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In an initial research proposal of December 1969, the scientists of the Coniferous Forest Biome (CFB), an ecosystem study centered in the Pacific Northwest and part of the larger International Biological Programme (IBP), expressed optimism that computer simulations and systems modeling could transform empirical knowledge of the carbon, water, and nutrient flows turned into viable forest management practice. The CFB's strategy aimed to use projections of the computer simulations and data from field study to constantly check and direct each other, resulting in a flexible, refined, and accurate understanding of forest ecosystems, as well as a reliable guide to forest management. To what extent did the CFB's research program, centered on a total system model, complete its cycle of field study, modeling, and validation? Despite the innovative strategies of the CFB modelers, ecosystem modeling lost its preeminent status among the goals of the CFB, due to different interpretations of the purpose and philosophy of ecosystem modeling and the practical limitations of administering a large research program. Instead, small field-based studies during the CFB yielded a number of ground-breaking discoveries. Although they diverged from the modeling objectives, these areas of fieldwork emerged from questions the forest's functions and cycling processes that the modeling efforts of the CFB required. Focusing on the work of CFB participants from Oregon State University and the USDA Forest Service in the H. J. Andrews Experimental Forest, this thesis addresses the relationship between the marginalization of the modeling objectives and the rising centrality of field-based forest studies in the CFB from 1969 to 1980. Given the ongoing legacy of CFB research at the Andrews Long-Term Ecological Research (LTER) site and the later implications of CFB findings in debates over forest policy and management, this thesis also seeks to evaluate the Coniferous Forest Biome as a whole and discuss the role of modeling and field work within large ecological research endeavors more generally. / Graduation date: 2006

Management, under the concepts of multiple-use and adaptive management, requires the assessment of potentials and limitations of the natural ecosystems to provide satisfaction to human needs, to protect long term productivity, and preserve biological diversity. Overstory-understory relationships were developed for ponderosa pine (Pinus ponderosa) ecosystems to help managers to evaluate herbage production potentials. Secondary data sources from the Beaver Creek and the Heber Watersheds were divided as follows: igneous soils, igneous clay loam, igneous loam-sandy loam, sedimentary soils, alluvium, and sandstone. Regression models were fitted to the raw data by the least squared method. The dependent variables were herbage production (lb/ac) by component; namely total, grass and grass like plants, forbs and half shrubs, and shrubs. The independent variables were total and ponderosa pine basal area (ft 2 /ac). Semilogarithmic models fitted the data from igneous soils, while logarithmic transformations of hyperbolic models fitted the data from sedimentary soils. For igneous soils, ponderosa pine basal area suffice for adequate predictions of herbage production. Significant differences were found between equations for alluvium and sandstone. The resulting equations for ponderosa pine integrate the core of the model UNDER. Mathematical functions developed elsewhere are included in UNDER to compute herbage production in pinyon-juniper and mixed conifer ecosystems. UNDER is linked to other simulators by MICROSIM. MICROSIM, a multiple-resource simulation model, is a tool to assist in the assessment of potentials of forest and woodlands of Arizona. MICROSIM is a menu driven program for IBM or compatibles it contains the module Flora, for estimating plant responses, and module Fauna, to evaluate impacts on animals. Further development of MICROSIM should include the linkage to more modules and models, and to Geographical Information Systems.

