Crested wheatgrass arrived in North America at the turn of the twentieth century through the foreign plant exploration missions sponsored by the United States Department of Agriculture. During the first two decades of the new century, scientists tested the grass at agricultural experiment stations. They determined it was useful for grazing and particularly valuable because it could grow in drought conditions with little or no care and would continue to produce high quality feed even after several years of heavy use. Beginning in the 1930s federally sponsored land utilization and agricultural adjustment programs sponsored the use of crested wheatgrass for soil conservation and weed control. The grass protected the soil on the land that had been entered into the acreage reserves and the conservation reserves programs of the federal soil bank. Also in the late 1930s and through the 1960s, rangeland managers used crested wheatgrass to improve forage productivity on public lands that were used for grazing. By the 1970s somewhere between 12 and 20 million acres of crested wheatgrass grew in North America in eleven western states, and in Saskatchewan and Alberta. By 1980 attitudes about agriculture and wilderness had changed in the United States and land management was focused on multiple uses and on protecting ecosystems and native species. Attitudes about grazing and agricultural landscapes had changed and many preferred nonagricultural landscapes and land uses. As a result, crested wheatgrass went from being considered one of the most valuable plants in North America to being considered an invasive weed, in some quarters. Debates in the last 25 years have tried to determine if, where, and how crested wheatgrass belongs in North America. This thesis explains the discourses, or interest groups, that are participating in the current conversation. One impulse is to use empirical evidence to determine whether or not introduced plants like crested wheatgrass belong, but the main contention of this thesis is that empirical studies alone will always be insufficient measures because belonging is also a subjective and experientially or emotionally derived measure.

The invasive species Saltcedar is affecting water and land resources throughout the western states of America. Because of great water use capabilities and other ecosystem detriments, Saltcedar has been targeted for treatment. For successful management of Saltcedar, individual landowners need to be aware of the costs and benefits of treating Saltcedar. Eleven of the most commonly reported treatment methods were evaluated for firm level economic feasibility. Evaluated on the basis of treatment cost, treatment effectiveness, Saltcedar water-use, and re-vegetation water-use, a production plan of ten years was created for each treatment method. Some treatment methods required re-treatment and were evaluated with re-treatments most commonly found in the literature. Of the treatment methods evaluated, five treatment methods were determined to be most feasible. Landowner valuation of environmental changes in the ecosystem brought on by Saltcedar is very important in the decision of which treatment method to implement. Personal valuation, over a period of ten years, will often determine which treatment methods are most cost efficient.

My thesis focuses on different ranch management practices and the effects they may have on bird communities in northern Texas. Specifically, I address the potential differences between ranches enrolled in the Audubon Conservation Ranching program versus ranches that are not enrolled. Each research chapter is meant to be an individual publication. Chapter 2 compares and contrasts two different ways to survey bird communities in prairie systems. Chapter 3 compares ranch management techniques, prairie health indicators, and bird communities. This study aims to observe any mechanistic drivers of rangeland health that may elicit a positive bird assemblage response.

Doctor of Philosophy / Department of Animal Sciences and Industry / K C Olson / The objective of this series of studies was to examine select factors that affect behavior of beef cattle grazing native-tallgrass rangelands. Mineral supplements with divergent palatability characteristics were delivered to beef cows grazing native tallgrass range during various seasons of the year in order to measure mineral intake, frequency of supplement use, and duration of supplement use. We concluded that molasses-based mineral supplements influenced the activities of grazing cows more strongly than salt-based mineral supplements. These influences extended to the amount of supplement consumed as well as to the frequency, duration, and timing of use. Diet selection preferences of experienced, multiparous beef cows and naïve, primiparous beef cows grazing dormant, native tallgrass pastures were examined also during a short-term winter grazing bout. Naïve, primiparous cows selected more forbs and fewer grasses than experienced, multiparous cows. Previous research indicated that preference for broadleaf plants generally increased with grazing experience; however, these conclusions were based on research with greater-quality forages than those evaluated in our study. The differences in diet selection patterns between experienced, multiparous cows and naïve, primiparous cows during a short-term winter grazing period could be indicative of differences in long-term foraging strategies. In addition, the botanical composition of diets grazed by lactating beef cows with suckling calves and non-pregnant, non-lactating beef cows grazing either burned or unburned native tallgrass prairie during summer were evaluated. There were no differences in botanical diet composition between lactating cows suckling calves and non-lactating cows. In contrast, total graminoid selection was greater on burned (74.2%) than unburned pastures (71.8%) and total forb selection was greater on unburned (28.2%) than burned pastures (25.8%). We interpreted these data to suggest that forage selection preferences of beef cows can be altered with spring burning of native tallgrass pastures. Effects of large, round hay bale feeding method on intake of smooth bromegrass hay and eating behavior by beef cows were examined on dormant tallgrass prairie pastures during winter. Three large, round hay bale-feeding systems were evaluated: bales fed in a ring feeder, bales unrolled on the ground, and bales chopped with a flail-type hay processor (20-cm particle length) and deposited on the ground. Hay intake, hay refusal, frequency of use, and duration of use were not influenced by hay-feeding method. Foraging behaviors of beef cows in our studies were influenced by supplement type, cow age, and prescribed burning of rangeland. Conversely, foraging behaviors of beef cows were not influenced by lactation or by hay-feeding method.

Master of Science / Department of Agronomy / Walter H. Fick / Rough-leaved dogwood (Cornus Drummondii Meyer) is one of the most invasive woody plants in rangeland of Kansas. Reduced prescribed burning due to drought and urban encroachment probably has contributed to its spread. Herbicides are commonly recommended for control of rough-leaved dogwood, but minimal data exists for recommendation development. Ten herbicide treatments were applied in June during late flowering at two locations in northeast Kansas in 2005 and 2006. Each treatment was replicated three times in a randomized complete block design with individual plot sizes of 3 x 3 m. Herbicides were applied with hand sprayers in 1017 L/ha solution. Visual evaluations of defoliation were made about 1 and 12 months after treatment (MAT) and mortality was estimated about 1 year after treatment (YAT). Defoliation 1 MAT varied among herbicides with significant location by year and herbicide by year interactions. Treatments providing greater than 70% defoliation 12 MAT both years were triclopyr + 2,4-D (1.12 + 1.06 kg ae/ha), triclopyr + fluroxypyr (1.83 + 0.61 kg ae/ha), picloram + fluroxypyr (0.41 + 0.41 and 0.82 + 0.82 kg ae/ha), and picloram + 2,4-D + triclopyr (0.66 + 2.44 + 2.44 kg ae/ha). Rough-leaved dogwood is difficult to control with a single herbicide application, but treatments exist that will substantially reduce stands. Tebuthiuron pellets (Spike 20P) are another control measure recommended for rough-leaved dogwood. Treatments of 4.4 kg ai/ha (3/4 oz per 100 square feet) tebuthiuron pellets were applied in December 2004. A visual estimate of control indicated tebuthiuron reduced dogwood cover by 65% compared to a 3% decrease on untreated plots. Dogwood density was reduced by 2.2 stems/m2 (P<0.08). Total woody plant cover increased on untreated plots by 6.2 percentage units, but was decreased by 20.9 percentage units on tebuthiuron treated plots. Other woody plants decreased in both treated and untreated plots. Shading by a large elm tree likely caused variation between replications including increases in cover and density of rough-leaved dogwood on treated plots. Tebuthiuron is a photosynthetic inhibitor that often is not effective on shaded plants. Tebuthiuron pellets applied at 4.4 kg ai/ha was an effective control option for unshaded rough-leaved dogwood.

