<p>The aim of this work is to analyze if a change in the benthic community can be detected two years after a restoration of a small stream. The samples were taken in a small stream at Tinnerö Eklandskap just south of Linköping. In addition to the restored area, two reference sites upstream and downstream of the restored area were sampled to compare to the restored site. The method used for sampling of benthic fauna in the stream was kick sampling. ASPT, Berger-Parker and Renkonen-indices were used to find out if there was any difference between the reference areas and the restored area. In addition to indices, rank-abundance curves and species lists were made to see if there was any trend difference between the different areas. The only index that showed a difference between the different areas was Berger-Parker diversity index. The reason why there were no greater differences between the areas may be due to the fact that two years is too short to allow time for the benthos to re-colonize the restored area.</p>

The aim of this work is to analyze if a change in the benthic community can be detected two years after a restoration of a small stream. The samples were taken in a small stream at Tinnerö Eklandskap just south of Linköping. In addition to the restored area, two reference sites upstream and downstream of the restored area were sampled to compare to the restored site. The method used for sampling of benthic fauna in the stream was kick sampling. ASPT, Berger-Parker and Renkonen-indices were used to find out if there was any difference between the reference areas and the restored area. In addition to indices, rank-abundance curves and species lists were made to see if there was any trend difference between the different areas. The only index that showed a difference between the different areas was Berger-Parker diversity index. The reason why there were no greater differences between the areas may be due to the fact that two years is too short to allow time for the benthos to re-colonize the restored area.

Some of the major problems we face today are human induced changes to the nitrogen (N), phosphorus (P) and carbon (C) cycles. Predicted increases in rainfall and temperature due to climate change, may also increase dissolved organic matter (DOM) inflows to freshwater ecosystems in the boreal zone. N, P, C and light, are essential resources that most often limit phytoplankton (PPr) and bacterial production (BP) in the pelagic zone of lakes. PPr and BP not only constitute the total basal C resource for the pelagic aquatic food web, but also influence ecosystem function and biogeochemical cycles. In this thesis I studied how N, P, C and light affect the relative and absolute rates of PPr and BP, along a wide latitudinal and trophic gradient using published data, and in two in situ mesocosm experiments in a clear water oligotrophic lake. In the experiments I manipulated bottom-up drivers of production and top-down predation to examine how these factors interact to affect pelagic food web structure and function. The most important predictors of PPr globally (Paper I) were latitude, TN, and lake shape. Latitude alone explained the most variation in areal (50%) and volumetric (40%) PPr. In terms of nutrients PPr was primarily N-limited and BP was P-limited. Therefore bacteria and phytoplankton were not directly competing for nutrients. BP:PPr was mostly driven by PPr, therefore light, N, temperature and other factors affecting PPr controlled this ratio. PPr was positively correlated with temperature, but not BP, consequently, higher temperatures may reduce BP:PPr and hence the amount of energy mobilised through the microbial food web on a global scale. In papers II and III interaction effects were found between C-additions and top-down predation by young-of-the-year (YOY) perch. Selective predation by fish on copepods influenced the fate of labile C-addition, as rotifer biomass increased with C-addition, but only when fish were absent. Interaction effects between these top-down and bottom-up drivers were evident in middle of the food web, which is seldom examined in this type of study. Although the energy pathway from bacteria to higher consumers is generally longer than from phytoplankton to higher trophic levels, increased BP still stimulated the biomass of rotifers, calanoid copepods and YOY fish. However, this appeared to be mediated by intermediate bacterial grazers such as flagellates and ciliates. Light was an important driver of crustacean zooplankton biomass (paper IV), but the light:nutrient hypothesis was inadequate to predict the mechanisms behind the decrease in zooplankton biomass at low light. Instead, it appeared that reduced edibility of the phytoplankton community under low light conditions and reduced BP most strongly affected zooplankton biomass. Thus, the LNH may not apply in oligotrophic lakes where PPr is primarily N-limited, Daphnia is rare or absent and mixotrophic phytoplankton are abundant. N, P, C and light manipulations have very different effects on different parts of the pelagic food web. They influence the relative rates of PPr and BP, affect phytoplankton community composition, alter the biomass of higher trophic levels and change pathways of energy transfer through the pelagic food web. This thesis adds valuable information as to how major changes in these resources will affect food web structure and function under different environmental conditions and future climate scenarios. / Lake ecosystem response to environmental change

