Part I. (1) An elementary treatment of the motion of charged particles in a magnetic field is presented. The concept of guiding centre motion is introduced, and is used in outlining the theory of particle drifts. (2 ) The motion of charged particles in the geomagnetic field is discussed, and the concept of adiabatic invariance introduced. (3) Mc Iliwains coordinates for mapping the distribution of charged particles trapped in the geometric field are defined and briefly discussed. (4) A survey of present knowledge of the Van Allen radiation zones is made. Particular attention is given to the distribution, characteristics, and variability of the trapped radiation. (5) The Cape Town magnetic anomaly, the Brazil radiation anomaly and the South Atlantic radiation anomaly are discussed. The electrons entering the South Atlantic radiation anomaly are shown to be those monitored over Iowa by the US satellite Injun I. Part 11. (1) It is shown how the geomagnetic field can, at high altitudes and over relatively short distances, be approximated by the field of a monopole. A new method is developed which enables one to plot the energy absorbed from an electron (which moves in a monopole field in the atmosphere) against altitude, given the initial energy and pitch angle of the electron. Some numerical computations using this method are described, and the results discussed. These results are used, in conjunction with US satellite Injun I for the Iowa region, to estimate the energy inout to the atmosphere in the South Atlantic radiation anomaly. The main approximations and simplifying assumptions made in this treatment, are discussed. (2) Geophysical effects generally recognized to be connected with the precipitation of charged particles are discussed. In the course of this discussion the two main theories of the connection between the radiation zones and the auroras are examined. (3) A preliminary discussion, based on the work summarized in point (1) of detectable geophysical events associated with the precipitation of electrons into the South Atlantic radiation anomaly is given. It is concluded that auroral emission, X-ray bursts, and ionospheric ionization in the E region should be more frequent and pronounced in the South Atlantic radiation anomaly than in any other region of comparable invariant latitudes and that the effects of atmospheric heating by precipitated electrons should be detectable over the anomaly. (4) An assessment is made of the value of the method referred to in point (1). Suggestions for its modification and extension are put forward. It is suggested that if more extensive rocket and satellite data on the low energy component of the electron flux become available, this method can be employed in a rigorous theoretical investigation of the South Atlantic radiation anomaly. Summary, p. 98-100.

Tropical wetlands such as the Pantanal help regulate global biogeochemical cycles, but climate change is modifying these environments. Controls on environmental changes can potentially be assessed from ancient, well-dated lacustrine sedimentary records. An integrated field and laboratory approach was undertaken to study the limnogeology of Lake Uberaba in the northern Pantanal, and test whether the lake has preserved a reliable record of environmental change in its strata. This study was designed to understand how the basin accumulates sediment and to assess its sensitivity to hydroclimatic variability. The data showed that modern Lake Uberaba is a highly dynamic, freshwater fluvial-lacustrine basin. Modern lake floor sediments are largely siliciclastic silts, with limited organic matter content and abundant sponge spicules. This sedimentary composition reflects the lake’s open hydrology and well-mixed water column. Limited data from sediment cores indicates that Lake Uberaba may have formed ~1760 CE, following an abrupt transgression over an oxidized floodplain depositional environment. The stratal contact between lacustrine and floodplain deposits suggests the presence of an erosional unconformity, the timing and duration of which remains unknown. The environmental change favoring lake formation appears to be linked to increased effective precipitation provided by the Intertropical Convergence Zone (ITCZ) in the northern Pantanal.

Diel dissolved oxygen (DO) data were used to characterize seasonal, inter-annual, and longitudinal variation in production and respiration for the James River Estuary. Two computational methods (Bayesian and bookkeeping) were applied to these data to determine whether inferences regarding DO metabolism are sensitive to methodology. Net metabolism was sensitive to methodology as Bayesian results indicated net heterotrophy (production < respiration) while bookkeeping results indicated net autotrophy (production > respiration). Differences in net metabolism among the methods was due to low seasonal variation in respiration using the Bayesian method, whereas bookkeeping results showed a strong correlation between production and respiration. Bayesian results suggest a dependence on allochthonous organic matter (OM) whereas bookkeeping results suggest that metabolism is dependent on autochthonous OM. This study highlights the importance in considering the method used to derive metabolic estimates as it can impact the assessment of trophic status and sources of OM supporting an estuary.

