In spite of the relative simple vegetation structure, the Grassland biome has surprisingly high species diversity. The Grassland biome is also the most transformed biome in South Africa, with cultivation having the largest impact. When croplands are abandoned, secondary succession leads to low diversity Hyparrhenia hirta dominated grassland. A combination of two seed mixtures, two seeding densities and two establishment methods was established in plots on a recently abandoned cropland at Suikerbosrand Nature Reserve to evaluate their effect on secondary succession. The rip plots, where more resources were available between the rip lines, have shown higher densities of relic weeds as well as local perennials, showing some progressive successional movement. However, Hyparrhenia hirta was one of the non-sown perennials increasing in the rip plots. Hyparrhenia invasion and relic weeds were best controlled in the plough plots. Although Hyparrhenia was successfully controlled in plough plots, no secondary succession occurred in these treatments. / Agriculture, Animal Health & Human Ecology / M. Tech. (Nature Conservation)

Thesis (PhD)--Stellenbosch University, 2012. / ENGLISH ABSTRACT: The restoration of natural capital has ecological, hydrological and economic benefits. Are these benefits greater than the costs of restoration when compared across a range of dissimilar sites? This study examines the impact of restoration at eight case study sites distributed throughout South Africa. The benefits of restoration include improved grazing values and crop yields, improvements in water yield and quality, soil carbon improvements, wild products, lumber, fuelwood and electricity. The impact of restoration on all forms of natural capital (i.e. cultivated, replenishable, renewable and non-renewable) is therefore quantified. The costs of restoration include depreciation on capital expenditure, labour costs, equipment and bond refinancing costs. The literature review done during this study presents three frameworks. The first framework classifies social science using the classification scheme of Burrell and Morgan. It shows that system dynamics modelling and neoclassical economics share the same epistemological and ontological characteristics, both of these fall within the naturalistic paradigm, which also characterises most of scientific research. System dynamics modelling and neoclassical economics, however, digress in the Flood and Jackson classification scheme, which is the second framework for classifying social science. Neoclassical economics is characterised by a small number of elements and few interactions between the elements. Systems dynamics modelling, on the other hand, is characterised by a large number of elements and many interactions between the elements. The nature-freedom ground motive is subject to a number of criticisms, including the fact that it introduces dualistic thinking into the analysis, as well as that it does not adequately address normative or moral issues. The framework of Dooyeweerd, the third framework, is presented as a means of transcending the nature-freedom ground motive. Although the nature-freedom ground motive is largely utilised in this study, the analysis does transcend the traditional economic approach in a number of areas. These include, for example, a focus on transdisciplinary methods, disequilibria, adopting a case study approach, and empirical estimation instead of theoretical models. The restoration case studies in this study are examples of individual complex systems. Eight system dynamics models are developed to model interactions between the economic, ecological and hydrological components of each of the case studies. The eight system dynamics models are then used to inform a risk analysis process that culminates in a portfolio mapping exercise. This portfolio mapping exercise is then used to identify the characteristics and features of the different case study sites based on the risk profile of each sites. This study is the first known application of system dynamics, risk analysis and portfolio mapping to an environmental restoration project. This framework could potentially be used by policymakers confronted with budgetary constraints to select and prioritise between competing restoration projects. / AFRIKAANSE OPSOMMING: Die restorasie van natuurlike kapitaal het ekologiese, hidrologiese en ekonomiese voordele. Maar is hierdie voordele groter as die kostes verbonde aan restorasie wanneer dit oor verskeie ongelyksoortige terreine vergelyk word? Hierdie studie bestudeer die impak van restorasie op agt verskillende studie terreine versprei regoor Suid-Afrika. Die voordele van restorasie sluit die volgende in: beter weiding waardes en oes opbrengste, verbeterde water lewering en water kwaliteit, verbetering van grondkoolstof, wilde produkte, hout, brandstofhout en elektrisiteit. Die impak van restorasie op alle vorme van natuurlike kapitaal (gekultiveerd, aanvulbaar, hernubaar en nie-hernubaar) is daarom gekwantifiseer. Die kostes van restorasie sluit in ‘n vermindering in kapitaal uitgawes, arbeidskoste, toerusting en verband herfinansieringskoste. Die literatuurstudie hou drie raamwerke voor. Die eerste raamwerk klassifiseer sosiale wetenskappe volgens die Burrel en Morgan klassifikasie skema. Dit wys daarop dat dinamiese stelsel modellering en neoklassieke ekonomie dieselfde epistemologiese en ontologiese eienskappe deel; beide val binne die naturalistiese paradigma, wat dan ook meeste wetenskaplike navorsing tipeer. Stelseldinamiese modellering en neoklassieke ekonomie wyk egter af na die Flood and Jackson klassifikasie skema, wat die tweede raamwerk is waarvolgens sosiale wetenskappe geklassifiseer word. Neoklassieke ekonomie word gekenmerk aan 'n klein aantal elemente en 'n beperkte hoeveelheid interaksie. Stelseldinamiese modellering het egter 'n groot aantal elemente met veel meer interaksies tussen hierdie elemente. Die natuur-vryheid grondmotief is onderworpe aan 'n aantal punte van kritiek, insluitende die feit dat dit dualistiese denke in analise inbring. Verder spreek dit ook nie voldoende die normatiewe of morele kwessies aan nie. Die raamwerk van Dooyeweerd, wat dan die derde raamwerk is, word voorgestel as 'n wyse waarop die natuur-vryheid grond-motief getransendeer kan word. Alhoewel die natuur-vryheid grondmotief grootliks gebruik word in hierdie studie, transendeer die analise die tradisionele ekonomiese benadering op 'n aantal gebiede. Hierdie gebiede sluit die volgende in: 'n fokus op transdissiplinere metodes, onewewigtigheid, 'n gevallestudie benadering, en empiriese skatting in plaas van teoretiese modelle. Die restorasie gevallestudies wat in hierdie studie gebruik word is voorbeelde van individuele komplekse sisteme. Agt dinamiese stelsel modelle word ontwikkel om die interaksies tussen ekonomiese, ekologiese en hidrologiese komponente in elke gevallestudie te modelleer. Hierdie agt stelseldinamiese modelle word dan gebruik in 'n risiko analise proses wat uitloop op 'n portefeulje plot oefening. Hierdie portefeulje plot oefening word dan gebruik om eienskappe en kenmerke van verskeie gevallestudie terreine te identifiseer gebaseer op die risiko profiel van elke terrein. Hierdie studie is die eerste bekende toepassing van dinamiese stesels, risiko analise en portefeulje plot tot 'n omgewingsrestorasie projek. Hierdie raamwerk kan potensieël gebruik word deur beleidskrywers wat met begrotings beperkinge gekonfronteer word om tussen restorasie projekte te kies en om hulle te prioritiriseer.

