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Red Alder (Alnus rubra) is a native coastal hardwood in British Columbia and has evolved a symbiotic relationship with the nitrogen-fixing actinomycete, Frankia. This research uses δ15N signatures in soils, wood and litter to assess the contribution of nitrogen-fixing Red Alder to the components of stand nitrogen budgets. The stands used in this study are part of the B.C. Ministry of Forests’ long-term Experimental Project 1121.01 which examines the interactions between conifers and Red Alder. Planted in 1994, the Holt Creek site contains stands of Douglas-fir and Red Alder in five proportions (Red Alder: Douglas-fir proportions: 100/0, 50/50, 25/75, 11/89, 0/100). Increment cores from 5 trees per species per plot were taken along with soil and litter samples and analyzed for essential mineral elements and δ15N. I hypothesized that Red Alder would enhance soil nitrogen stocks and elevate δ15N signatures and that these changes would be observable in the δ15N signature of the tree rings of both species. Forest floor soil under Red Alder in the 100/0 plot was enriched in total nitrogen, and δ15N was elevated. This was due to the addition of nitrogen-rich litter, like followed by nitrogen discrimination in the forest floor during the process of nitrate leaching or denitrification. The litter of the two species did not differ in δ15N. The effect of forest floor nitrogen enrichment was visible in the tree rings of Douglas-fir in the 50/50 stand confirming that the effect of fixed-nitrogen can be observed in non-fixing species. Red Alder tree ring δ15N exhibited an unexpected non-linear relationship with time that could be due to reduced nitrogen fixation associated with declining tree vigour or negative feedback from low soil pH. This research provides insight into nitrogen fixation by Red Alder over time and its influence on pure and mixed stand nitrogen budgets. / Graduate / 2023-09-09

Osteoarthritis (OA) of the knee is common in Western society. OA of the patellofemoral (PF) compartment of the knee is prevalent in adults greater than 55 years of age. Isolated radiographic PFOA is present in 13.6-24% of females and 11-15.4% of males with knee pain. Biomechanical factors such as tibiofemoral alignment and high joint stress are associated with the development and progression of PFOA. PF joint stress is high when the quadriceps contracts with the knee in a position of extreme flexion, such as rising from sitting. The purposes of this study were to determine 1) the triplanar biomechanics of the hips and knees during sit-to-stand (STS) for persons with PFOA versus age- and gender-matched control subjects, 2) the impact of PFOA on physical performance, perceived functional status, and pain, and 3) the relationship between knee kinematics during STS and physical performance. The biomechanics of STS was examined using a video-based motion analysis system and two force plates. Physical performance was measured with the Timed Up and Go (TUG) and Fifty-Foot Walk (FFW) tests. Perceived functional status, pain, and stiffness were measured using the Western Ontario and McMaster University Osteoarthritis Index (WOMAC) questionnaire. Knee pain was measured with a Visual Analog Scale (VAS) following STS, TUG, and FFW. Persons with PFOA required a significantly longer time to perform STS and the TUG as compared to healthy controls. The PFOA group demonstrated greater hip flexion and knee abduction versus the control group. The hip and knee moments were significantly different with the PFOA group demonstrating greater hip extension, hip abduction, hip external rotation, knee extension, and knee adduction moments. Persons with PFOA were found to have significantly less perceived physical function, greater stiffness, and greater pain. Pain following STS, TUG, and FFW were all greater in the PFOA group. No significant association was found between any knee angle and time to perform the TUG or the FFW. These results indicate that dynamic malalignment of the TF joint is present during STS in persons with PFOA. This may contribute to the increased pain and decreased function in persons with PFOA. / Physical Therapy

Self-thinning quantifies the reduction in tree numbers due to density-dependent mortality. Maximum size-density relationships (MSDRs) are a component of self-thinning that describe the maximum tree density per unit area obtainable for a given average tree size, often quadratic mean diameter (D). An MSDR species boundary line has been defined as a static upper limit of maximum tree density -- D relationships that applies to all stands of a certain species within a particular geographical area. MSDR dynamic thinning lines have been defined as the maximum tree density obtainable within an individual stand for a particular D which have been shown to vary relative to planting density. Results from this study show that differences in boundary levels of individual stands cause the MSDR species boundary line slope estimate to be sensitive to the range of planting densities within the model fitting dataset. Thus, a second MSDR species boundary line was defined whose slope is the average slope of all MSDR dynamic thinning lines. Mixed-models are presented as a statistical method to obtain an estimate of the population average MSDR dynamic thinning line slope. A common problem when modeling self-thinning is to determine what observations are within generally accepted stages of stand development. Segmented regression is presented as a statistical and less subjective method to determine what observations are within various stages of stand development. Estimates of D and trees per acre (N) where MSDR dynamic thinning lines begin and end on the logarithmic scale were used as response variables and predicted as a function of planting density. Predictions of MSDR dynamic thinning line beginning and ending D and N are used in an alternative MSDR dynamic thinning line slope estimation method. These models show that the maximum value of Reineke's Stand Density Index (SDI) varies relative to planting density. By relating planting density specific Zone of Imminent Competition Mortality boundaries to a MSDR species boundary line, self-thinning was found not to begin at a constant relative SDI. Thus, planting density specific Density Management Diagrams (DMD) showed that self-thinning began at 40 to 72% for planting densities of 605 and 2722 seedlings per acre, respectively. / Ph. D.

The response of loblolly pine (Pinus taeda L.) to fertilization and weed control at stand establishment, using various formulations of conventional and controlled release N fertilizers was evaluated in a greenhouse study and at two field trials in the Virginia Piedmont, in 2002 and 2003. The greenhouse study evaluated five fertilizer treatments; 1) check with no fertilizer; 2) granular ammonium nitrate (10-10-10 + micro-nutrients) applied to the soil surface; 3) granular methylene urea (40-0-0) applied to the soil surface; 4) methylene urea (20-10-5) in tablet form applied in the planting hole; and 5) isobutylidenediurea (IBDU) (9-9-4) in tablet form applied in the planting hole. Equal amounts of N and P were applied. Fertilization significantly increased seedling root collar diameter and volume at the end of the first growing season in the greenhouse study. Differences in diameter and volume were still significantly different late in the second growing season, however at the last measurement the differences were no longer significant. An analysis of transformed growth curves for 2003 indicated that the ammonium nitrate treated seedlings had a significantly steeper slope than all other treatments. In field trials, at an old-field site and a cut-over site, the same fertilizer products were tested, except granular diammonium phosphate (18-46-0) was used substituted for the ammonium nitrate. Higher rates of N and P were used in the field trials. Complete weed control increased seedling volume by over 700 % after two growing seasons at the reforested old-field site, however fertilizer effects were not significant. At the reforested cut-over site an interaction between weed control and fertilizer treatments was observed. The MU and DAP granular, and the IBDU tablet treatments each had significantly greater seedling volume than the check and the other tablet controlled release fertilizer. IBDU tablets appear to have high fertilizer efficiency, due to the slow release nature and are safe to place in close proximity to the seedling root system. / Master of Science