The movement and distribution of elephants can be influenced by environmental factors over time (Foley, 2002). Examining how features in the landscape such as vegetation productivity, water sources and anthropogenic activities drive the movement of elephants can help in understanding patterns of movement. It can also help to inform the establishment and alignment of protected areas, wildlife corridors and identification of tourism hotspots as well as policy interventions to manage Human-Elephant Conflict (HEC). The Okavango Panhandle in Botswana is a HEC hotspot and the focus of My study. A number of strategies to address HEC are underway in the area, however one longer term strategy that has been proposed in this area involves provision of artificial water sources to influence elephant movements and keep animals away from fields during the cropping season. However, an improved understanding of how elephants utilize their habitats in relation to natural ephemeral surface water and other factors that influence their movements from dryland habitats to the Okavango Delta resources is needed to inform such management decisions. My study seeks to establish the role of ephemeral surface water on elephant distribution in the eastern Okavango Panhandle, Botswana as well as assess the movement distribution of elephants in relation to the seasonality, proximity and spatial extent of water presence represented by ephemeral surface water. Time series analysis of water extent on ephemeral surface water of the eastern Okavango panhandle will be developed and overlaid with elephant movement datasets. Elephant collar data from 15 elephants (5 males and 10 females) in the eastern Okavango Panhandle, Botswana have been analysed and Home Range (HR) sizes estimated using Kernel Density Estimation (KDE). The relative importance/probability of environmental variables in determining elephants' movement based on the Utilization Distribution (UD) were computed using Generalized Linear Mixed Models (GLMMs). I utilized a remote sensing spectral index, namely the Automated Water Extraction Index (AWEI) to delineate ephemeral surface water in dryland (excluding permanent waters) of the study area. The results reveal that during the wet season, elephants were evenly spread out all over the study area until the early dry season (April-June) when the ephemeral waterholes dried up. Elephants moved southwards towards the permanent waters of the Okavango River, where there are many human settlements and farms. Male HR sizes were found to be bigger than those of female elephants. Wet season (early and late) home range sizes were also bigger when compared to dry season (early and late) HR size. Mean daily distances were computed to investigate the effect of season on elephant daily distances and the distances ranged between 5km and 6.8km in the late wet and in the early wet and late dry season respectively. The Resource Selection Function (RSF) analysis shows that water adjacent sites are preferred over distant ones and both sexes prefer areas with high NDVI, with this preference being more pronounced in males. The seasonal variation of water use is notable in that it affirms the importance of proximity to water for elephants and has implications for their management and HEC. For example, I found that ephemeral surface water has a significant role in influencing elephant spatial use in the area, particularly during the early and late wet season. As ephemeral pans dried and NDVI (vegetation greenness) decreased, elephants started to move closer to the Okavango Delta and consequently human settlements and fields. However, further investigations into the timing of movements away from ephemeral waterholes and the influence of other environmental factors on elephant movements in the area would be needed before any recommendations can be made regarding artificial water provision in this area.

Plant-pollinator interaction assembly can be represented as so called pollination network. Using these networks, assembly of plant-pollinator communities and geographic patterns in pollination networks can be studied. During last years, a great progress in describing patterns in structure of plant-pollinator interaction webs and in describing their spatial variability was made. However, these results are mostly based on data from previous studies, which are not seamlessly comparable due to their nonuniformity. My thesis aims at the methodological concepts of study on pollination networks. Based on meta- analysis of previously published data and on data from four years detailed study of plant-pollination system from one locality, I show several methodological problems caused by dissimilarities in sam- pling method of current pollination-network data. These dissimilarities could be responsible for some observed patterns in the assembly of pollination networks. Thus, I propose a uniform method for observing pollination networks suitable for macroecolo- gical meta-analyses (pollination network samples). I tested this method in the field and I described 49 pollination networks from the central and eastern Europe by using such method. In these ne- tworks, no nonrandom structure character such as...

