We study the Sunyaev-Zel'dovich effect (SZE) signature in South Pole Telescope (SPT) data for an ensemble of 719 optically identified galaxy clusters selected from 124.6 deg(2) of the Dark Energy Survey (DES) science verification data, detecting a clear stacked SZE signal down to richness lambda similar to 20. The SZE signature is measured using matched-filtered maps of the 2500 deg(2) SPT-SZ survey at the positions of the DES clusters, and the degeneracy between SZE observable and matched-filter size is broken by adopting as priors SZE and optical mass-observable relations that are either calibrated using SPT-selected clusters or through the Arnaud et al. (A10) X-ray analysis. We measure the SPT signal-to-noise zeta - lambda relation and two integrated Compton-y Y500-lambda relations for the DES-selected clusters and compare these to model expectations that account for the SZE-optical centre offset distribution. For clusters with lambda > 80, the two SPT-calibrated scaling relations are consistent with the measurements, while for the A10-calibrated relation the measured SZE signal is smaller by a factor of 0.61 +/- 0.12 compared to the prediction. For clusters at 20 < lambda < 80, the measured SZE signal is smaller by a factor of similar to 0.20-0.80 (between 2.3 sigma and 10 sigma significance) compared to the prediction, with the SPT-calibrated scaling relations and larger lambda clusters showing generally better agreement. We quantify the required corrections to achieve consistency, showing that there is a richness-dependent bias that can be explained by some combination of (1) contamination of the observables and (2) biases in the estimated halo masses. We also discuss particular physical effects associated with these biases, such as contamination of. from line-of-sight projections or of the SZE observables from point sources, larger offsets in the SZE-optical centring or larger intrinsic scatter in the lambda-mass relation at lower richnesses.

Understanding the formation and evolution of galaxies is a crucially important task in modern astronomy. It is well known that galaxy formation is strongly affected by the environments they reside in. Galaxy clusters, as the densest large-scale structures in the Universe, thus serve as ideal laboratories to study how galaxy formation proceeds in dense environments. Clusters already began to form at $z>2$, therefore to directly witness the early stage of galaxy formation in dense environments, it is necessary to identify progenitors of clusters (`protoclusters') and study their galaxy constituents within. In this thesis, I present two observational studies on high-redshift protoclusters at $z>3$. Utilizing multiwavelength data and different galaxy selection techniques, significant galaxy overdensities are found in the two protoclusters, which are predicted to evolve into Coma-like clusters by present day. Various types of galaxies are identified in the protocluster, such as normal star-forming galaxies, massive quiescent galaxies and post-starburst galaxies. Together with extreme and rare sources such as giant Lyman-alpha nebulae and brighest cluster galaxy, they paint a picture of how different galaxy populations trace the underlying dark matter halos. Finally, the environmental impact on galaxy properties appears to be a subtle one for these protoclusters, which might depend on the galaxy population one chooses to study.

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This thesis addresses the challenge of understanding the typical star formation histories of red sequence galaxies, using linestrength indices, mass-to-light ratios and redshift evolution as complementary constraints on their stellar age distribution. We first construct simple parametric models of the star formation history that bracket a range of scenarios, and fit these models to the linestrength indices of low-redshift cluster red-sequence galaxies. For giant galaxies, we con firm the downsizing trend, i.e. the stellar populations are younger, on average, for lower σ galaxies. We find, however, that this trend flattens or reverses at σ < 70km/s. We then compare predicted stellar mass-to-light ratios with dynamical mass-to-light ratios derived from the Fundamental Plane, or by the SAURON group. For galaxies with σ ~ 70 km/s, models with a late 'frosting' of young stars and models with exponential star formation histories have stellar mass-to-light ratios that are larger than observed dynamical mass-to-light ratios by factors of 1.7 and 1.4, respectively, and so are rejected. The single stellar population (SSP) model is consistent with the Fundamental Plane, and requires a modest amount of dark matter (between 20% to 30%) to account for the difference between stellar and dynamical mass-to-light ratios. A model in which star formation was 'quenched' at intermediate ages is also consistent with the observations, although in this case less dark matter is required for low mass galaxies. We also find that the contribution of stellar populations to the 'tilt' of the Fundamental Plane is highly dependent on the assumed star-formation history: for the SSP model, the tilt of the FP is driven primarily by stellar-population effects. For a quenched model, two-thirds of the tilt is due to stellar populations and only one third is due to dark matter or non-homology. Comparing to high redshift cluster data, we find again the SSP and quenched models, as well as a model where strangulation begins at intermediate ages after a period of constant star formation, are preferred. They predict the recent faint-end build up of the red sequence, along with observed dwarf-to-giant ratios. Only the SSP model appears to predict the observed M/L evolution, but only if selection effects are carefully modeled.