The main aim of this study was to test rehabilitation interventions in a system altered by heavy grazing, and to develop methods of monitoring these interventions.

The main aim of this study was to test rehabilitation interventions in a system altered by heavy grazing, and to develop methods of monitoring these interventions.

In spite of the relative simple vegetation structure, the Grassland biome has surprisingly high species diversity. The Grassland biome is also the most transformed biome in South Africa, with cultivation having the largest impact. When croplands are abandoned, secondary succession leads to low diversity Hyparrhenia hirta dominated grassland. A combination of two seed mixtures, two seeding densities and two establishment methods was established in plots on a recently abandoned cropland at Suikerbosrand Nature Reserve to evaluate their effect on secondary succession. The rip plots, where more resources were available between the rip lines, have shown higher densities of relic weeds as well as local perennials, showing some progressive successional movement. However, Hyparrhenia hirta was one of the non-sown perennials increasing in the rip plots. Hyparrhenia invasion and relic weeds were best controlled in the plough plots. Although Hyparrhenia was successfully controlled in plough plots, no secondary succession occurred in these treatments. / Agriculture, Animal Health and Human Ecology / M. Tech. (Nature Conservation)

The Working for Water (WfW) Programme is a public works programme designed to clear South Africa of invasive alien vegetation and to restore lowwater consuming indigenous vegetation in the areas that have been cleared. Funds to clear alien invasives were initially secured on the basis that such a programme would increase water runoff, facilitate biodiversity and ecosystem functioning, and provide social benefits through job creation. The economic merits of the Programme, in terms of increased water yields, has been established in the Western Cape and KwaZulu-Natal, but questioned in the Eastern and Southern Cape. However, there are economic aspects of the studies carried out in the Eastern and Southern Cape that merit more attention than was given them; one of these being the issue of non-water benefits. Preliminary figures emanating from contingent valuation pilot studies conducted at six WfW projects sites indicated that one of these non-water benefits, namely the biodiversity and ecosystem resilience benefit, could be substantial. As such, the primary objective of the present study was to apply the contingent valuation method (CVM) to value people’s preference for indigenous vegetation. This value was intended to serve as a proxy for increased biodiversity and ecosystem resilience at three WfW sites. Despite the controversy surrounding the CVM, it has been found that it is a credible valuation tool. The CVM’s merits lie in its versatility and in the fact that it is the only method available which is capable of obtaining estimates of both nonuse and use values, thus making it applicable for valuing biodiversity. The primary aim of a CVM study is to determine an estimate of the total willingness-to-pay (WTP). In this study, the total WTP figure was calculated by multiplying the median WTP for the local WfW Programme by the total number of user households. The respective total WTP amounts are shown in Table 1. It was anticipated that respondents would be willing to pay more for the national WfW Programme, than for the less inclusive good (i.e. the local WfW Programme). The results correspond with this expectation at the Port Elizabeth and Underberg sites. However, due to strategic factors Worcester respondents were willing to pay more for the local WfW Programme than for the national Programme.