Knowledge about a species’ reproductive parameters such as breeding behaviours is a vital building block for essential conservation actions, especially for endangered species. Despite this, there is a considerable knowledge gap about the snow leopard (Panthera uncia) maternal behaviours, as well as the timing of den independence for the cubs. It has been assumed that female snow leopards change their behaviours post-denning and that the cubs leave their den together with their mothers around two to three months of age. However, until this day no quantitative data has been used to analyse female behaviours post-denning and when the cubs leave their den. I analysed pre- and post-denning activity for seven GPS-collared snow leopard females in Tost Mountains of southern Mongolia during the years 2010 to 2019. With linear mixed models and generalized linear mixed models, I found that female snow leopards with small cubs changed their behavioural patterns and space use by decreasing their monthly home range size, compared to females with older or no cubs. When the cubs became six months old, there were no detectable differences in these behaviours which suggests that the cubs started to travel continuously with their mother at the age of 5-6 months. The rate at which the collars successfully acquired positions decreased considerably during the early phase of denning when the female spent considerable time at the den sites where the collars could not communicate with the satellites. The age of the cubs when the female left the den ranged from 21-61 days (mean =44 days), suggesting that snow leopard females use their dens for 1.5 ± 0.5 months. This study provides the first estimate of the extent of den use by snow leopards, as well as the first estimates of post-denning behavioural patterns for snow leopard females and their cubs.

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Harmonia axyridis is a ladybird native to Asia, but introduced widely as a biocontrol agent. It is invasive and detrimental to native species in North America, which meant its arrival in Britain was met with concern. Establishment was seen as an opportunity to track the spread of an invasive alien species (IAS) whilst also monitoring impacts on native species. The aims of this thesis were to examine the responses of native British ladybirds to the arrival of H. axyridis, to establish the effect of the IAS on native ladybirds when compared to other drivers, and to investigate the possible facilitation of the H. axyridis invasion by natural enemy release. Modelling ladybird distributions with life-history and resource-use traits found that species predatory on a wide range of prey families had larger range sizes than those which ate fewer prey types. This suggests that the wide diet breadth of the IAS is likely to have played a critical role in the species’ rapid spread. Dietary niche overlap between H. axyridis and native ladybirds showed positive correlation with declines of native ladybirds. This indicates that the IAS is playing an important role, but the significance of urbanisation suggests habitat destruction is also significant. Abundance of H. axyridis was influenced by habitat type and aphid abundance, but not by the native ladybird community, suggesting the spread of the IAS will not be slowed by biotic resistance. Harmonia axyridis is attacked by native parasitoids, but at a much lower rate than is the native Coccinella septempunctata, in line with natural-enemy release theory. There was no evidence of attack rate increasing with time since arrival in an area. Overall, H. axyridis is an extremely successful IAS, with detrimental effects on native ladybirds which are likely to continue.

It has been more than two centuries since the latitudinal pattern of increase in taxonomic richness from poles to equator was first documented. After two centuries of research, and with more than two dozen hypotheses proposed, an understanding of the mechanisms underlying this pattern and their relative importance remains incomplete. Factors such as evolutionary history, area and latitude associated variables such as temperature, solar energy, climatic stability and seasonality are known to influence species richness by affecting geographic range size and location over ecological and evolutionary time. Understanding the forces that affect geographic range size is, therefore, integral to our understanding of latitudinal patterns in species richness. Using woody plants as a study system, my dissertation deciphers the latitudinal pattern, if any, in species richness within the evergreen forests of the Western Ghats. These wet evergreen forests form an evolutionarily distinct biogeographic zone, which has remained isolated from its counterparts. This has resulted in a high percentage of endemism among the evergreen woody plants and, therefore, the global geographic ranges of most of these plants are restricted within the boundaries of the Western Ghats. The first main objective of this dissertation is to understand the determinants of geographic range size in the evergreen woody plants of the Western Ghats. Further, the Western Ghats are characterized by a sharp climatic gradient in temperature and rainfall seasonality that is not correlated with mean annual temperature or annual rainfall. This allows a direct test of the hypotheses and predictions that are based on climatic seasonality, without the confounding effect of other climatic correlates of latitude. Therefore, the second main objective of this dissertation is to understand the mechanisms underlying latitudinal patterns in species richness of evergreen woody plants in the Western Ghats. Regional species richness is an outcome of two factors- local species richness of each location within the region and turnover in species composition among the locations, which in turn are a result of patterns in range size, range location and range overlap. To address these two objectives, I first test the effect of climatic niche of a species in determining geographic range size and then examine the effect of latitude associated climatic seasonality on range location and range overlap. Next, I link the observed pattern in range geometry to latitudinal patterns in species turnover and finally to latitudinal patterns in species richness. While the first part of my dissertation study deals with factors that generate spatial variation in species richness, the second part deals with the factors underlying spatial variation in species composition. Environmental heterogeneity and dispersal are considered the most important determinants of species turnover i.e. change in species composition. However, their relative importance in structuring in diverse plant communities within tropical regions across different scales is poorly understood. Hence, the third objective of this dissertation is to understand the processes that influence change in species composition of woody plants within the Western Ghats. Geographic range size and population size are important attributes of species rarity, which are directly linked to their extinction risk. Hence, data on distribution and population status of species can help us focus our efforts on those species that require conservation attention. This is achieved through carrying out species threat assessments based on attributes such as range and population size and then assigning then to a threat category. A majority of species endemic to the Western Ghats have not yet been assessed, largely due to lack of data on their population and distribution status. Therefore, the fourth and the final part of my dissertation explores the application of information on species range size and abundance in prioritizing species for conservation. To address these objectives, I sampled the wet evergreen forests of the Western Ghats along a series of locations distributed across its entire latitudinal gradient. Based on 156 plots, covering a latitudinal gradient of more than 1200 km and comprising of more than 20,000 occurrence locations belonging to more than 450 species of woody plants, I derived quantitative estimates of latitudinal gradients in range size, local and regional richness as well as species turnover. I used a combination of statistical and simulation approaches to discern the mechanisms underlying large-scale pattern in species ranges, richness and turnover. My dissertation is structured as follows.