Master of Science / Department of Agronomy / Walter H. Fick / Patch-burn grazing is a relatively new concept in terms of rangeland management. While numerous benefits have been associated with this system, in the tallgrass prairie of Kansas, cattle production and sustainability of rangeland are critical. In 2006, 253 ha at the KSU Bressner Range Research Unit in Woodson County, Kansas were subjected to spring patch-burn grazing (using one-third portions) and traditional full-burn grazing. Each treatment within the split-block design was replicated four times for 3 years. The objectives were to evaluate whether livestock performance would be compromised under this grazing system, to monitor the health of the rangeland, and to observe the usefulness of this tool as a potential control of the invasive plant sericea lespedeza [Lespedeza cuneata (Dumont) G. Don]. In regards to cattle performance, burn treatments had no significant difference in average daily gain (p≥0.10) in any of the 3 years. On average, cattle utilized 61% of the current year’s forage production in patch-burned portions, which was higher (p≤0.10) than that of unburned (30%) and full-burn (41%) portions. Results of the botanical composition show forb and woody plant composition did not differ between treatments, however differences (p≤0.10) were present in grass composition. Total annual grasses increased 19.1 percentage units under patch-burn and 2.1 units under full-burn, while total perennial grasses decreased 18.4 and 1.1 units, respectively. When evaluated by treatment area (one-third portion), results indicated that the 3-year cycle did allow enough time for recovery. At 2 years after treatment (2-YAT), no significant difference in composition(p≥0.10) was present between initial patch-burn portions and the full-burn pastures. Finally, in only 1 year of the study did cattle statistically consume a greater percentage of sericea lespedeza plants (p≤0.10) in the patch-burned portions (92%) than in full-burned pastures (35%). Biomass utilization did not differ (p≥0.10) between burn treatments. Surprisingly, there was a trend for the number of plants in the sampled areas of the patch-burn portions to decrease throughout the cycle. However, at the conclusion of the 3-year cycle, sericea densities did not differ(p≥0.10) between treatments. Patch-burn grazing shows promise as a potential management tool for Kansas land managers.

Thesis (MSc)--University of Stellenbosch, 2000. / ENGLISH ABSTRACT: The composition and state of soil-stored seed banks on- and off the mesa Tafelberg, in the Nama Karoo rangelands of the Eastem Cape, South Africa, were investigated within the context of a broader restoration ecology project "Restoration of degraded Nama Karoo rangelands: the role of conservation islands'". The premise for this seed bank study was that restoration of degraded semi-arid rangelands is possible through applied management programs based on the methodology and practice of ecological restoration. Broadly acknowledged properties of non-equilibrium environments (e.g. unpredictable climates and varying degrees of disturbance) and soilstored seed banks (e.g. spatial and temporal distributions) formed the basis for investigating. the general environment and the properties.. of existing seed, banks, in the Tafelberg locality. The potential role of hills as refugia for palatable plant species was an under-lying element of the investigation. Following a brief investigation, of historical and contemporary research and policy on rangeland degradation in semi-arid regions of the world, the fundamental need for comprehensive and applied seed bank research in the Nama Karoo is emphasised. Within the framework of the umbrella project, the seed bank study examined local environmental criteria commencing with an investigation into seed bank- and vegetation habitats. Chemical and physical properties of soils from twenty two sites on- and off Tafelberg were described. Substantial soil habitat variation, between the top, the north west slopes and plains and the south east slopes and plains of Tafelberg, was identified. Micro-site variation between open-canopy (interplant spaces) and closed-canopy (under plant cover) microhabitats was found to be significant Primary soil habitat differences were linked to soil organic matter content which was found to be low on the plains relative to the top and slopes; and, low in open-canopy micro-sites relative to closed-canopy sites. Soil texture and nutrient composition on- and off Tafelberg was found to be highly variable with significant differences between the top and the plains as well as between the north west and south east plains. The slopes were found to be intermediate (showing some level of gradient) between the top and the plains. Soil nutrient variation was interpreted as a function of textural and parent-material properties of soils. Anthropogenic factors for accelerated erosion, deposition, leaching and salt-crust formation were also considered. It was concluded that while properties of soils on- and off Tafelberg are inherently related to parent soils, changes to textural and nutrient properties may be occurring and these changes may have been exacerbated by high levels of grazing. An investigation (focusing on small shrubs) of plant phenological response, in relation to rainfall and grazing gradients, identified trends of peak budding- and flowering seasons following rainfall during summer and autumn respectively. A continuum of seeding activity, with peaks in late autumn and early winter, was construed from quarterly data. Flush vegetative growth was noted for most small shrubs during spring, autumn and summer surveys. Since some form of activity related to reproductive output (flush growth, budding, flowering or seeding) was apparent at almost all times of the year, it is argued that high intensity disturbance (including grazing) might impact negatively on plant survival, leading to reduced reproductive input (i.e. seeds) into future generations. Plant communities on the top and plains are described noting significant differences between vegetation on top of Tafelberg (comprising primarily high production, palatable grass and shrub species) and that on the plains (comprising mainly spinescent, ephemeral, toxic and low production species). Given the high grazing pressure on the plains (relative to the less utilised slopes and top of Tafelberg), differences in vegetation composition are discussed in relation to studies elsewhere that describe degraded rangelands. It is concluded firstly that the plains surrounding Tafelberg are degraded, secondty that long term over-utilisation has altered vegetation composition and finally that inter-grazing rest periods of three- or six months alone may not restore vegetation diversity nor desirable plant species to the plains. Results of germination trials (investigating soil-stored seed banks) from two sampling episodes (spring and autumn 1998) revealed that seed banks in soils removed from the plains, slopes and top of Tafelberg followed distribution patterns observed in above-ground vegetation. Species-specific data was not finalised for this thesis since not all seedlings matured and flowered within given time constraints. In order to compare seedling emergence data, plant categories were developed that distinguished ephemerals versus persistent (perennial) species and dicotyledonous species versus grasses and other monocotyledonous