Bears and wolves are large mammalian predators who fill similar biological niches and have acquired similar cultural significance throughout Western history. Although superficial similarities exist between them in Charles G. D. Roberts's short animal stories, Roberts uses anthropomorphism to differentiate between these two species. This thesis uses a historical-cultural approach to provide the context for determining what was known or believed about these animals during Roberts's life and what contemporaneous theories were likely to have influenced Roberts's writing. The present literary analysis of bears and wolves in Roberts's stories shows that the species are primarily differentiated through the degree of anthropomorphism attributed to their individual members. Roberts anthropomorphizes bears more than his other species, and this contributes to the bears representation of the positive potential of animality. By contrast, Roberts minimizes anthropomorphization of wolves, who represent the negative potential of animality. In Roberts's work, humans who live in the wilderness must become either bear-like or wolf-like. Those who embrace bears positive animal potential are those who belong in the natural world, while those who practice the wolves' negative animal potential are denied a place in the natural order. Humans ultimately prove themselves to be superior animals through their use of technology, but must also demonstrate positive qualities, such as morality, in order to show that they belong in nature. Roberts's binary of animality speaks to a conflict that continues today, between the desire to accept the animality that is part of human nature and simultaneously to deny the baser aspects of that animality.

Perceived predation risk alters animals’ behavior. This shift in behavior often comes at the cost of attaining resources. Generally, African elephants (Loxodonta africana) experience little predation pressure; however, the risk of predation by lions (Panthera leo) increases other prey species are less abundant. In elephant herds, related females and their offspring travel together in family groups, led by the eldest female. Response to predation pressure was examined by playing lion calls to the population of 437 elephants at the Main Camp Section of Addo Elephant National Park (AENP) in South Africa. Unfamiliar lion calls from a single male and two males, static, and running water were played from a remote speaker to elephants at waterholes. These trials were recorded by video. Behaviors of elephants were then extracted from video into focal observations of thirty second segments before, during and after a sound was played. I analyzed these data using parametric t-tests and non-parametric randomization tests. When no sound was played, elephants did not alter their behavior. Water elicited low levels of distress behaviors. Elephants behaved in a threatened or annoyed manner toward static. Elephants changed their behavior more in response to lion calls than to the controls, namely by decreasing drinking and increasing walking and distress behaviors. I also examined how individuals differed in their responses to the lion calls based on a number of demographic factors. Adult and subadult females performed more social behaviors after lion calls when the matriarch was absent than when she was present. Furthermore, when group size was larger and more calves were present, females decreased drinking and increased time exhibiting distress behaviors. Based on this and other studies it can be concluded that elephants of different demographics perceived similar levels of elevated risk when hearing lion calls. Landscape of fear models are useful for assessing habitat use by prey species in response to real and perceived predation risk. The present study corroborates findings from a study in East Africa that elephants perceive threat from lions based on calls alone and appear to distinguish levels of threat by the number of lions calling.

Predation is a strong dective force on invertebrate prey. Asellus aquaticus differs in pigmentation reed and submerged vegetation habitats in lakes. Light pigmented individuals al vegetation dominated by Chara sp. while dark pigmented individuals dominate in the reeds. These differences have been hypothesized to result from background matching. Predation pressure from fish is belived to be highest in Chara sp., while invertebrate predators are more common in the reeds. In this study I investigatedif predation from perch and damselfly larvae create different se1ection pressure on pigmentation and size of the Asellus, and if selection is affected by the structure of the habitat. The study vas carried out in aquaria in a loboratory. Regarding predation from perch a tendancy to be selective against dark pigmented individuals in Chara substrate was seen. Mortality increased with body size in Asellus, regardless of pigmentation. In the experiment mortality of Asellus decreased with body length. The results illdicate that it is not as important for Asellus to be cryptic in the reeds since it is not exposed to visual predation on the same level as in the Chara sp. where it is preferable to be small and cryptic. In the reed, large individuals are probably favorued since common predators, such as damselfly larvae, are size-limited in their prey choice.

The tropical palm Mauritia flexuosa has highly nutritious fruits and is an important food resource for both humans and wildlife throughout its geographic range in South America. Unsustainable harvesting threatens wild populations. Mauritia f. occurs primarily in wetlands called Aguajales where it can become the dominating canopy species. Seed predation and dispersal can dramatically affect the survival and distribution of plant species in tropical rainforests (Janzen 1970, Connell 1971, Bleher & Böhning-Gaese 2001, Paine & Beck 2007, Mari et al. 2008). Increased knowledge of seed predation and germination requirements is essential for successful management of this commercially and ecologically important palm. Four experiments were conducted in Manu National Park in southeastern Peru to study: (1) Seed survival in the Aguajal, (2) Quantify seed predators on dry land, (3) Insect visitors and consumers of fruits and seeds, and (4) Germination in greenhouse experiments. Seed survival was significantly higher below water compared to on dry micro sites within the Aguajal. Seeds and fruits placed on dry land were preyed upon by both insects and mammals. Terrestrial insects were the most important predators. Different insects visited fruits and seeds, indicating a successive breakdown of different tissues. Seed survival was also higher below water and/or soil in the greenhouse experiment. This may suggest that the distribution of Mauritia f. is highly influenced by seed predation and that water protects seeds from their insect enemies.