Orientador: Clarisse Palma da Silva / Resumo: Áreas úmidas são ambientes na interface terrestre e aquática, onde sazonalmente a disponibilidade de água pode estar em excesso ou em escassez. A história geológica da bacia amazônica está intimamente relacionada com a presença de áreas úmidas em grandes extensões espaciais e temporais e em variadas tipologias. Dentre as tipologias de áreas úmidas presentes na Amazônia as áreas alagáveis ao longo das planícies de inundação dos grandes rios são possivelmente as que possuem maior extensão territorial. Esta tese aborda o papel das áreas úmidas para a diversidade de árvores na Amazônia. As florestas que crescem em áreas úmidas possuem menor diversidade de espécies arbóreas em relação às florestas em ambientes terrestres (i.e., florestas de terra-firme); possivelmente devido às limitações ecológicas e fisiológicas relacionadas a saturação hídrica do solo e as inundações periódicas. Entretanto, nas áreas úmidas da Amazônia já foram registradas 3,515 espécies de árvores (Capítulo 2), uma quantidade comparável à da diversidade na Floresta Atlântica. Em relação às florestas de terra-firme da Amazônia, as espécies de árvores que ocorrem em áreas úmidas tendem a apresentar maiores áreas de distribuição e amplitudes de tolerâncias de nicho ao longo da região Neotropical (Capítulo 3). A composição florística e a distância filogenética entre espécies arbóreas nas florestas de várzea da Amazônia central mudam amplamente entre localidades (Capítulo 4). O gradiente ambiental contido entre as ... (Resumo completo, clicar acesso eletrônico abaixo) / Abstract: Wetlands are in the interface of terrestrial and aquatic environments, where seasonally water availability may be in excess or scarcity. Geological history of Amazon basin is closely linked with a huge temporal and spatial extents of wetlands. Nowadays, floodplains (i.e., Vázea and Igapó) are the wetlands with greatest coverage in Amazon. The present thesis is focused on the role of wetlands to tree species diversity in Amazon. Wetland forests have lower tree species diversity than upland forests (i.e., Terra-Firme); most likely due to ecological and physiological limitations. Notwithstanding, in Amazonian wetland forests 3,515 tree species already were recorded, (Chapter 2), which is comparable to tree species diversity in the Atlantic Forest. Wetland tree species show greater ranges sizes and niche breadth compared to tree species do not occur in wetlands (Chapter 3). Floristic compositional turnover and phylogenetic distances between floodplain forests in Central Amazon is high (Chapter 4). The most influential driver of floristic compositional turnover was the geographic distances between localities, whereas phylogenetic distances is driven mainly by the environmental gradients between forests. Furthermore, in general, the most abundant species are those that shows greater co-occurrence associations (Chapter 5). Co-occurrence structure is influenced by biotic interactions like facilitation and competition among species, but also by niche similarities indicated in the evol... (Complete abstract click electronic access below) / Doutor