In alpine regions worldwide, climate change is dramatically altering ecosystems, affecting biodiversity across habitats and taxonomic scales. For streams, the associated recession of mountain glaciers and snowfields, paired with altered precipitation regimes, are driving shifts in hydrology, species distributions, and basal resources – often threatening the very existence of some habitats and biota. Globally, alpine streams harbor particularly substantial species and genetic diversity due to significant habitat insularity and environmental heterogeneity: however, anthropogenic warming threatens to homogenize habitats through the reduction of the cryosphere, thereby reducing biodiversity from micro- to macroscopic organisms and genes to communities. Still, alpine stream biodiversity, particularly in North America, is poorly understood, making it difficult to predict future changes without baselines for comparison. For my dissertation, I used genetic tools to assess biodiversity in alpine streams of the central Rocky Mountains in North America. Here, I begin by reviewing the current state of alpine stream biology from an organismal perspective. Next, I provide two perspectives on macroinvertebrate diversity. The first, a population genetic comparison of three highly similar species, is followed by a fine-scale genomic study of one species, Lednia tumana. I follow these largely macroinvertebrate-centric chapters with a modern synthesis of the microbial ecology of mountain glacier ecosystems. Finally, I conclude with a study of microbial diversity that addresses how microbial diversity is shaped by geography, habitat, and hydrological source in North America. Collectively, this research refines existing themes in alpine stream biology by revealing unexpected differences in population genetic patterns among closely related species, the influence of recent deglaciation on population genetic structure and demographic history of a threatened stonefly, and clarification of the environmental drivers shaping microbial diversity.

L'étude de l'industrie lithique des populations qui occupaient la Patagonie et la Terre de Feu nous a permis de discuter des dynamiques culturelles mises en œuvre par les chasseurs terrestres et maritimes, de part et d'autre du détroit de Magellan, à l'Holocène moyen et tardif. L'approche techno-économique développée pour les industries de la grotte Fell, Punta Santa Ana 3, Marazzi 1, Cabo Monmouth 20, Oosin Aike et Cabo San Vicente a été axée sur deux éléments: d'abord, les objets dont les matières, parfois d'origine non locale, constituent des témoignages de transport et d'interaction (obsidiennes); ensuite, les méthodes élaborées de débitage, avec prédétermination (méthode Levallois, Clactonien, laminaire et discoïde), dont les modalités peuvent être liées aux processus complexes de partage et de transmission des connaissances. Une discontinuité dans la distribution archéologique de l'obsidienne verte est observée entre 4500 et 2500 ans BP et nous permet d'envisager l'hypothèse d'une migration des peuples de Patagonie du nord. La comparaison des distributions spatiales et chronologiques des méthodes de débitage Levallois montrent des interactions et contacts entre groupes, aussi bien que des conceptions partagées, transmises et généralisées dans la Fuego-Patagonia. Ces observations sont en lien direct avec les premiers nomades de la mer, dont le rôle clé, à partir de 7000 BP, supprime la notion de barrière biogéographique relative pour le détroit de Magellan. Ainsi, on remarque l'existence d'un fond culturel commun. Les variations correspondraient plutôt à des évolutions mesurables en termes d'intensités d'interactions et d'adaptations à des systèmes écologiques. / The study of the lithic industries of the peoples that inhabited Patagonia and Tierra del Fuego bas allowed the discussion of cultural dynamics, displayed by terrestrial and marine hunter-gatherers, at one or the other side of Magellan strait, during the Middle and Late Holocene. The techno-economic approach used to study the materials from Fell cave, Punta Santa Ana 3, Marazzi 1, Cabo Monmouth 20, Oosin Aike and Cabo San Vicente, was organized following two axis: first, objects of exotic raw materials that establish evidence of transport and interaction (obsidians); second, elaborated core reduction methods and débitage with predetennination (Levallois, Clactonian, blade, and discoid reduction), were the action modes can be related to complex processes of knowledge sharing and transmission. The observed discontinuity of green obsidian archaeological distribution between 4500 and 2500 years BP bas permitted considering the hypothesis of a north Patagonian immigration. The comparison of spatial and chronological distributions of Levallois core reduction methods, show interactions and contact between human groups. Likewise they display the existence of shared conceptions, as in an integrated, transmitted and generalized knowledge corpus for Fuego-Patagonia. These results have direct rapport with the key role of the first marine nomads since c. 7000 BP and the removal of the biogeographic barrier notion relative to the Magellan strait. Thus, worthy ofremark is the existence of a common cultural background and that differences in distribution would correspond to measurable changes regarding interaction intensity and adaptation to ecologic systems.