The jaguars (Panthera onca) of Tortuguero National Park, Costa Rica, sometimes kills and eats green sea turtles (Chelonia mydas), they also, though less often, kill and eat leatherback sea turtles (Dermochelys coriacea). The three species are considered endangered and are listed in CITES. It was the aim of this study to find out more about the jaguars behaviour in the area. To discern any patterns of jaguar and turtle activity on the beach the number of tracks per eighth of a mile was recorded on a daily basis for 26 days and then analyzed. It was also considered to be of interest to determine how many jaguars could be responsible for the predation of sea turtles. In addition to this average beach width was measured for each eighth of a mile. There was a noticeable difference in jaguar activity on the beach between days of recording. Analysis found that the beach width could possibly have a small positive effect on jaguar activity. No correlation was found between jaguar and turtle activity. It is believed that the reason that there was no correlation between jaguar activity and turtle activity was due to most of the tracks used to estimate turtle activity had originated from leatherback turtles, which are not as often predated by jaguars as the green turtle. An estimation of five or six jaguars was made using photographs of pugmarks and a method of track discrimination together with information from personnel from the Jalova station.

The aquatic isopod Asellus aquaticus can in some lakes be found as two different ecotypes; one in the habitat dominated by reed and one in stonewort stands. These ecotypes have been shown to differ in size, color and behavior. The reed ecotype is larger, darker and more active compared to the stonewort ecotype. In the two habitats there are different dominating predators: Invertebrate predators in the reed habitat and fish in the stonewort habitat. This project aims to examine how the presence of invertebrate predators and fish affect the two ecotypes of the isopod in regard to behavior and size composition in different substrates. To examine the effect on behavior the activity of isopods collected from Lake Tåkern was measured without and with chemical cues from perch and damselfly larva. The result of the behavior experiment showed no difference between the ecotypes nor the treatments. The lack of differences in the behavior indicates that there could be a variation between lakes. The effect on the size composition was examined by subjecting groups of isopods to predation by perch or damselfly larva in different substrate. The mean length of the group was measured before and after the trials. The size decreased significantly for the reed ecotype in stonewort substrate when subjected to predation by perch. The size for the stonewort ecotype increased significantly in reed substrate with damselfly larva as predator. The effect on size supports that the predators are the cause of the size difference between the ecotypes.

Growth rate is an important life history trait, which impacts fitness indirectly through its effect on the age and size at maturity, as well as directly through costs associated with accelerated growth such as increased predation risk. Genetic variation and plasticity in growth are widespread in nature, and local adaptation of growth rate may evolve due to divergent selection in different environments, for example related to predation risk, temperature or time constraints. I studied local adaptation of larval life history in the moor frog Rana arvalis, in a local network of ponds close to Uppsala. Local adaptation of growth rate and survival was studied in a reciprocal transplant experiment between ponds with different habitat characteristics. Meanwhile, differences among the populations in intrinsic growth, activity and response to predation were studied in a common garden experiment in the laboratory, where tadpoles were raised in the presence or absence of a predator and tested in direct predation trials. In the field, differences in growth among populations were found, independent of which pond the tadpoles were raised in, indicating that the ponds were similar growth environments. Survival differences among the populations depended on the pond, but local populations did not do better than foreign ones. In the laboratory, similar patterns in growth rate were found. All populations were highly plastic in their response to predation, having lower growth and activity in the predator-induced treatment and decreased mortality in the predation trials. Tadpole size was an important factor in escaping predation. One population clearly grew faster than the others in the field and in the lab, which could be explained in terms of its habitat of origin but was most likely related to the relatively late hatching of this population. Future studies are necessary concerning the possible costs of this accelerated growth and the importance of breeding phenology. Apart from the one differential population, I did not find evidence of local adaptation in the field or in the laboratory. The influence of habitat characteristics on tadpole life history was difficult to study, due to the limited number of ponds and many environmental differences among them. However, this thesis was a valuable pilot study concerning the design of experiments to study factors promoting and constraining local adaptation in landscape mosaics. An understanding of local adaptation at the scale at which gene flow occurs is important for the conservation of populations in fragmented landscapes as well as for the study of ecological speciation.