Rehabilitation is a dynamic process influenced by factors related to more than one field of ecology. It is therefore necessary to consider all these components when assessing the rehabilitation, although in the initial stages the successful revegetation of the disturbed areas is the most important criterion. Richards Bay Minerals, on whose mining site this project was carried out, is dredge mining heavy minerals on the north coast of Natal, where they have rehabilitated mined areas since 1978. This project has been carried out to establish: 1) The success of their dune forest rehabilitation using quantitative techniques. 2) The available seed bank in their rehabilitation stands. 3) The similarities in the succession taking place in rehabilitation stands compared to the revegetated stands in the vicinity of Richards Bay. 4) The best methods for creating alternative vegetation communities, especially grasslands, with a high species diversity on the mined tailings. This study reviews only the success of rehabilitation of the natural vegetation but other studies focusing on the insect, reptile, mammal and bird populations are also being undertaken by other researchers. No particular method of determining the success of vegetation rehabilitation has been chosen by restoration ecologists. Therefore in this study a broad range of quantitative techniques were used to show whether successional changes are occurring in the vegetation and physical environment. The results obtained from sampling the rehabilitated vegetation have shown that both the species richness and diversity are increasing as the returned vegetation matures. Levels of soil properties such as Sodium, Phosphate, Calcium and percentage organic matter have also risen with increasing stand age. Community complexity is also increasing with stand age, and TWINSPAN and DECORANA plots have separated out the differently aged stands based on their differences. A "pilot" study was done on the seed bank present in the rehabilitation stands. This has shown the presence of large amounts of early successional, mostly herbaceous species. Seeds of later successional and woody species were scarce which may be a result of the sampling intensity used. However seeds of late successional ground cover species were found in the older stands. Comparisons between the natural revegetation of disturbed areas in the vicinity of Richards Bay and the rehabilitation stands revealed similarities in both species composition and complexity. Species richness and diversity values are comparatively similar for the younger revegetated and older rehabilitation stands, and lWlNSPAN and DECORANA analysis techniques clustered the samples recorded from these areas in close proximity on their relative plots. The oldest revegetated sites contain a number of species found in the rehabilitated vegetation but as Acacia karroo has thinned-out in these stands many of these other species are now mature individuals. Attempts at rehabilitating an area of grassland at Richards Bay Minerals has not produced satisfactory species diversity and experimental manipulations were used to try and increase the diversity of the existing Eragrostis curvula dominated community. Of the several treatments used for the manipulation, a combination of burning and further topsoiling was the most successful in reducing Eragrostis importance and in increasing the species richness. Grassland topsoil spread directly onto the bare tailings produced an extensive vegetation covering over a short period but species richness was not significantly greater than for the existing Eragrostis dominated grassland, and further treatments and management needs to continue if this technique is to be employed. Only a limited amount of alien infestation of the rehabilitated areas was evident from the sampling undertaken during this research. As the removal of alien plants is an ongoing process and the rehabilitation stands are continuously monitored to identify any new invaders, this is not expected to become a problem. From the results of work done overseas and the rehabilitation carried out in South Africa it appears that it is possible to return natural vegetation communities on mined areas. That this is a lengthy process is to be expected but by manipulating the vegetation and continuously monitoring the process it may be possible to speed up development. Areas in need of further research have been identified based on the findings of this project. This will help to reinforce the undertaking of management proposals that will enhance the vegetation recovery and the success of the rehabilitation programme.