Cette thèse présente une quantification du CO2 dans le manteau lithosphérique Pannonien par l'étude de xénolites de péridotites remontées par le volcanisme alcalin de la Bakony, au nord du lac Balaton. Les études texturales et compositionnelles de xénolites ont mis en évidence un métasomatisme polyphasé du manteau lithosphérique. Le dernier épisode métasomatique résulte de la percolation de liquides silicatés de type adakite, issus de la fusion d'une lithosphère subductée. Lors du métasomatisme, ces fluides primaires ont réagi avec des amphiboles métasomatiques, pour former les magmas parents de la suite calco-alcaline observée à la surface du bassin. Le bilan CO2 de ces magmas et du manteau sous Pannonien a été contraint par une approche innovante couplant : (1) de la microtomographie au rayonnement synchrotron, (2) des mesures NanoSIMS, Raman et microthermométriques, et (3) des modélisations thermodynamiques. Les concentrations en CO2 des fluides primaires ont été quantifiées entre 9.0 et 25.4 wt. %, validant ainsi une source riche en fluides. Des teneurs de ~2000 ppm de CO2 ont été estimées comme représentatives du manteau lithosphérique Pannonien. Cet important réservoir lithosphérique de CO2 est probablement drainé par le système tectonique décrochant de la Mid Hungarian Zone, entrainant l'omniprésence de CO2 mantellique dans le Bassin pannonien. / This work presents a quantitative investigation of the CO2 in the lithospheric mantle by the study of peridotite xenoliths brought up to the surface by alkaline volcanism in the Pannonian Basin (Central Europe). Textural and geochemical studies of mantle xenoliths highlight a polyphased metasomatism of the lithospheric mantle. The last metasomatic event is related to the percolation of silicate melts of adakite-like compositions, originated from slab melting. During metasomatism, primary fluids reacted with metasomatic amphiboles to form parental melts of the calc-alkaline series observed at the surface of the Pannonian Basin. The CO2 budget of adakite-like magmas and of the mantle below the Pannonian Basin was then constrained by an innovative approach on mantle xenoliths using (1) synchrotron X-ray microtomography, (2) NanoSIMS, Raman spectroscopy and microthermometry, and (3) thermodynamic models. The CO2 concentrations in adakite-like melts were estimated between 9.0 and 25.4 wt. %, in agreement with a fluid-rich source. CO2 concentrations of ~2000 ppm were determined as representative of the Pannonian lithospheric mantle. This significant CO2 lithospheric reservoir is probably tapped by the major shear zone of the Mid Hungarian Zone, resulting in the omnipresence of mantle CO2 in the Pannonian basin.

The equilibrium theory of island biogeography provides a useful model for understanding patterns of species richness on island systems and analogous fragmented terrestrial habitats. However, like all models, it is limited in its ability to explain island species richness patterns when nutrients move across ecosystem boundaries. Recently, enhancements to the theory have been proposed, including the subsidized island biogeography hypothesis. This hypothesis suggests that nutrient subsidies from the marine environment may impact the productivity and diversity of small islands. Sea wrack (dead, shore-cast seaweed) is a recognized vector of marine-nutrient subsidies to islands in regions of low in situ productivity, but little is known about the mechanisms surrounding sea wrack accumulation in productive, temperate environments. In this research I explore the spatial and temporal distribution of sea wrack on islands along British Columbia’s temperate Central Coast. Through an observational study I investigate three broad factors that could affect sea wrack deposition: climatic patterns, physical characteristics of shorelines, and the amount of nearby donor habitat. I surveyed sea wrack biomass and species composition, as well as the biogeographical characteristics of shorelines across 455 sites on 101 islands. I returned to a subset of sites on a bi-monthly basis to document temporal changes in wrack biomass and species composition. My results demonstrate that sea wrack accumulations were present at sites that were not composed of rock substrate, and that had wide, wave protected shorelines and high amounts of nearby donor ecosystem habitat. Additionally, sea wrack biomass and species composition was ubiquitous throughout all seasons. These results suggest that sea wrack can be considered a press subsidy as it is a consistent vector of nutrients to beaches along the Central Coast. Ecological research on macrophytes, macroalgae and sea wrack often requires the conversion of wet biomass to dry, to create consistency across investigations. This is a laborious process. Here, I present the results of wet-dry calibrations for 12 common macrophyte and macroalgae species collected from the Northeast Pacific Ocean. Future investigators can use the correction factors derived from these results for estimating dry biomass, reducing the need to conduct wet-dry calibrations for each new macrophyte, macroalgae, or sea wrack study. / Graduate