plants. Samples from the top and the middle to upper slopes showed a high percentage of palatable and persistent grass and shrub species present in soil-stored seed banks while over twothirds of plants germinating from plains' soil samples were ephemeral species and most of these were both tiny «Scm) and short-lived «3 months). Most of the perennial species germinating from plains' samples were seedlings of Pentzia incana, Chrysocoma ciliata and succulents (mostly Mesembryanthemaceae) but few perennial grasses were present. On the other hand, roughly 94% of seedlings germinating from samples from the top and 63% of seedlings germinating from two slopes of Tafelberg respectively were persistent- grass or shrubby species. Roughly 89% of ephemeral species recorded from slopes' samples germinated from the lowest footslope sites. Seed densities were closely linked to micro-habitats with roughly three-fold differences between open- (lower seed density) and closed-canopy (higher seed density) micro-sites. Multivariate analysis of variance (MANOVA) indicated that most of the variation in seed density data was explained at the micro-habitat level. Seasonality played a secondary role with significant twoway interaction between the two effects. It is concluded that both factors must be considered when developing restoration programmes that aim to improve both overall plant cover and improved plant species diversity. While the original or "pristine" state of vegetation composition and seed banks on the plains is unknown, concems were raised regarding the apparently degraded state of total above- and below-ground plant diversity on these plains. A likely consequence of habitat degradation is that species with specific soil-, nutrient ratio-, aspect- and altitudinal requirements from the top and slopes of Tafelberg (as well as from nearby plains' refugia) may not be able to establish in degraded habitats on the plains. The identification of pioneer plant species (tolerant of habitat degradation) that allow increased vegetation cover and safe-sites for seedling germination of desirable plant species is recommended. Restoration programmes will need to be coupled with strict grazing management principles that allow seedling germination. establishment and successful reproductive output of desirable plants for future rangeland regeneration. Common and salient features of the soil habitat, plant pbenological response and germination trial studies are brought together in an examination of habitats and related seed bank diversity on- and off Tafelberg. Acknowledging the brevity of this research study, but utilising case studies from elsewhere and integrating both lines of questioning, the conclusion is again reached that the plains surrounding Tafelberg are degraded through decades and probably centuries of grazing by domestic livestock. It is considered crucial that restoration through improvement (or rehabilitation) of habitats and increased seedling safe-sites be considered. Both climate and grazing management appear to play an irrevocably linked role in shaping vegetation composition in rangelands. While rangelands are intrinsiCalfy adapted to surviving extremes of climatic variability found in non-equilibrium regions it is argued that the impacts of grazing, particularly during times of drought and climate change, are slowly reducing the intrinsic "buffer-capacity" of rangelands to withstand these changes and extremes. It seems apparent from research elsewhere that restoration through passive management is slow and probably not economically achievable within a viable time frame. The instifution and promotion of integrated and strategic programmes that identify and address issues of land degradation and land use change in semi-arid rangelands is recommended. The input and endeavours of different authorities, ministries and a broad public participation incentive are encouraged in these proposed programmes in order to ensure broadly-based input into long term sustainability and conservation of the considerable biological diversity of these regions. Seed bank assessment is considered to be a valuable means of indicating restoration potential and ranqeland condition with potential for the identification of both degraded and conservationworthy areas. Finally, some limitations and challenges of this study are examined through a process of firstly identifying alternative approaches to research methodologies and secondly through proposing recommendations for future research projects. While alternative methods could have been applied for the purposes of accomplishing this study it is concluded that, within the given time- and other constraints, the appropriate methods were applied. / AFRIKAANSE OPSOMMING: Die samestelling entoestand van groncbbewaarde saadbanke.op en van die mesa Tafelberg af, in die Nama Karoo weiveld van die Oos-Kaap, Suid-Afrika, is ondersoek binne die konteks van 'n wyer herstelekologieprojek: "Restourasie van oorbeweide Nama Karoo weiveld: die rol van bewannqseilande'". Die uitqanqspunt van hierdie saadbankondersoek was dat dit moontlik is om beskadigde semidroë weiveld te herstel deur middel van toegepaste bestuursprogramme wat gebaseer is op die metodologie en praktyk van ekologiese herstel. Algemeen erkende eienskappe van nieekwilibrium- omgewings (byvoorbeeld onvoorspelbare klimate en wisselende mates van versteuring) en grond-bewaarde saadbanke (byvoorbeeld ruimtelike en temporale verspreidings), het die basis gevorm vir die ondersoek van die algemene omgewing en die eienskappe van bestaande saadbanke in die Tafelberg omgewing. Die potensiële rol van heuwels as skuilplek vir aanvaarbare plantspesies was 'n onder1iggende element van die ondersoek. Na 'n kort ondersoek van historiese ~n kontemporêre navorsing en beleid oor weiveldbeskadiging in semi-droë streke van die wêreld, is die fundamentele behoefte aan omvattende en toegepaste saadbanknavorsing in die Nama Karoo beklemtoon. Binne die raamwerk van die oorkoepelende projek het die saadbankondersoek plaaslike omgewingskriteria bestudeer, beginnende met 'n ondersoek na saadbank- en plantegroeihabitats. Chemiese en fisiese eienskappe van grond vanaf twee en twintig terreine op en van Tafelberg af is beskryf. Omvattende grondhabitatwisseling tussen die kruin, die noordwestelike hange en die vlaktes en die suidoostelike hange en vlaktes van Tafelberg is geïdentifiseer. Mikroterreinwisseling tussen die oop-dak (tussenplantruimtes) en toe-dak (onderplantdekking) -mikrohabitats is as beduidend bevind. Primêre grondhabitatverskille is gekoppel aan die inhoud van die grond se organiese materiaal, wat op die vlaktes as laag bevind is vergeleke met die kruin en hange; en laag bevind is in oop-dak-mikroterreine vergeleke met toe-dak-terreine, Daar is gevind dat die grondtekstuur en voedingstofsamestelling op en van Tafelberg af aansienlik varieer, met beduidende verskille tussen die kruin en die vlaktes, en ook tussen die noordwestelike en suidoostelike vlaktes. Daar is gevind dat die hange intermediêr is (toon 'n mate van gradiënt) tussen die kruin en die vlaktes. Die wisseling in grondvoedingstowwe is vertolk as 'n funksie van teksturele en ouermateriaaleienskappe van grond. Antropogeniese faktore vir versnelde erosie, neerslag, loging en soutkorsvorming is ook oorweeg. Die gevolgtrekking is gemaak dat terwyl die eienskappe van grond op en van Tafelberg af inherent aan ouergrond verwant is, kan veranderinge aan tekstuur en voedingstofeienskappe voorkom, en hierdie veranderinge kon deur hoë vlakke van weiding vererger gewees het. 