Thesis (MScConsEcol(Conservation Ecology and Entomology)--University of Stellenbosch, 2006. / Anglo American Corporation’s Namakwa Sands heavy minerals mining and beneficiation operation has been strip-mining a heavy mineral deposit, rich in the commercially valuable minerals ilmenite, rutile and zircon, since September 1994. The mine is located in the vicinity of Brand-se-Baai on the west coast of South Africa, approximately 385 km north of Cape Town. Strip-mining causes total destruction of natural ecosystems through the removal of vegetation and soil in the area where mining is being undertaken. Namakwa Sands has been rehabilitating mined out areas as the mining front moves forward. Due to the difficulty of rehabilitating mined out areas as a result of harsh environmental factors, Namakwa Sands has initiated various research projects to gain an understanding of the baseline conditions and ecosystem function in order to increase plant cover and biodiversity on post-mined areas. This on-going research and the development of rehabilitation and mining techniques have resulted in the implementation of four rehabilitation techniques varying in investment of topsoil replacement, seeding and plant translocation. This study assesses the success and effectiveness of these techniques in terms of various vegetation and soil parameters. In addition, those parameters that are considered useful for monitoring are identified. This study indicated that topsoil replacement and plant translocation facilitate the return of similarity, species richness, species diversity and vegetation cover to post-mined areas. The rehabilitation site that had the greatest amount of biological input (topsoil replacement and plant translocation) appeared to be the most successful technique in facilitating vegetation recovery similar to reference sites. In comparison, the site that had the least amount of biological input performed the worst and requires adaptive management, e.g. reseeding and / or plant translocation. Namakwa Sands should continue to replace topsoil in all future rehabilitation efforts and, when possible (e.g. after sufficient winter rain), continue to translocate species in multi-species clumps. In terms of species selected for translocation, Othonna cylindrica, Ruschia versicolor and Lampranthus suavissimus should be considered for future large-scale translocation projects. Zygophyllum morgsana appears to be more difficult to re-establish under the current climatic conditions (below average rainfall). The long-term viability of rehabilitated Z. morgsana populations needs to be determined before considering this species for any future large-scale translocation purposes. No translocated Asparagus spp. individuals survived and should therefore not be considered for any further translocation purposes. The grass Ehrharta calycina, which is dominant in the site seeded, should continue to be considered for future seeding. Species and functional diversity appear to be the most limiting factors within all the rehabilitation sites and Namakwa Sands will not be able to meet their long-term objective of small-stock farming if diversity and the number of palatable species do not increase significantly. Adaptive management should seriously be considered in order to speed up this process. Alternatively, an appropriate grazing strategy, which is related to the Tetragonia fruticosa dominated vegetation within rehabilitation sites, would need to be determined and adopted.More time is needed to ameliorate the rehabilitated soil profiles to the same level as in reference sites, especially with regard to carbon, pH and sodium levels. In order to increase organic matter within rehabilitation areas, Namakwa Sands should consider creating clumps with cleared vegetation from the mining front. Since the long-term rehabilitation goal has not been achieved, Namakwa Sands will need to continue to monitor plant and soil changes until it has been achieved. The objectives of the current rehabilitation programme are limited and Namakwa Sands should develop additional objectives relating to the structure and function of the natural vegetation. This will give a better indication of whether rehabilitation sites are progressing towards the desired end point and if adaptive management is required. In addition, the current monitoring programme (vegetation survey) implemented at Namakwa Sands could be improved by increasing the vegetation parameters to be monitored. It is recommended that the following vegetation parameters be monitored as part of the long-term monitoring programme: species composition and similarity, species richness, species diversity, vegetation cover, species dominance, vertical structure and functional diversity of the vegetation (clumps and inter-clumps). It is also recommended that carbon, pH and sodium of soil profiles be monitored as part of the long-term monitoring programme. These parameters should not be seen as exhaustive as this study only considered various vegetation parameters and soil chemistry between rehabilitation and reference sites. The results of other studies on the fauna, mycorrhiza, insects, etc. should also be taken into consideration and the monitoring parameters expanded accordingly.