The regular occurrence of hurricane-associated winds has been an important factor in shaping the structure and composition of the forest ecosystems of the Florida Everglades. Forest communities in the Everglades are adapted to hurricane disturbances, but increased frequency and/or intensity of hurricanes may lead to decline or even collapse of these communities. The overall objective of this project is to understand the patterns, pace, and mechanisms of the recovery process to Hurricane Andrew damage in four Everglade forest communities: pinelands, hardwood hammocks, bayhead tree islands, and cypress domes. This study combines long- and short-term field measurements and experimental garden studies to determine how the four woody plant community types recover from hurricane impacts. Most of the community types were adversely affected by storm damage in the short-term (3 years post-hurricane) through altered growth rates and canopy defoliation, however these effects were relatively short lived and were not visible in shifts in species composition after the long-term (20 year) recovery period. Only in the most diverse communities over the long-term there was a delayed mortality in damaged individuals that drove a diversity loss. This loss was not present over the short-term recovery time period. Using individual damage extent and short-term recovery growth rates, I developed a simplified model that accurately predicted surviving individual stem size over long-term recovery periods of Taxodium distichum within cypress domes and select hardwood hammock species. The shadehouse experiments demonstrated the importance of nutrient availability to growth of seedlings of canopy dominants. Recruits of these species responded to changing environmental conditions associated with storm impact through a variety of strategies in accordance with their adaptive traits. Synergistically, the combined parts of this dissertation demonstrate directional community and species-specific shifts that vary over time scales. Storm impacts have the potential to alter community composition and diversity within impacted systems, and in particular the Everglades ecosystem.

Hydroscape structure can play a critical role in animal behavior, abundance, and community structure dynamics. Hydroscape configuration can be dynamic and can change quickly in ephemeral systems. However, ephemeral freshwater wetlands are among the most impacted systems in the world and restoration efforts often rely on incomplete information when establishing management objectives. Further understanding how alterations in hydroscape structure in dynamic systems affect animals is critical for conservation and management success. To determine impacts that changing hydroscape conditions can have on consumers in freshwater wetlands, I examined the effects of a large-scale physical model on fish behavior, abundance, and community structure. The physical model incorporated the restoration of sheetflow, canal-fill treatments, and the removal of a decades-old levee that divided two water management areas in the central Everglades. Small fishes modified directional movement behaviors and speed of movement before and after alterations took place, though behavioral responses varied widely by species. Density and community structure of small fishes did change as a function of canal-fill and levee removal treatments. Behaviors of large fishes were also affected by hydroscape alterations, as well as hydroscape configuration beyond the limited footprint of the physical model. Large fish abundance was altered by hydroscape alteration, particularly among certain species. Composition of the large fish community changed before and after hydroscape alteration, though magnitude of responses were site-specific. Effects of hydroscape structure proximity on trophic dynamics were examined using exclosure cages that excluded large predators but allowed access for small consumers. Exclosures were stratified according to proximity to a deep-water canal. Predator avoidance behaviors in small consumers were limited but present. Differences in behavior between sites may also be caused by differences in structure across sites and limited differences in nutrient quality. Behavioral, population, and community responses to hydroscape alteration can be valuable metrics to assess the success of hydroscape restoration. While results can vary across individuals, species, and sampling sites, effects can still be detected even at the scale of the hydroscape. My research has detailed the potential effects of restoration plans across the greater Everglades and can be extended to other ephemeral wetland restoration programs.

Degraded pollinator habitat may have far-reaching consequences for recovery of imperiled flowering plant populations. Studies indicate that broad-spectrum insecticides used in mosquito abatement are detrimental to non-target invertebrates, including pollinators. A decline in efficient pollinators can reduce plant fitness by decreasing reproductive output and constraining genetic diversity, a challenge for rare plants. In 2015, I monitored flower visitation and fruit set of three imperiled plant species throughout protected areas on three islands in The Lower Florida Keys. These islands consist of conservation land fragmented by intermittently dispersed residential neighborhoods that seasonally spray insecticides for mosquito control. Flowers open at treatment sites had decreased flower visitor activity following insecticide applications, but only species that require invertebrate agents for pollen transfer had significantly reduced fruit set. Implications of mosquito insecticides near conservation lands may pose immediate threats to invertebrate pollinators and flowering plants that require pollinators for reproduction, although long-term threats to genetic diversity are unknown for automatic self-pollinating species.