'n Ondersoek (wat op klein struike gefokus het) van plantfenologiese reaksie met betrekking tot reënval en weidingsgradiënte het tendense van piekbot- en blomseisoene na reënval gedurende die somer en herfs onderskeidelik geïdentifiseer. 'n Kontinuum van saadskietaktiwiteit, met piektye in laat herfs en vroeë winter, is van kwartaallikse data saamgestel. Groeistuwing is vir die meeste klein struike waargeneem gedurende lente-, herfs- en someropnames. Aangesien die een of ander vorm van aktiwiteit met betrekking tot voortplantingsgroei (groeistuwing, bot, blom of saadskiet) op feitlik al die tye van die jaar sigbaar was, word daar geredeneer dat hoëintensiteitversteuring (ook weiding) 'n negatiewe impak op plantoorlewing kan hê, wat sal lei tot verminderde voortplantingsinset (m.a.w. sade) in toekomstige geslagte. Plantgemeenskappe op die kruin en vlaktes word beskryf met beduidende verskille tussen plantegroei op die kruin van Tafelberg (wat hoofsaaklik bestaan uit hoëproduksie, smaaklike gras- en struikspesies) en dié ap die vlaktes (wat bestaan uit hoofsaaklik doringagtige, efemere, toksiese en laeproduksie-spesies). Gegee die hoë weidingsdruk op die vlaktes (vergeleke met die minder benutte hange en kruin van Tafelberg), word verskille in die samestelling van plantegroei bespreek met betrekking tot studies elders wat beskadigde weiveld beskryf. Die gevolgtrekking word eerstens gemaak dat die vlaktes om Tafelberg beskadig is, tweedens dat langtermyn-oorbenutting die samestelling van die plantegroei verander het, en laastens dat interweiding-rusperiodes van drie tot ses maande alleen dalk nie die diversiteit van plantegroei of die verlangde plantspesies op die vlaktes kan herstel nie. Die uitslae van kiemingstoetse (wat ondersoek ingestel het na grond-bewaarde saadbanke) van twee steekproefepisodes (lente en herfs 1'998)het getoon dat saadbanke in grond wat van die vlaktes, hange en kruin van Tafelberg verwyder is, die verspreidingspatrone volg wat in bogrondse plantegroei waargeneem is. Spesie-spesifieke data is nie vir hierdie tesis gefinaliseer nie, aangesien nie alle saailinge binne die gegewe tydsbeperkinge gegroei en geblom het nie. Ten einde saailing-verskyningsdata te vergelyk, is plantkategorieë ontwikkel wat efemere en langdurige (meerjarige) spesies en dikotiele spesies en grassoorte en ander monokotiele plante onderskei. Steekproewe van die kruin en teen die middelste en boonste hange het 'n hoë persentasie van aanvaarbare en standhoudende gras- en struikspesies getoon wat in grond-bewaarde saadbanke teenwoordig is, terwyl meer as twee derdes van plante wat in die vlaktes se grondmonsters ontkiem het, efemere spesies was, en die meeste daarvan was klein «5cm) en met 'n kort leeftyd «3 maande). Die meeste van die meerjarige spesies wat van die vlaktes se steekproewe kom, was saailinge van Pentzia incana, Chrysocoma ciliata en vetplante (hoofsaaklik Mesembryanthemaceae), maar min meerjarige grasse was teenwoordig. Daarenteen was onderskeidelik ongeveer 94% van saailinge wat van monsters van die kruin en 63% van saailinge wat van twee van die hange van Tafelberg ontkiem het, langdurige gras- of struikspesies. Ongeveer 89% van die efemere spesies wat van die hange se steekproewe aangeteken is, het op die laagste voethang-terreine ontkiem. Saaddigthede toon 'n noue verband met mikrohabitats, met ongeveer drievoudige verskille tussen oop- (laer saaddigtheid) en toe-dak (hoër saaddigtheid) -mikroterreine. 'n Multivariaatontleding van variansie (MANOVA) het aangedui dat die meeste van die variasie in saaddigtheidsdata op die mikrohabitat-vlak verduidelik is. Seisoenaliteit het 'n sekondêre rol gespeel, met beduidende tweerigting-interaksie tussen die twee uitwerkings. Die gevolgtrekking word gemaak dat albei faktore oorweeg moet word wanneer herstelprogramme ontwikkel word wat ten doel het om algehele plantbedekking en die diversiteit van verbeterde plantspesies te verhoog. Terwyl die oorspronklike of "ongerepte" toestand van die plantegroei se samestelling en saadbanke op die vlaktes onbekend is, is kommer uitgespreek oor die klaarblyklik beskadigde toestand van die totale bo- en ondergrondse plantdiversiteit op hierdie vlaktes. 'n Waarskynlike gevolg van habitatbeskadiging is dat spesies met spesifieke grond-, voedingstofverhouding-, aspek- en seevlak-vereistes van die kruin en hange van Tafelberg (asook van nabygeleë vlakteskuilings) nie in staat sal wees om in beskadigde habitats op die vlaktes te vestig nie. Die identifikasie van pionierplantspesies (verdraagsaam vir habitatbeskadiging) wat verhoogde plantegroeidekking en veilige terreine vir saailingontkieming van verlangde plantspesies toelaat, word aanbeveel. Herstelprogramme sal gekoppel moet word aan streng weidingsbeginsels wat saailingontkieming, die vestiging en suksesvolle voortplantingsproduksie van gewenste plante vir toekomstige weiveldherstel moontlik maak. Algemene en belangrike eienskappe van die grondhabitat, plantfenologiereaksie en kiemingsproefondersoeke word saamgevoeg in 'n ondersoek van habitats en verwante saadbankdiversiteit op en van Tafelberg af. Met erkenning van die kortstondigheid van hierdie navorsingsondersoek, maar met benutting van gevallestudies van elders en die integrasie van albei vraaglyne, word daarweertot die slotsom gekom dat die vlaktes om Tafelberg beskadig is deur dekades en waarskynlik eeue se beweiding deur mak lewende hawe. Dit word as uiters belangrik beskou dat herstel deur verbetering (of rehabilitasie) van habitats en 'n groter aantal saailing beveiligingsterreine oorweeg moet word. Dit lyk asof klimaat sowel as weidingsbestuur 'n onherroeplik gekoppelde rol speel in die vorming van die samestelling van plantegroei op weiveld. Terwyl weiveld intrinsiek aangepas is by die oorlewing van uiterste klimaatswisseling wat in nie-ekwilibriese streke aangetref word, word daar geredeneer dat die impak van beweiding, veral gedurende droogtetye en klimaatsverandering, stadigaan die intrinsieke "buffervermoë" van weiveld verminder om hierdie veranderinge en uiterstes te weerstaan. Dit blyk uit navorsing elders dat herstel deur passiewe bestuur stadig en waarskynlik nie ekonomies haalbaar is binne 'n lewensvatbare tydsraamwerk nie. Die instelling en bevordering van geïntegreerde en strategiese programme wat kwessies van grondbeskadiging en verandering van grondgebruik in semi-droë gebiede identifiseer en oplos, word aanbeveel. Die insette en pogings van verskillende owerhede en ministeries en deelname deur die breë publiek word aangemoedig in hierdie voorgestelde programme ten einde insette met 'n breë basis in die langtermyn~volhoubaarheid en bewaring van die aansienlike biologiese diversiteit van hierdie streke te verseker. Saadbankbeoordeling word beskou as 'n waardevolle manier om die herstelpotensiaal en weiveldtoestande met die potensiaal vir die identifisering van beskadigde sowel as bewaringswaardige gebiede aan te dui. Laastens word 'n paar beperkings en uitdagings van hierdie studie ondersoek deur 'n proses van eerstens die identifisering van alternatiewe benaderings tot navorsingsmetodologieë, en tweedens deur die voorstel van aanbevelings vir toekomstige navorsingsprojekte. Terwyl alternatiewe metodes toegepas kon gewees het ten einde hierdie studie af te handel, is die gevolgtrekking dat die toepaslike metodes binne die gegewe tyd5- en ander beperkings toegepas is.