This thesis addresses the challenge of understanding the typical star formation histories of red sequence galaxies, using linestrength indices, mass-to-light ratios and redshift evolution as complementary constraints on their stellar age distribution. We first construct simple parametric models of the star formation history that bracket a range of scenarios, and fit these models to the linestrength indices of low-redshift cluster red-sequence galaxies. For giant galaxies, we con firm the downsizing trend, i.e. the stellar populations are younger, on average, for lower σ galaxies. We find, however, that this trend flattens or reverses at σ < 70km/s. We then compare predicted stellar mass-to-light ratios with dynamical mass-to-light ratios derived from the Fundamental Plane, or by the SAURON group. For galaxies with σ ~ 70 km/s, models with a late 'frosting' of young stars and models with exponential star formation histories have stellar mass-to-light ratios that are larger than observed dynamical mass-to-light ratios by factors of 1.7 and 1.4, respectively, and so are rejected. The single stellar population (SSP) model is consistent with the Fundamental Plane, and requires a modest amount of dark matter (between 20% to 30%) to account for the difference between stellar and dynamical mass-to-light ratios. A model in which star formation was 'quenched' at intermediate ages is also consistent with the observations, although in this case less dark matter is required for low mass galaxies. We also find that the contribution of stellar populations to the 'tilt' of the Fundamental Plane is highly dependent on the assumed star-formation history: for the SSP model, the tilt of the FP is driven primarily by stellar-population effects. For a quenched model, two-thirds of the tilt is due to stellar populations and only one third is due to dark matter or non-homology. Comparing to high redshift cluster data, we find again the SSP and quenched models, as well as a model where strangulation begins at intermediate ages after a period of constant star formation, are preferred. They predict the recent faint-end build up of the red sequence, along with observed dwarf-to-giant ratios. Only the SSP model appears to predict the observed M/L evolution, but only if selection effects are carefully modeled.

Using the Chandra X-ray Observatory, Martini et al. (2006) found that the AGN fraction of galaxy clusters was five times higher than previous optical studies suggested. Using visual observations only, Dressler et al. (1985) estimated the AGN fraction of field galaxies to be 5%, while that of clusters was thought to be 1%. To understand the role that the environment plays in AGN fueling, the author studied a variety of environments, ranging from the field to groups to clusters. Will the AGN fraction of groups also be higher than that of the field? The author demonstrates how the AGN fraction of groups compares to that of clusters. In the following sections, the author describes the mechanics of X-ray astronomy, the group environment, and the characteristics of active galactic nuclei. The author briefly describes the possible mechanisms for AGN fueling.

We present an extremely deep CO(1-0) observation of a confirmed z = 1.62 galaxy cluster. We detect two spectroscopically confirmed cluster members in CO(1-0) with signal-to-noise ratio >5. Both galaxies have log (M-star/M-circle dot) > 11 and are gas rich, with M-mol/(M-star + M-mol) similar to 0.17-0.45. One of these galaxies lies on the star formation rate (SFR)-M-star sequence, while the other lies an order of magnitude below. We compare the cluster galaxies to other SFR-selected galaxies with CO measurements and find that they have CO luminosities consistent with expectations given their infrared luminosities. We also find that they have gas fractions and star formation efficiencies (SFE) comparable to what is expected from published field galaxy scaling relations. The galaxies are compact in their stellar light distribution, at the extreme end for all high-redshift star-forming galaxies. However, their SFE is consistent with other field galaxies at comparable compactness. This is similar to two other sources selected in a blind CO survey of the HDF-N. Despite living in a highly quenched protocluster core, the molecular gas properties of these two galaxies, one of which may be in the process of quenching, appear entirely consistent with field scaling relations between the molecular gas content, stellar mass, star formation rate, and redshift. We speculate that these cluster galaxies cannot have any further substantive gas accretion if they are to become members of the dominant passive population in z < 1 clusters.