Thesis (MScFor )--Stellenbosch University, 2003. / ENGLISH ABSTRACT: Species-rich Renosterveld vegetation does not return to old agricultural fields even after many years (10-20) of abandonment. In most cases the fields are slowly taken over by non-indigenous species, particularly alien pasture grasses and African lawn grass. While poor survival of indigenous seedlings due to competition, grazing or any other establishment constraints, might explain the failure of natural vegetation to colonise old fields, here I am testing the hypothesis that recolonisation by indigenous plant species is limited by seed dispersal. I have quantified changes in seed density with distance from the natural vegetation into old fields using seed traps and soil seed bank assessment. I have also looked at seed dispersal in the dung of large herbivores. Shrubs (with the exception of Eytropappus rhinocerotis) were poorly represented in the seed rain indicating that they might be seed limited. Dicerothamnus rinocerotis (a dominant shrub in Renosterveld) and certain geophytes (that were recovered in the soil bank) have proven that seed dispersal is not the primary constraint to their return to ploughed fields. Dicerothamnus rinocerotis produces a large number of seeds that are dispersed long distances into the old field and a number of geophytes were well represented in the soil bank. Large herbivores are dispersing a lot of herbaceous forbs in their dung but make very little contribution to the dispersal of indigenous tussock grass and shrubs. Seeds of indigenous Renosterveld tussock grasses (e.g Tribolium hispidum) were restricted to the edge of the natural vegetation. My research has shown that some indigenous seeds, particularly those dispersed in wind and through large mammals, do reach the old ploughed field. I therefore conclude that seed dispersal is not the primary constraint to the natural recovery of populations of some indigenous plant species on old fields, but may limit recovery of certain indigenous geophytes, grasses and shrubs. / AFRIKAANSE OPSOMMING: Spesie-ryke Renosterveld plantegroei hervestig nie op ou landerye nie, selfs ná baie jare (l0-20) van verlating. Oor die algemeen word hierdie grond stadig oorgeneem deur uitheemse spesies, veral uitheemse weidingsgrasse en kweekgras. Terwyl swak oorlewing van inheemse saailinge as gevolg van kompetisie, weiding of enige ander vestigingsbeperkinge, die mislukte vestiging van natuurlike plantegroei op ou benutte grond kan verduidelik, toets ek hier die hipotese dat hervestiging deur inheemse plantspesies deur saadverspreiding beperk word. Ek het veranderinge in saaddigtheid oor afstand vanaf die natuurlike plantegroei na ou landerye gekwantifiseer, deur gebruik te maak van saadopvangsvalle en sand saadbank beraminge. Ek het ook gekyk na saadverspreiding in die mis van groot herbivore. Struike (behalwe Eytropappus rhinocerotis) was swak verteenwoordig in die "saadreën", wat mag toon dat struike saadbeperk is. Eytropappus rhinocerotis ('n dominante struik in Renosterveld) en sekere geofiete (wat in die saadbank gevind is) het bewys dat saadverspreiding nie die hoofbeperking is vir hulle terugkeer na geploegde lande nie. Eytropappus rhinocerotis produseer 'n groot hoeveelheid saad wat oor groot afstande oor die ou landerye versprei word en 'n aantal geofiete was goed verteenwoordig in die saadbank. Groot herbivore versprei 'n groot hoeveelheid kruidagtige forbe in hulle mis, maar maak 'n baie klein bydrae tot die verspreiding van inheemse polgras en struike. Saad van inheemse Renosterveld polgrasse (bv. Tribolium hispidum) was beperk tot die some van die natuurlike plantegroei. My navorsing het getoon dat sommige inheemse saad, veral dié wat deur die wind en deur groot herbivore versprei word, wel die ou geploegde landerye bereik. Ek maak dus die gevolgtrekking dat saadverspreiding nie die hoofbeperking is vir die natuurlike herstel van populasies van sommige inheemse plantspesies op ou landerye nie, maar mag die herstel van sekere inheemse geofiete, grasse en struike beperk.

Globally, ecosystems provide services of almost twice the value of global gross national product (Costanza et al., 2006). The Buffalo City Municipality (BCM), South Africa contains biodiversity of national and international importance (Pierce, 2003; Pierce et al., 2005). Despite this, the municipality continues to experience loss of both urban and rural biodiversity (Buffalo City Municipality, 2006a). This study sought to determine the status of biodiversity, and the potential for ecosystem services to contribute to conservation, within the BCM. Biodiversity features, including ecosystem type, species of special concern and biodiversity processes, were identified and mapped using a GIS to produce a biodiversity priority index for the BCM. Current transformation status was then mapped to determine the level of ecosystem degradation within the BCM. Priority biodiversity areas as well as individual biodiversity features were spatially overlain against current transformation status and protected areas and analysed using a GIS to determine the level of degradation and protection of BCM biodiversity. In total 3.5 % of total BCM biodiversity was protected. Of the 24 ecosystem types, 11 (45%) had less than 1% under protection, while 16 (67%) had less than five percent protected. Not restorable areas, thus completely lost to biodiversity conservation, comprised just less than a quarter of the total BCM area while un-impacted areas comprised just 12.3%. Twenty five ecosystem services were identified as being provided by intact natural ecosystems within the BCM. The natural capital providing these services was identified and mapped to produce an ecosystem service index (ESI) using a GIS. This ecosystem service index and the biodiversity priority index were overlain to determine their level of correlation. Overall ESI correlation with priority biodiversity was weak although several individual ecosystem services, including carbon sequestration, showed correlation. Using the above data layers an implementation plan and conservation framework was proposed to assist the coordination of local conservation action within the BCM. It is concluded that ecosystem services are a potentially useful tool for conservationists at the local level seeking to ensure that biodiversity has relevance to and receives protection from broader society.