Thesis (MSc)--University of Stellenbosch, 2002. / ENGLISH ABSTRACT: Arid lands comprise some 47.2 % of the world's land surface. Between 32 % and 35 % of these drylands are subject to some form of desertification and land degradation. South Africa is an arid country with water being a major limiting resource. The Nama-Karoo biome is an extensive tract of semi-arid vegetation comprising some 22.7 % of South Africa, characterised by low rainfall and high temperatures. Grazing is the most common form of land use in the country, and particularly in this biome. Aridity and grazing are two factors that make large areas of South Africa (including the Nama-karoo) susceptible to land degradation. The Nama-Karoo biome has been neglected in past research and is an understudied part of South Africa's vegetation. This study is a component of a bigger umbrella project entitled "Restoration of degraded Nama-karoo rangelands: the role of conservation islands". The aim of this project was to assess the role played by isolated hills and mountains (mesas, inselbergs, etc.) in conserving the remnant biological diversity of the Nama-karoo rangelands of Namibia and South Africa. Vegetation composition, seed banks, seedling recruitment, pollination and grazing patterns and intensity on and off isolated mesas in the eastern parts of the Nama-karoo were investigated in the South African component of this project. I report on patterns of 1) plant species diversity and 2) plant communities across the Middelburg District, Eastern Cape, South Africa. Three isolated mesas (Tafelberg, Folminkskop and Buffelskop) and their surrounding plains were selected for this study. Permanent plots were established in broad transects extending from the plains to southeastern slopes, plateaux, north-western slopes and north-western plains of each mesa. Plots were 25 m2 in size, and ten 1 m2 blocks within the 25 m2 plot were randomly selected and sampled. Species composition was recorded and cover values determined for all plants in every subplot. In the eastern Nama-karoo mesas were not found to be higher in Alpha Diversity than their surrounding plains. Plains habitats were mostly equally as diverse as mesa habitats, with some plains habitats being higher in diversity than mesa habitats. Greater differences between mesas and their surroundings were found in a parallel study in the northern parts of the Nama-karoo in Namibia, indicating that mesas are more distinct islands of diversity with an increase in aridity. In the Middelburg District, a greater presence of woody phanerophytes occurred on mesa habitats in comparison with plains habitats, a possible result of the more moist microhabitats on mesas. Cooler, moister conditions on the south-eastern slopes of the mesas led to higher Alpha Diversity there in comparison with the warmer and drier north-western slopes, a phenomenon commonly found in arid areas. In terms of Alpha Diversity, the largest mesa sampled, Tafelberg (450 m above surroundings), was not significantly more diverse than the two smaller mesas (Folminkskop and Buffelskop both being 200 m above surroundings). Tafelberg and Buffelskop were equally high in Beta Diversity, with Folminkskop being much less diverse. Increased Beta Diversity for Tafelberg was explained by the increased size of the mesa (providing a more diverse microhabitat) relative to Folminkskop, which has the same shape and geology but is much smaller in size. High Beta Diversity values for Buffelskop were explained by the presence of degraded communities on the northwestern slopes and plains, while the south-eastern slopes were some of the most diverse habitats sampled in the landscape. Mesas and plains shared few species. Proportionally fewer species were shared between the plateau and slopes of the larger Tafelberg in comparison with the smaller two mesas. Isolation of the plateau could possibly become more distinct with an increase in size of the mesa. However, Tafelberg had more species in common with the surrounding plains than did the smaller mesas. It is suggested that bigger mesas support a more diverse array of microhabitats enabling plains species to occupy selective sites on the slopes of the mesa. Average Shannon-Wiener Alpha Diversity Indices for Middelburg were higher than that of the Nama-karoo in general. This could be explained by the higher precipitation in the eastern parts of the Nama-karoo relative to the rest. Diversity in Middelburg compared favourably with that in other arid lands of North America and Australia and was found to be similar in patterns and determinants of diversity compared to arid lands such as those of the New World. Regarding plant community composition, mesas were found to be distinctly different from their surrounding plains, with no shared communities between mesas and plains. The distribution of communities across the landscape was mainly attributed to a soilmoisture gradient. Mesas, compared to plains, have shallower, rockier soils coupled with very little run-off and naturally higher precipitation due to elevation. This results in a more mesic habitat. Plains, compared with mesas, have little rock cover, high percentages of bare soil and higher run-off rates than mesas, resulting in a more xeric habitat. Habitat differences such as these probably existed before the impact of domestic livestock but overgrazing has probably exacerbated the differences. The dominance structure of plant communities on the plains has probably changed in favour of toxic, spinescent or unpalatable plant species due to selective grazing by livestock. For the two dolerite-capped mesas (Tafelberg and Folminkskop) aspect and the expected cooler, moister conditions on south-eastern slopes as factors determining community composition were overridden by soil type and associated nutrient status. On Buffelskop (sandstone mesa), aspect and slope overrode soil type and associated nutrient status as determinants of community compositions. The potential to use mesas as a source of seeds and propagules to restore degraded plains habitats is low. Approximately 28 % of species were shared between mesas and their surroundings, and not all of these species could be regarded as good colonisers. Generalist, palatable species occurring across the landscape, such as Eragrostis obtusa, Felicia muricata, F. ovata, Fingerhuthia africana, Jamesbrittenia tysonii, Limeum aethiopicum, Pentzia incana and Selago albida could have some potential for future restoration attempts. Mesas are distinct in composition and can be regarded as islands of one vegetation type (mesas) in a sea of a different vegetation type (plains). Mesa habitats are not generally used for grazing by livestock, due to their general inaccessibility, and are not threatened by development. Plant communities in the Middelburg area were very similar In composition to communities identified in other parts of the Nama-karoo. Results from this study suggest that the plains are at present more degraded than the mesa habitats, but it is impossible to conclude whether or not these rangelands have stabilised given current stocking rates and climatic conditions. The Braun-Blanquet classification system proved an effective method to describe plant communities in the semi-arid Nama-karoo. All the plots used during the duration of the study were marked by metal stakes and are therefore of a permanent nature, so that they can be resampled in future. This might possibly shed some light on questions related to resilience, stability and degradation of the karoo. Restoring rangeland in the future is an important option, however, few farmers would, in the short term, be able to afford costly restoration techniques. It is recommended that farmers inspect the condition of their veld on a regular basis, and adjust stocking rates accordingly. Veld should