The minimum ecologically viable population size (MEVP) of a species describes the minimum size at which the species itself or another species in the same ecosystem goes extinct as a result of the loss of inter-specific interactions. The MEVP shows a good potential for use as a tool for exploring the mechanisms behind species extinctions, but presently only a small amount of research has been done that takes advantage of this. In this study the MEVP is used to investigate what properties of species can be used as good indicators of ecological importance. 100 large computer generated food webs were created with an assembly model and the reduction in density that was necessary to induce an extinction event in the web was subsequently determined for each species within the webs. This change in density was then correlated with 28 different properties, measured for each species. The results show that properties that measure how well connected a species is, as well as measures of the species role as a prey item in the web are the ones with the greatest potential to find species with high MEVP. Further, the results put emphasis on the importance of regarding the web as a whole when working with species extinctions, while also highlighting the usefulness of the MEVP concept.

The colour choice of spiders and the effects of competition on this phenomenon were examined in the laboratory using the sit-and-wait predator goldenrod crab spider (Misumena vatia, (Thomisidae)). This species can  be both white and yellow (could also change colour)  and it was here examined whether individuals preferred to sit on flowers which has the same colouration as their bodies. I found that M.vatia showed preference for flowers that matched their own body colour, but when competition was introduced this preference disappear. It seems, however, that position is a factor that can influences the substrate decisions of M. vatia, where the edge of the flower is preferred.

This research aims to investigate the effects that dredging may have on benthic fauna. The dredging took place in the stream Ireån on northern Gotland during the early summer 2010. My area of research was taking place in a scenic area downstream in Ireån close to the Baltic Sea. I chose this area due to the dredging that started just upstream of this area and continued further upstream. Thus it is not the direct impact of the dredging that is the focus for this paper but the side-effect the dredging may have on the area downstream. In this research I used the M42-method and BQI- index in order to evaluate the results. The effects on the dredging were weaker than expected, with only minor changes occurring within the benthic fauna (BQI-value before dredging: 3.35, BQI-value after dredging: 3.51).

<p>Ecological models have problems showing the positive relationship between diversity and stability found in nature. Theory states that complex food webs have high extinction risks and low stability. However, persistent food webs found in nature are large and complex containing many interconnections between species. There are many possible mechanisms enabling persistent food webs such as; complex interaction patterns, asynchronous fluctuations of species densities, environmental fluctuations and spatial distribution. These factors have not been used in classical models. In this study, coloured environmental 1/f noise and dispersal between subpopulations were incorporated into a diamond shaped food web based on a model by Vasseur and Fox 2007. Contradictions between theoretical and empirical results regarding food webs can be resolved by detailed analyses of models, withholding stabilizing mechanisms. Weak environmental 1/f noise generated an increased coefficient of stability but the stabilizing effect of noise can be questioned because of a decreased mean food web biomass and reduced stabilizing effect when reddened. However, detailed studies of the food web revealed that noise can redistribute density proportions between species, evading lowest species density and thereby increase food web resistance to demographic stochasticity and catastrophes. Noise induced density proportion shifts imply that large population sizes are no insurance towards future increase in environmental variance. Synchrony of species environmental responses and dispersal between subpopulations can both have major influences on stability and extinction risk of smaller food webs indicating that spatial structure could be one of the dominating factors stabilizing complex food webs found in nature.</p>

Shallow sheltered Baltic Sea bays are ecologically important habitats that harbour a unique vegetation community and constitute vital reproduction areas for many coastal fish species. Knowledge about the invertebrate community in these bays is, however, limited. This thesis examines the macroinvertebrate community in shallow sheltered Baltic Sea bays and how it is affected by: (1) the natural morphometric isolation of bays from the sea due to post-glacial land uplift; and (2) differences in vegetation types. The invertebrate biomass and number of taxa was found to decrease with increased bay isolation. The taxon composition changed from dominance by bivalves and gastropods in open bays to a community composed of a larger proportion of insects in isolated bays. Stable isotope analysis indicated epiphytes and periphyton as the major energy resources for most of the examined consumers, but the relative importance of these in relation to larger plants decreased for some consumers with increased bay isolation. A comparison of invertebrate abundance between plants revealed a close relationship with morphological complexity of the plants. More complexly structured plants had higher invertebrate abundance than plants with simpler morphology. The results suggest that management of these coastal habitats should be dynamic and take into consideration the natural change in invertebrate community resulting from the slow bay isolation process. In addition, the results imply that changes in the aquatic vegetation due to anthropogenic influences could induce changes in the invertebrate community as the plant habitat structure is altered. A changed invertebrate community may in turn affect higher trophic levels since invertebrates are important food for many fish and waterfowl species. / At the time of the doctoral defense, the following papers were unpublished and had a status as follows: Paper 2: Submitted. Paper 4: In press.