This thesis presents the results of a study to examine long-term forest dynamics in the high-altitude mountains of West-Central Mexico. Vegetation dynamics on temporal scales ranging from 10² to 10³ years were reconstructed in order to provide essential information on the temporal variability of ecological patterns and processes in these forests; information that is of direct relevance for their current and future conservation and management strategies. Vegetation and palaeoecological methods undertaken included fossil and modern pollen analysis, vegetation surveys, microfossil charcoal analysis, magnetic susceptibility, inorganic and organic geochemistry, radiocarbon and ²¹⁰Pb dating. These were used to evaluate the long-term dynamics of three forest types; Pine Forest, Cloud Forest and Transitional Forest on timescales spanning the past 4260, 1340 and 1230 years respectively. The main drivers of change were climate and disturbance events induced by climate fluctuations, for example increased fire frequency. The reconstructed records indicate that the sequences from the Cloud Forest and the Transitional Forest spanned two wet and one dry climatic interval while the Pine Forest sequence spanned two dry and two wet periods. The impact of these climatic fluctuations was significant on all three forest types and resulted in variations in forest diversity, taxonomic turnover and successional change. The climate change episodes observed in these records seem to be the local manifestation of climatic events that were occurring throughout Mexico at these intervals in time. Human influences were evident in the three forests through the appearance of cultural taxa, particularly during the driest period (~ 1200 yr BP). There is little evidence from these records, however, to suggest a widespread clearance of the landscape for agriculture. Results from this study support the current conservation and management recommendations for Cloud Forest to exclude timber extraction, grazing and agricultural activities from this forest type. In the Pine Forest, human interventions such small-scale agriculture, prescribed burning and silvicultural actions are in agreement with the longterm pine ecology and as such, total exclusion of human activities is not necessary. For the Transitional Forest, results from this study suggest that there needs to be the establishment of adequate plans to reduce frequent fires to arrest the development of prone-to-fire taxa.

This paper explores the characterization of a riparian bottomland forest in north central Texas in two ways: field study, and computer simulation with the model ZELIG. First, context is provided in Chapter One with a brief description of a southern bottomland forest, the ecological services it provides, and a history of bottomland forests in Texas from the nineteenth century to the present. A report on a characterization study of the Lake Ray Roberts Greenbelt forest comprises Chapter Two. The final chapter reviews a phytosocial study of a remnant bottomland forest within the Greenbelt. Details of the ZELIG calibration process follow, with a discussion of ways to improve ZELIG's simulation of bottomland forests.

Effective stewardship of forest ecosystems make it imperative to measure, monitor, and predict the dynamic changes of forest ecology. Measuring and monitoring provides us a picture of a forest's current state and the necessary data to formulate models for prediction. However, societal and natural events alter the course of a forest's development. A simulation environment that takes into account these events will facilitate forest management. In this thesis, we describe an efficient parallel implementation of a land cover use model, Mosaic, and discuss the development efforts to incorporate spatial interaction and succession dynamics into the model. To evaluate the performance of our implementation, an extensive set of simulation experiments was carried out using a dataset representing the H.J. Andrews Forest in the Oregon Cascades. Results indicate that a significant reduction in the simulation execution time of our parallel model can be achieved as compared to uni-processor simulations.

White-tailed deer are ecosystem engineers that dramatically alter forest understory vegetation. Consequently, deer can impact many species in a forest through both direct and indirect effects. One species that deer may indirectly affect is the gypsy moth, whose pupae are preyed upon by the white-footed mouse. Through alterations to understory habitat of mice, deer may reduce mouse predation on gypsy moth pupae. In this study, I tested for indirect effects of deer on the gypsy moth by comparing mouse abundance, vegetation properties, and predation on pupae inside, and outside, of long-term deer exclosures. Overall, I did not find evidence for indirect effects of deer on the gypsy moth. There was little effect of the exclosures on mouse abundance, predation rates, and habitat measures. High mouse abundances, which likely resulted from a large acorn mast the previous year, may be obscuring indirect effects that would be detected at lower mouse abundances.

This study explored the application of compost onto a small loblolly pine tree forest in northeast Texas. Its purpose was to determine if the application of various amounts of compost would provide for accelerated rates of growth for the trees. Soil parameters were also monitored. A total of 270 trees were planted and studied in a northeast Texas forest ecosystem. Compost rates of 5, 25, and 50 tons per acre with either soil or compost backfill were utilized and compared to a control without compost. Nonparametric and parametric ANOVA and Chi-Square tests were utilized. The results indicated that greater application rates retained greater moisture and higher pH levels in the soil. Compost applications also yielded a greater survival rate as well as larger tree height and diameter when compared to the control. The 25 ton/acre application backfilled in native soil achieved the greatest average in height and diameter when compared to the averages for the control plot. Greater growth differences for the 25S application can be attributed to additional nutrients coupled with a stable pH consistent with native soil acidity.