be rested on a regular basis, and assessment should precede the movement of stock to a camp. Most farmers plough denuded areas of veld to encourage restoration, and it is recommended that this practice be combined with reseeding, followed by rest during and shortly after germination of the seed. / AFRIKAANSE OPSOMMING: Ariede areas beslaan tans 47 % van die aarde se land oppervlak. Tussen 32 % en 35 % van hierdie areas is onderworpe aan een of ander vorm van land degredasie. Die Nama- Karoo bioom is 'n ekstensiewe semi-ariede area wat naastenby 47 % van Suid-Afrika beslaan. Die karoo word gekenmerk deur hoë temperature en lae reënval. Suid-Afrika is 'n ariede land en water is 'n beperkende faktor. Weiding is die algemeenste boerderypraktyk, en meer so in ariede areas soos bv. in die karoo. Die droë klimaat en hoë voorkoms van weiding as boerderypraktyk maak ekstensiewe areas (insluitende die karoo) van Suid-Afrika vatbaar vir land degredasie. Hierdie studie is deel van 'n breër projek genaamd: "Restorasie van gedegradeerde Nama-karoo veld: Die rol van bewaringseilande". Die doel van hierdie projek was om die rol te bepaal van ge-isoleerde koppies en berge (mesas, inselberge, ens.) in die bewaring van biologiese diversiteit in die Nama-karoo van Suid-Afrika en Namibië. Plantegroeisamestelling, saadbanke, saailing oorlewing, bestuiwing en weidingsintensiteitlpatrone op koppies en hulle omringende vlaktes in die oostelike dele van die Nama-karoo is ondersoek in die Suid-Afrikaanse komponent van die studie. Ek rapporteer oor patrone van 1) plant diversiteit en 2) plantegroeigemeenskappe in die Middelburg Distrik, Oos-Kaap, Suid-Afrika. Drie ge-isoleerde koppies (Tafelberg, Folminkskop en Buffelskop) en hulle omliggende vlaktes is geselekteeer vir die doel van die studie. Permanente persele is uitgelê in 'n breë lyn vanaf die suid-oostelike vlaktes na die suid-oostelike hang, oor die plato's, noord-westelike hange en noord-westelike vlaktes vir al drie koppies. Persele was 25 m2 in grootte, waarvan tien 1 m2 sub-persele geselekteer is binne die groter 25 m2 blok. Spesies samestelling en bedekkingswaardes is bepaal vir alle plante in al die sub-persele. Mesas was nie hoër in Alpha Diversiteit as hulle omringende vlaktes nie. Vlaktes was meestal net so hoog in diversiteit as die mesas, en somtyds selfs hoër. Groter verskille in diversiteit is gevind in 'n paralelle studie in die noordelike dele van die Nama-karoo in Namibië. Dit dui aan dat mesas moontlik meer definitiewe eilande van diversiteit is in droër gebiede. Meer houtagtige fanerofiete het voorgekom op die mesas in vergelyking met die omliggende vlaktes, 'n moontlike gevolg van 'n natter habitat op die mesas. Suid-oostelike hange was hoër in alfa-diversiteit in vergelyking met noord-westelike hange. Dit is toegeskryf aan die koeler, natter mikrohabitat van die suid-oostelike hange. Die groter mesa (Tafelberg - 450 m bo die vlakte), was nie meer divers as die twee kleiner mesas nie (Folminkskop en Buffelskop is sowat 200 m bo die vlakte). Tafelberg en Buffelskop was ewe hoog in Beta Diversiteit, terwyl Folminkskop heelwat laer was. Hoër Beta Diversiteit vir Tafelberg kan verklaar word deur die groter oppervlak en hoogte van die mesa (voorsien 'n meer diverse mikrohabitat) in vergelyking met Folminkskop. Folminkskop het dieselfde vorm en geologiese geskiedenis as Tafelberg, maar is veel kleiner. Buffelskop was hoog in Beta Diversiteit as gevolg van 'n gedegradeerde gemeenskap op die noord-westelike hang en vlaktes, terwyl die suidoostelike hang baie hoog was in diversiteit. Min spesies kom op beide mesas en vlaktes voor. Minder spesies is gedeel deur die plato en hange van Tafelberg in vergelyking met die plato's en hange van Folminkskop en Buffelskop. Isolasie van die plato kan moontlik hoër wees in groter mesas. Tafelberg (slegs mesa) het meer spesies in gemeen met sy omliggende vlakte as die ander twee mesas. Groter mesas soos Tafelberg kan moontlik 'n meer diverse spektrum van mikrohabitatte bevat, wat sekere vlakte spesies in staat stel om te oorleef op mesa hange. Gemiddelde alfa diversiteit vir die Middelburg Distrik was hoër as die van die res van die karoo. Dit kan verklaar word deur die hoër reënval in die oostelike dele van die karoo relatief tot die meer westelike dele. Diversiteit in Middelburg vergelyk goed met die van ander ariede lande in Noord-Amerika en Australie, terwyl patrone en faktore wat diversiteit bepaal soortgelyk was aan die van gemeenskappe in ariede lande van die Nuwe Wêreld. Mesas en vlaktes het drasties verskil in die samestelling van hulle onderskeie plantegroeigemeenskappe, alhoewel hulle soortgelyk was in diversiteit. Geen plantegroeigemeenskappe het voorgekom op beide mesas en vlaktes nie (gedeelde plantegroeigemeenskappe). Die verspreiding van plantegroeigemeenskappe oor die landskap was toegeskryf aan 'n grond-water gradient. Mesas het, oor die algemeen, vlakker, meer rotsagtige grond en 'n hoër reënval met minder afloop as vlaktes. Vlaktes het minder rotse, 'n hoër persentasies kaal grond en vinniger afloop tempo's na reën as mesas. Hierdie faktore het tot gevolg dat mesas 'n natter habitat verteenwoordig in vergelyking met die meer ariede vlaktes. Habitatsverskille soos dié het heel moontlik klaar bestaan voor die aankoms van vee, maar oorbeweiding het die verskille in habitat tussen mesas en vlaktes groter gemaak. Selektiewe beweiding deur vee het waarskynlik tot gevolg gehad dat plante wat giftig, doringagtig of onsmaaklik is, toegeneem het in plantegroeigemeenskappe, ten koste van meer smaaklike spesies. Grondtipe en grondsamestelling het aspek en verwagte koeler kondisies op die suidoostelike hange onderdruk as bepalende faktore vir plantegroeigemeenskap samestelling op Tafelberg en Folminkskop (beide bedek met 'n doleriet laag). Aspek en steilte van die hange het grondtipe en grondsamestelling onderdruk as bepalende faktore vir plantegroeigemeenskappe op Buffelskop. Die potensiaal om mesas te gebruik as bronne van saad vir die rehabilitasie van die vlaktes was laag. Mesas en hulle omliggende vlaktes het naastenby 28 % van spesies in gemeen gehad, maar nie al hierdie spesies was goeie koloniseerders nie. Sekere smaaklike plante wat op beide mesas en vlaktes voorgekom het, kan potensiaal hê vir toekomstige restorasie doeleindes, bv. Eragrostis obtusa, Felicia muricata, F. ovata, Fingerhuthia Africana, Jamesbrittenia tysonii, Limeum aethiopicum, Pentzia incana en Selago albida. Mesas is uniek in samestelling en kan beskou word as een plantegroeitipe in 'n see van 'n ander tipe (vlaktes). Mesas word nie oor die algemeen bewei nie (as gevolg van hulle steil hange en bergagtigheid) en word gevolglik nie bedreig deur huidige ontwikkeling nie. Plantegroeigemeenskappe in Middelburg is soortgelyk aan die van ander dele van die Nama-Karoo bioom. Resultate van die studie dui aan dat die vlaktes huidiglik meer gedegradeer is as mesas, maar dit is onbekend of die veld gestabiliseer het of steeds besig is om verder te degradeer in huidige weidings- en klimaatskondisies. Die Braun-Blanquet klassifikasiemetode IS effektief vir die beskrywing van plantegroeigemeenskappe in ariede areas. Al die persele in die studie area is gemerk met metaal paaltjies en is dus permanent. Dit maak dit moontlik om in die toekoms die persele weer te ondersoek. 'n Langtermyn datastel kan moontlik lig werp op die stabiliteit en degredasie van karoo veld. Restorasie van veld is 'n moontlikheid vir die toekoms, maar min boere kan duur restorasie metodes bekostig in die kort termyn. Boere moet hulle lande gereeld ondersoek en drakrag aanpas by die kondisie van die veld. Baie boere ploeg kaal kolle in die veld op 'n gereelde basis, en daar word voorgestel dat die praktyk gekombineer word met die saai van geskikte saad, gevolg deur 'n rusperiode tydens en na ontkieming daarvan.