Ecological models have problems showing the positive relationship between diversity and stability found in nature. Theory states that complex food webs have high extinction risks and low stability. However, persistent food webs found in nature are large and complex containing many interconnections between species. There are many possible mechanisms enabling persistent food webs such as; complex interaction patterns, asynchronous fluctuations of species densities, environmental fluctuations and spatial distribution. These factors have not been used in classical models. In this study, coloured environmental 1/f noise and dispersal between subpopulations were incorporated into a diamond shaped food web based on a model by Vasseur and Fox 2007. Contradictions between theoretical and empirical results regarding food webs can be resolved by detailed analyses of models, withholding stabilizing mechanisms. Weak environmental 1/f noise generated an increased coefficient of stability but the stabilizing effect of noise can be questioned because of a decreased mean food web biomass and reduced stabilizing effect when reddened. However, detailed studies of the food web revealed that noise can redistribute density proportions between species, evading lowest species density and thereby increase food web resistance to demographic stochasticity and catastrophes. Noise induced density proportion shifts imply that large population sizes are no insurance towards future increase in environmental variance. Synchrony of species environmental responses and dispersal between subpopulations can both have major influences on stability and extinction risk of smaller food webs indicating that spatial structure could be one of the dominating factors stabilizing complex food webs found in nature.

The climate affects animal populations through several processes. These processes includereproduction, phenology and the success of hunting. By influencing the phenology of species theclimate also affects the way species interact. If a climate-induced phenology change promotes abasal species to bud earlier in the spring for example, this will affect the links to other species inthe food web due to a time lag in the tropic levels above. This dependence, that a predator has tobeing synchronous to its prey, is called the match/mismatch hypothesis (MMH). Studies haveindicated that species higher up in the food web adapt slower than species lower in the food webwith shorter generation times (which creates a temporal mismatch).A climate-induced phenology change in basal species could therefore be expected to affect thedensities and extinction rates of species higher up in the food web.In this study a declining conversion efficiency for predators was used to model the effects ofclimate change on triangular food webs with three trophic levels. This was done by using ageneralized Lotka-Volterra model. The results indicate that, in a food web with three trophiclevels, the densities of herbivores and carnivores drops in response to an increased change ofclimate. The extinction rates of carnivores also increase rapidly in response to an increasedclimate change. / Klimatet påverkar djurpopulationer genom ett flertal processer. Dessa processer inkluderarreproduktion, fenologi och jaktframgång. Genom att påverka arters fenologi påverkar klimatet ävenhur arter interagerar med varandra. Om en klimatinducerad fenologiförändring stimulerar en basalart att knoppa tidigare på våren kommer det att påverka länkar till andra arter i näringsväven pågrund av en tidsförskjutning till andra trofinivåer. Detta beroende för ett rovdjur att varasynkroniserad med sitt byte kallas för match/mismatch-hypotesen (MMH). Studier har indikerat attarter högre upp i näringsväven anpassar sig långsammare än arter lägre ned i näringsväven medkortare generationstid. En klimatinducerad fenologiförändring i basala arter kan därför förväntaspåverka densiteten och utdöendehastigheten för arter högre upp i näringsväven. Studien använderen avtagande rekryteringsförmåga för att modellera klimatförändringars effekt på trianguläranäringsvävar med tre trofinivåer. För att göra detta har en generell Lotka-Volterra modell används.Från resultaten av den här studien dras också slutsatsen att, i en näringsväv med tre trofinivåer,densiteten av herbivorer och karnivorer avtar som svar på en ökad klimatförändring.Utdöendehastigheten för rovdjur ökar också snabbt som svar på en ökande klimatförändring.