Bears and wolves are large mammalian predators who fill similar biological niches and have acquired similar cultural significance throughout Western history. Although superficial similarities exist between them in Charles G. D. Roberts's short animal stories, Roberts uses anthropomorphism to differentiate between these two species. This thesis uses a historical-cultural approach to provide the context for determining what was known or believed about these animals during Roberts's life and what contemporaneous theories were likely to have influenced Roberts's writing. The present literary analysis of bears and wolves in Roberts's stories shows that the species are primarily differentiated through the degree of anthropomorphism attributed to their individual members. Roberts anthropomorphizes bears more than his other species, and this contributes to the bears representation of the positive potential of animality. By contrast, Roberts minimizes anthropomorphization of wolves, who represent the negative potential of animality. In Roberts's work, humans who live in the wilderness must become either bear-like or wolf-like. Those who embrace bears positive animal potential are those who belong in the natural world, while those who practice the wolves' negative animal potential are denied a place in the natural order. Humans ultimately prove themselves to be superior animals through their use of technology, but must also demonstrate positive qualities, such as morality, in order to show that they belong in nature. Roberts's binary of animality speaks to a conflict that continues today, between the desire to accept the animality that is part of human nature and simultaneously to deny the baser aspects of that animality.

Benthic invertebrates play important roles as feeding resources for many organisms in different food webs. Shifts in predation of these organisms can generate cascading effects and potentially lead to the disappearance of one or more species from a site. Cascading effects can bring impacts to organisms who aren’t even directly involved, why studies in this field are important for understanding sudden changes in ecosystems. I examined the predation from fish and waterfowl on benthic invertebrates in the shallow and eutrophic Lake Tåkern in the plains of Östergötland County, southern Sweden. The study was experimental and used exclosures (three types, eight replicates) in the shape of 130-liter cages to examine the composition of invertebrates. It was conducted during early spring, a relatively unexamined period for this kind of study. I found no significant differences in the control treatment compared to any of the other treatments regarding biodiversity. The invertebrate fauna was dominated by a small number of species, with a relatively patchy spread throughout the sediment area. Furthermore, the results indicate that the fish hadn’t had the time to properly activate their predatory habits due to prolonged winter temperatures. The waterfowl were considered too absent during the test period to affect the invertebrate community. Future studies in this area are recommended to work with a greater sampling area, to reduce the influence of extreme values.

The northern fur seal (NFS) population on the Pribilof Islands (PI) is currently declining while the population on the Commander Islands (CI which includes Bering and Medny Islands) is stable. The reasons for the different population trajectories remain unknown. Comparing differences in behavioral ecology and predation pressure between these two populations could provide an explanation. This study examined lactating NFS female behavior to determine: 1) summer foraging patterns (trip duration, trip direction, dive depth) of animals from two nearby rookeries on Bering Island, 2) winter migration from Medny and Bering Islands relative to patterns of ocean productivity, and 3) the potential impact of killer whale predation on population dynamics. Data were collected from 2003 to 2010 using visual observations and telemetry. Twenty-one satellite transmitters, 29 time-depth recorders and 17 geolocation recorders were deployed. Shore-based observations of killer whale predation and photo-identification were conducted near the CI rookeries in 19992010. During lactation, both mean foraging trip duration and mean maximum diving depth (3.4 plus/minus 1.3 days and 17.7 plus/minus 6.8 m, respectively) for NFS adult females (n = 28) did not significantly change among years. Although foraging areas of NFS from the two rookeries on Bering Island overlapped, the mean direction of travel from Severo-Zapadnoe rookery was significantly (p<0.01) different compared with Severnoe rookery. The foraging patterns suggested that these females had a reliable food source that did not change despite potential environmental changes or the effects of fisheries. During their winter migration, NFS females from the CI traveled to the Transition Zone Chlorophyll Front (32° N-42° N) in the North Pacific Ocean. Their winter migration routes and the location of overwinter foraging areas were positively correlated with high ocean productivity (near surface chlorophyll a concentration). Over 82 percent (n=17) of these females spent 38 months near the eastern coast of Hokkaido, Japan and followed the coastal high productivity areas on their way back to the CI. Transient killer whales in groups of 2-12 individuals were repeatedly observed preying mostly on NFS males during the summer. The simulation model showed little impact on population dynamics as long as male fur seals were the primary prey. However, if the number of killer whales increased or they changed their diet to include females and pups, then the NFS population on the CI could decline. The winter migration of NFS from CI and PI are similar. Lactating NFS from the PI exhibit greater summer foraging effort (longer average trip duration and bout duration; greater number of deep dives) compared with females from the CI.