Thesis (MSc)--University of Stellenbosch, 2003. / ENGLISH ABSTRACT: Arid and semi-arid rangelands of the Nama-Karoo Biome are believed to have changed considerably since the arrival of domestic livestock in the veld. Severe grazing pressure is considered to be one of the prime factors responsible for the perceived degradation of vegetation and concurrent soil degradation. To understand the process of degradation and to make further recommendations for future veld restoration, a large-scale project was undertaken in the Eastern Cape. This project focused on the role that landscape heterogeneity plays in providing refuges for plant species. The key question asked in this project was: What role do these hypothetically less grazed mesas play in the conservation of rangelands in the Nama-Karoo of the Eastern Cape? This study, a component of the larger project, investigated grazing and soil landscape gradients on- and off- the three mesas (Tafelberg, Folminkskop and Buffelskop) in the Middelburg District of the Eastern Cape, South Africa and the possible influences that they might have on the veld. The flats surrounding the mesas were mostly used as grazing camps in contrast to the plateaux of the mesas, which had varied levels of accessibility. In the absence of direct observations, the primary objective of the study was to test the differential dung pellet abundance and impact of animals on different parts of the landscape. As such, dung pellet counts in this study were correlated with surrogates of soil physical properties including bare ground, trampling and litter cover. Variation in dung pellet density was found at the different habitats (flats, slopes and plateaux) of all mesas. The flats to the north-eastern and south-western of Tafelberg mesa were found to be more heavily utilized by livestock and herbivores, while the plateaux and southeastern slopes of Folminkskop and Buffelskop were also utilized by grazers. The Tafelberg mesa was the only study site that was consistent with the hypothesis which stated that grazers would be less concentrated on the plateau compared to the surrounding flats due to its inaccessibility, whilst the high mean dung pellet density on the plateaux of the smaller Folminkskop and Buffelskop mesas due to easier access contradicted the original hypothesis. It appeared that dung pellet density did not clearly turn out to be an indicator of habitat use in this study, but showed where slopes and plateaux were accessible to herbivores, as in the case for the Buffelskop mesa, a higher abundance of dung pellets were found suggesting that higher intensities of habitat use took place. During this study a strong pattern was observed of bare soil patches on the flats surrounding the mesas. There was a decrease in percentage of bare soil along the gradient of the three mesas with a high percentage of bare soil on the plateaux of the mesas. A significant correlation was found between bare soil and dung pellet density. However, the plateaux of Folminkskop and Buffelskop had a high percentage of bare soil compared to the plateau of Tafelberg mesa. Farmers mainly used these smaller mesas as grazing camps for their livestock and herbivores. A positive correlation between bare soil and litter cover of the different habitats was evident in this study. A lower percentage of litter cover at these sites was associated with a high percentage of bare soil. Litter is very important in a healthy vegetation community in terms of nutrient cycling and fertile patches. A detailed assessment of soil chemical and physical properties would reveal, firstly, if vegetation change is better explained by soil or grazing effects and, secondly, if changes in soil have resulted from land use. Differences in macro- and micro-site variations between open-canopy (between shrubs) and closed-canopy (under shrubs) sites for each habitat were determined to differentiate between local scales due to land use and landscape scales due to geomorphology. The results suggested that carbon, phosphorus, calcium, magnesium, copper and manganese levels in soils at landscape scale better explain vegetation changes between habitats. At a local scale (open- and closed-canopy sites) land use was responsible for little changes in soils. Changes in only soil potassium, zinc and boron elements were actually a consequence of local scales due to land use. The soil nutrient content on the slopes appeared to be intermediate between the flats and plateaux of all three mesas. It appeared that dolerite capped Tafelberg and Folminkskop mesas had high silt and clay content, while Buffelskop (sandstone) mesa had a lower silt and clay content. Consequently, the texture and parent material of the soils contributed to the variations in soil nutrient composition between these mesas. High infiltration rate together with low nutrient content on the flats clearly showed that these flats, surrounding the mesas were degraded. The high infiltration rates were caused by high activity by livestock and other indigenous animals on the flats which breaks the surface crusting of bare soil and improve infiltration. It was concluded that high levels of grazing at these sites have also altered the textural and soil properties. Endozoochory dispersal and the deposition of dung pellets in areas of small patch disturbances play an important role in veld regeneration in degraded areas. Dung pellets collected from permanent study sites on the southeastern and northwestern flats and slopes, and all the study sites on the plateau of Tafelberg mesa, was sown in seedling trays, watered and monitored for seedling germination. Species list were then compared to below-ground soil seed bank data and above-ground vegetation data collected by other researchers at the same permanent study sites. Higher seedling percentages were recorded from dung pellets collected on the flats than on the plateau. A total of sixteen species were found to germinate in dung pellets collected on the flats compared to ten species germinating in dung pellets collected on the slopes and two species on the plateau of Tafelberg mesa. The seeds that germinated represent a variety of palatable grasses and shrub species. Aristida sp., Eragrostis bicolor, Eragrostis ch/orome/as and Eragrostis obtusa were palatable grasses recorded for dung pellets collected on the flats. Of the species recorded, Aristida sp., Chenopodium sp. and Pentzia sp. were found in dung pellets but were not recorded in parallel soil seedbank and vegetation studies. Successful restoration of veld conditions requires strict grazing management practices. Germination of seed in dung pellets might be considered to be a valuable means of indicating restoration potential and rangeland conditions for the identification of both degraded and conservation worthy areas. With appropriate land management skills and restoration measures, these challenges can be constructively and creatively faced. / AFRIKAANSE OPSOMMING: Daar word geglo dat die ariede en sernl-arlede weivelde van die Nama-Karoo bioom aansienlik verander het sedert die aankoms van mak lewende hawe in die veld. Swaar weidingsdruk word beskou as een van die vernaamste faktore verantwoordelik vir die waarneembare oorbeweiding van plantegroei en die gelyktydige grond agteruitgang (degradasie). Ten einde die proses van degradasie beter te verstaan, en om verdere aanbevelings te kan maak vir toekomstige veld herstel, is huidiglik 'n grootskaalse projek in die Oos-Kaap aan die gang oor die rol wat landskap ongelyksoortigheid speel in die voorsiening van skuiling aan plant spesies. Hierdie studie ondersoek weiding en grondlandskap gradiënte (op en af) van die drie mesas (Tafelberg, Folminkskop en Buffelskop) in die Middelburg streek van die Oos- Kaap, Suid-Afrika en die moontlike invloed wat hierdie gradiënte op die veld het. Die vlaktes was meestal gebruik vir weidingskampe in teenstelling met die kruin van die mesas. Hierdie studie het duidelike patrone vir habitat-gebruik op die vlaktes, hange en kruine van al die mesas aangetoon. Die vraag wat gestel was, was watter rol speel hierdie hipoteties minder beweide mesas in die bewaring van weivelde in die Nama- Karoo van die Oos-Kaap? Hierdie studie toets nie direk vir