The present single species geometric forest plantations in the Scottish landscape suggest that foresters were not interested in conservation issues and landscape aesthetics. This thesis argues that the appearance of the forests is not so much the result of the foresters' lack of interest in conservation and nature but the social, economic and political pressures that underpinned their creation as well as the Scottish physical environment. Scottish foresters have had a long-standing interest in conservation issues that dates back to the colonial roots of Scottish forestry in mid- 19th century India. The concept of conservation was introduced in Scotland through foresters returning from their service in India and other parts of the Empire. The root of the interest in landscape aesthetics dates back even to the 18th century when Scottsh landowners started to plant trees, both exotic and native, to beautify their estates. By the second half of the 19th century influential landowners became concerned about the fact that Scotland could not produce sufficient timber to provide for its own needs. They also thought that forestry could provide jobs in the Highlands of Scotland and thus contribute to strengthening the social and economic fabric of rural Scotland. To increase timber production and improve the rural economy, influential landowners lobbied for the creation of a forestry agency. It was from these roots - aesthetics, conservation and social and economic concerns - that forestry policy in Scotland developed. It was only after the First World War, when Britain was confronted with severe timber shortages, that a state forestry organisation, the Forestry Commission, was created. Its initial task was to create a strategic timber reserve but over time conservation objectives came on board forest policy. The lands available for forestry were poor upland areas where only a handful introduced conifers were able to survive the harsh condition and, because of their fast growth, created in a relatively short time-span the desired timber reserve. It was for this reason that the forests created by the Forestry Commission were mainly made up of fast growing conifers introduced from the Pacific coast of North America. Technical improvements such as the introduction of mechanical ploughing and the use of fertilisers expanded the range of planting and pushed planting even further uphil. The coincidence of ploughing and the use of conifers on a large scale also led to an increasing monotonous appearance of the new plantations. It was this monotonous and artificial appearance that attracted the first opposition to the planting of conifers in the Lake District, but not in the Scottish Highlands. The development of an environmental policy as part of state forestry in Scotland was not so much driven by external pressures from conservation organisations but by a combination of economic and social pressures in the Highlands and the fact that many foresters ar.e sensitive to the environment in which they work. The general public and nature conservation organisations were until the 1970s not much concerned about the emergence of coniferous plantations in the landscape. Other more pressing environmental issues, such as the impact of build structures on the landscape, the use of herbicides, and the creation of nature reserves, occupied public environmental concern and nature conservation organisations alike. In the meantime the Commission developed the fundaments on which the broadleaf and conservation policies of the 1990s became based. It was the pressure from the Treasury and the wood processing industry that made it hard to change direction because hard economics were dominant. But when change came, the Commission was able to adapt to the new situation thanks to the deep-rooted interest of its staff in nature conservation.

Long term fire exclusion has been attempted in Catchment IX (CIX) at Cathedral Peak. Baseline vegetation of CIX was sampled in 1952, with follow-up surveys in 1973, 1986 and 2010. These took place at key times in terms of changes in fire history within the catchment. Complete fire exclusion was achieved between 1973 and 1986, but eight accidental fires burnt part of CIX thereafter. The woody component was resurveyed in 2010 after the latest of thirteen unintended fires had swept through CIX in 2007. This fire burnt about 90% of the catchment. The vegetation of the area not burnt was a distinct community and size structure indicating it had not been burnt by other accidental fires. Partial exclusion of fire over 58 years resulted in vegetation transforming from grassland to a grassland-fynbos-scrub forest mosaic. Erica evansii and Leucosidea sericea, a reseeder and resprouter respectively, were the two dominant species in CIX. These displayed expected responses to a single fire, resulting in dominance shifting from E. evansii (92% mortality) to L. sericea (1.6% mortality). The decrease in E. evansii individuals resulted in a relative increase in community contribution of species not affected by fire. Leucosidea sericea’s post-fire dominance in burnt plots was not apparent in fire-protected areas. A successional trend of colonisation of woody species, predominantly E. evansii and L. sericea, into grassland was observed. Despite occasional fires since 1986 vegetation did not revert to grassland. The emergent woody community was not homogenous. This was attributed to a combination of an irregular pattern of accidental burns and environmental variability within the catchment. The mosaic of distinct grassland, woodland, ecotonal and scrub forest communities are predicted to remain as such.