Bright Ly alpha blobs (LABs)-extended nebulae with sizes of similar to 100 kpc and Ly alpha luminosities of similar to 10(44) erg s(-1)-often reside in overdensities of compact Ly alpha emitters (LAEs) that may be galaxy protoclusters. The number density, variance, and internal kinematics of LABs suggest that they themselves trace group-like halos. Here, we test this hierarchical picture, presenting deep, wide-field Ly alpha narrowband imaging of a 1 degrees x. 0 degrees.5 region around a LAB pair at z = 2.3 discovered previously by a blind survey. We find 183 Lya emitters, including the original LAB pair and three new LABs with Ly alpha luminosities of (0.9-1.3) x 10(43) erg s(-1) and isophotal areas of 16-24 arcsec2. Using the LAEs as tracers and a new kernel density estimation method, we discover a large-scale overdensity (Bootes J1430+3522) with a surface density contrast of delta(Sigma) = 2.7, a volume density contrast of delta similar to 10.4, and a projected diameter of approximate to 20 comoving Mpc. Comparing with cosmological simulations, we conclude that this LAE overdensity will evolve into a present-day Coma-like cluster with log(M/M-circle dot) similar to 15.1 +/- 0.2. In this and three other wide-field LAE surveys re-analyzed here, the extents and peak amplitudes of the largest LAE overdensities are similar, not increasing with survey size, and implying that they were indeed the largest structures then and today evolve into rich clusters. Intriguingly, LABs favor the outskirts of the densest LAE concentrations, i.e., intermediate LAE overdensities of delta(Sigma) = 1-2. We speculate that these LABs mark infalling protogroups being accreted by the more massive protocluster.

We present a study of the relation between galaxy stellar age and mass for 14 members of the z = 1.62 protocluster IRC 0218, using multiband imaging and HST G102 and G141 grism spectroscopy. Using UVJ colors to separate galaxies into star-forming and quiescent populations, we find that, at stellar masses M*>= 10(10.85)M circle dot the quiescent fraction in the protocluster is f(Q) = 1.0(-0.37)(+0.00), consistent with a similar to 2x enhancement relative to the field value, f(Q) = 0.45(-0.03)(+0.03). At masses 10(10.2)M circle dot <= M* <= 10(10.85)M circle dot, f(Q) in the cluster is f(Q) = 0.40(-0.18)(+0.20), consistent with the field value of f(Q) = 0.28(-0.02)(+0.02). Using galaxy D-n(4000) values derived from the G102 spectroscopy, we find no relation between galaxy stellar age and mass. These results may reflect the impact of merger- driven mass redistribution-which is plausible, as this cluster is known to host many dry mergers. Alternately, they may imply that the trend in f(Q) in IRC 0218 was imprinted over a short timescale in the protocluster's assembly history. Comparing our results with those of other high- redshift studies and studies of clusters at z similar to 1, we determine that our observed relation between f(Q) and stellar mass only mildly evolves between z similar to 1.6 and z similar to 1, and only at stellar masses M* <= 10(10.85) M circle dot Both the z similar to 1 and z similar to 1.6 results are in agreement that the red sequence in dense environments was already populated at high redshift, z greater than or similar to 3, placing constraints on the mechanism(s) responsible for quenching in dense environments at z >= 1.5.

We present a combined strong and weak lensing analysis of the J085007.6+360428 (J0850) field, which contains the massive cluster Zwicky 1953. This field was selected for its high projected concentration of luminous red galaxies. Using Subaru/Suprime-Cam BVR(c)I(c)i'z' imaging and MMT/Hectospec spectroscopy, we first perform a weak lensing shear analysis to constrain the mass distribution in this field, including the cluster at z = 0.3774 and a smaller foreground halo at z = 0.2713. We then add a strong lensing constraint from a multiply imaged galaxy in the imaging data with a photometric redshift of z approximate to 5.03. Unlike previous cluster-scale lens analyses, our technique accounts for the full three-dimensional mass structure in the beam, including galaxies along the line of sight. In contrast with past cluster analyses that used only lensed image positions as constraints, we use the full surface brightness distribution of the images. This method predicts that the source galaxy crosses a lensing caustic, such that one image is a highly magnified "fold arc" that could be used to probe the source galaxy's structure at ultra-high spatial resolution (< 30 pc). We calculate the mass of the primary cluster to be M-vir = 2.93(-0.65)(+0.71) x 10(15) M-circle dot. with a concentration of C-vir = 3.46(-0.59)(+0.70), consistent with the mass-concentration relation of massive clusters at a similar redshift. The large mass of this cluster makes J0850 an excellent field for leveraging lensing magnification to search for high-redshift galaxies, competitive with and complementary to that of well-studied clusters such as the HST Frontier Fields.