mis verspreiding as 'n plaasvervanger vir dier verspreiding in die landskap sedert werklike dier getalle nie oorweeg was nie. In die awesigheid van direkte waarneming, die primêre doelwit was om te toets die differensiaal misdigtheid en die impak van diere op verskillende dele van die landskap. Mis getalle in hierdie studie was gekorreleer met plaasvervangers van fisiese eienskappe insluitent onbedekte grond, vertrapping en droë plant material decking. Variasie in misdigtheid was gevind by die verskillende habitate (vlaktes, hange en kruine) van all die mesas. Die vlaktes aan die noord-oostelike en suid-westelike kante van die Tafelberg mesa was meer hewig benut deur lewende hawe en hêrbivore, terwyl die kruine en suid-oostelike hange van Folminkskop en Buffelskop ook gebruik was deur weidende diere. Die Tafelberg mesa was die enigste studie area wat konsekwent was met die hipotese, terwyl die hoë gemiddeld misdigtheid op die kruine van die kleiner Folminkskop en Buffelskop mesas, as gevolg van makliker toegang weerspreek die oorspronklike hipotese. Dit blyk dat misdigtheid duidelik opkom as 'n nie aanwyser habitat-gebruik in hierdie studie, maar wys well waar hange en kruine toeganklik was vir herbivore, soos in die geval van Buffelskop mesa, oorvloed van mis was gevind wat voorstel dat hoer intensiteit van habitat-gebruik voorgekom het. 'n Duidelike patroon is waargeneem tydens hierdie studie van onbedekte grond areas op die vlaktes rondom mesas. Daar was 'n afname in persentasie onbedekte grond langs die gradiënt van die drie mesas. Die kruine van Folminkskop en Buffelskop toon hoër persentasies onbedekte grond in vergelyking met die kruin van die Tafelberg mesa. Daar is 'n betekenisvolle korrelasie tussen onbedekte grond en misdigtheid gevind. Boere gebruik hierdie kleiner mesas hoofsaaklik as weidingskampe vir hulle lewende hawe en hêrbivore. In hierdie studie is daar 'n positiewe korrelasie gevind tussen onbedekte grond en droë plant material decking van die verskillende habitatte. Droë plantmateriaal bedekking speel 'n baie belangrik rol in 'n gesonde plant gemeenskap in terme van voedingsirkulering en vrugbare kolle. 'n Breedvoerige skatting van grond chemise en fisiese eienskappe wou openbaar eerstens of plantegroei verandering beter verklaar word deur grond of weiding effekte en tweedens, of verandering in grond veroorsaak was deur land-gebruik. Verskille in makro- en mikroterrein-wisseling tussen oop-blaredak (tussenplantruimtes) en toeblaredak (onderplantdekking) terreine vir elke habitat was bepaal om te onderskei tussen lokaal effek as gevolg van land-gebruik en landskap effek as gevolg van geomorfologie. Die resultate stel voor dat koolstof, fosfor, kalsium, magnesium, koper en mangaan vlakke in die grond by landskap effek beter plantegroei verandering tussen habitate verduidelik. By 'n lokaal effek (oop-en toe-blaredak terreine) land-gebruik was verantwoordelik vir min veranderinge in grond. Verandering in grond kalium, sink en baron elemente was werklik 'n gevolg van lokaal effek as gevolg van land-gebruik. Die nutrient inhoud van grond op die hange blyk intermediêr te wees tussen die van die vlaktes en kruine van al drie mesas. Hierdie studie toon ook dat die geologie en topografie van hierdie mesas die grond tekstuur en nutrient samestelling op en af van die mesas beinvloed. Dit wil voorkom asof die dolerite bedekte Tafelberg en Folminkskop mesas, hoë slik en klei materiaal bevat, terwyl Buffelskop (sandsteen) mesa 'n laer slik en klei inhoud het. Gevolglik, tekstueel en ouer materiaal grond dra by tot die variasie in grond nutrient samestelling tussen hierdie mesas. Hoë infiltrasie tempo tesame met lae voedingstof van die vlaktes dui daarop dat die vlaktes rondom die mesas oorbewei was. Hierdie hoë infiltrasie tempo word veroorsaak deur hoë dierlike aktiwieteite op die vlaktes deur lewende hawe en ander inheemse diere wat die oppervlakke van onbedekte grond versteur en infiltrasie verbeter. Die gevolgtrekking word gemaak dat die hoë weidingsvlakke van hierdie terreine ook die tekstuur en ander grond eienskappe verander het. Endozoochoreeuse verspreiding en die mis-afsetting in gebiede van klein areaversteurings, speel 'n belangrike rol in veld herlewing in oorbeweide gebiede. Mis versamel by permanente studie terreine op die SO en NW vlaktes en hange, as ook al die studie terreine op die kruin van die Tafelberg mesa, is gesaai in saailing bakke, besproei en gekontroleer vir saailing ontkieming. 'n Lys van spesies is vergelyk met onder-grondse saadbank gegewens en bo-grondse plantegroei gegewens wat deur ander navorsers by dieselfde studie terreine versamel was. Hoër saailing persentasies is gevind vir mis wat op die vlaktes versamel is as op die kruine. 'n Totaal van sestien geidentifiseerde spesies het uit die mis op die vlaktes ontkiem in vergelyking met die tien spesies wat ontkiem het uit die mis van die hange en twee spesies iut die mis van die kruin van Tafelberg mesa. Die ontkiemde saad verteenwoordig 'n verskeidenheid van vreetbare gras en struik spesies. Aristide sp., Eragrostis bicolor, Eragrostis chloromelas en Eragrostis obtusa is vreetbare grasse wat gevind was in die mis versamelop die vlaktes. Die spesies opname wys dat Aristida sp., Chenopodium sp. en Pentzia sp. Gevind was in die mis, maar nie in die parallelle grond saadbank en plantegroei studies nie. Die suksesvolle herstel van veldtoestande vereis "n streng weidingsbestuur toepassing. Die ontkieming van saad in mis kan beskou word as "n bekostigbare manier om veld herstel potentiaal te bepaal, asook weiveld toestande vir die indentifiseering van beide degradasie en bewaring van waardevolle areas. Met gepaste veldbestuur vaardighede en herstel maatreels, kan hierdie uitdagings konstruktief en kreatief aangespreek word.

Master of Science / Department of Agronomy / Walt Fick / Prescribed burning has been a common conservation practice on native prairie dating back to the days of pioneer settlement. Advantages include increased forage quality, reduction of undesirable plants, improved wildlife habitat, removal of accumulated dead plant litter and relatively low costs. While spring is the commonly accepted time to burn, little research has been conducted on late-summer and fall burning for specific objectives that include targeting undesirable plant species and measuring potential effects on soil physical properties. The first part of this study was to evaluate the effect that prescribed burning has on population dynamics of sericea lespedeza (Lespedeza cuneata [Dumont] G. Don), rough-leaf dogwood (Cornus drummondii Meyer), and additional woody species. Stem counts and cover estimates were taken from 20, 0.25-m[superscript]2 frames prior to and post-burn. Change in botanical composition, plant density, frequency, and Daubenmire canopy cover estimates were calculated. Sericea lespedeza plant frequency across all clay upland burns decreased 2.27% and increased 4.76% across all loamy/limy upland burns the first growing season post-burn. Dogwood densities increased 3.12 stems m[superscript]-[superscript]2 on spring burns compared to a decrease of 0.30 stems m[superscript]-[superscript]2 on unburned plots the first growing season post-burn. Changes in frequency of other woody species the first growing season post-burn showed significant interactions between burn treatment and ecological site, and between ecological site and year. A significant interaction between burn treatment and ecological site was found on total woody species plant composition changes two growing seasons post-burn for the first year of burn treatments. The secondary part of this study was to evaluate the effect of prescribed burning on soil bulk density and wet-aggregate stability. Soil samples were collected along the same line-transects used for vegetation sampling. Significant differences among mean weight diameters (MWD), percent water-stable aggregates (WSA), and WSA size fractions occurred between burned and unburned soils following burning in the fall of 2011. Monitoring plant and soil response to prescribed burning in different seasons may lead to adjustments being made in management of rangelands where sericea lespedeza, dogwood, and additional woody species occur.