tanto devido às inundações quanto devido ao alto risco de contaminação. / The urban tissue expansion and the urban area densification have a negative impact on water resources, in terms of quality and quantity of available water, since it increases the polutant load at the same time it changes the natural hydrological cycle, exposing population to risk. It is possible to reduce the risks by means of assessment, planning and correct management of risk areas in order to protect the communities. The ecohydrology, which takes into account the functional relationship between hydrology, aquatic systems and biota in watershed scale, considering quali-quantitative aspects and their interdependecies, enable us to assess, plan and manage risks in an advantageous way for humans and living beings as well. This work investigates the ecohydrology dynamics in space and time of Rio Monjolinho basin, located within the municipality of São Carlos (SP), Brazil, in the context of disaster risk management. The quantitative aspects are investigated using the SWMM (Storm Water Management Model) simulation model, and the simulation data generated are used to calculate the Risk Index (RI) and to map the flooding risk areas in the basin. Eleven classic water quality variables are experimentaly determined to assess the polutant load dynamics and its distribution in the sub-basins, enabling us to assess the biologic risks. The results show that some areas in the catchment are not just flood risk areas but also areas of high biological risk of contamination.

The presence of fractures, joints and karst features within rock strongly influence the hydraulic and mechanical behavior of a rock mass, and there is a strong desire to characterize these features in a noninvasive manner, such as by using ground penetrating radar (GPR). These features can alter the incident waveform and polarization of the GPR signal depending on the aperture, fill and orientation of the features. The GPR methods developed here focus on changes in waveform, polarization or texture that can improve the detection and discrimination of these features within rock bodies. These new methods are utilized to better understand the interaction of an invasive shrub, Juniperus ashei, with subsurface flow conduits at an ecohydrologic experimentation plot situated on the limestone of the Edwards Aquifer, central Texas. First, a coherency algorithm is developed for polarimetric GPR that uses the largest eigenvalue of a scattering matrix in the calculation of coherence. This coherency is sensitive to waveshape and unbiased by the polarization of the GPR antennas, and it shows improvement over scalar coherency in detection of possible conduits in the plot data. Second, a method is described for full-waveform inversion of transmission data to quantitatively determine fracture aperture and electromagnetic properties of the fill, based on a thin-layer model. This inversion method is validated on synthetic data, and the results from field data at the experimentation plot show consistency with the reflection data. Finally, growing hierarchical self-organizing maps (GHSOM) are applied to the GPR data to discover new patterns indicative of subsurface features, without representative examples. The GHSOMs are able to distinguish patterns indicating soil filled cavities within the limestone. Using these methods, locations of soil filled cavities and the dominant flow conduits were indentified. This information helps to reconcile previous hydrologic experiments conducted at the site. Additionally, the GPR and hydrologic experiments suggests that Juniperus ashei significantly impacts infiltration by redirecting flow towards its roots occupying conduits and soil bodies within the rock. This research demonstrates that GPR provides a noninvasive tool that can improve future subsurface experimentation.

Growth of riparian cottonwoods along regulated rivers can be limited by water availability. In this study we associate seasonal variation of environmental conditions and stream flows with water relations of a natural cottonwood grove located along a regulated river in southern Alberta. To link elements in the river-soil-plant-atmosphere continuum, river and groundwater levels and precipitation were monitored; sap flow was continuously measured with thermal dissipation probes in eight trees and stomatal conductance and leaf water potential were measured monthly; and weather conditions were monitored. From June through August, stomatal conductances at both leaf and canopy levels were increasingly limited by decreasing water availability. Artificially increasing the soil moisture in August resulted in an increase in sap flow and stomatal conductance at leaf and canopy levels. These responses can be attributed to seasonal changes in the water potential difference between soil and leaves or an alteration in hydraulic conductance, or a combination of both. / xiv, 135 leaves : ill. ; 29 cm