A grazing trial investigating the effect of season of grazing and stocking rate initiated at the Nortier Experimental Farm in 1988 provided an opportunity to assess the response of the veld to both grazing and environmental influences in an arid environment. The trial allowed an assessment of the relative influence of internal (equilibrium) and external (non-equilibrium) forces on the dynamics of an arid rangeland. This study involved the analysis of a nine year data set stretching from 1988 to 1996 and served to provide evidence supporting the existence of an equilibrium/non-equilibrium continuum in rangeland dynamics. The most significant implication of this result is that rangeland systems should not be classified as either equilibrial or non-equilibrial, but rather according to a continuum extending between equilibrium and non-equilibrium poles. The exact position of any system on this continuum is a function of the relative influence of internal and external forces on its species dynamics. The dynamics of the veld at the Nortier Experimental Farm showed significant response to both grazing and environmental variables suggesting conformity to both equilibrial and non-equilibrial paradigms. Both ordination and analysis of variance highlighted the importance of rainfall particularly in the fluctuations of the predominant grass species, Ehrharta calycina, which increased in abundance with rainfall. Partial ordination enabled the assessment of species variation following the removal of variation associated with rainfalL Partial ordinations revealed the gradual, directional movement of samples through multivariate space in response to grazing treatments. Individual plant species were also shown to be responding to grazing, the extent of which was influenced by season of grazing and stocking rate. Both the partial ordinations and the ANOVA showed Melothria sp., Tetragonia fruiticosa and Hermannia scordifolia as increasing and Ruschia caroli as decreasing in absolute abundance in response to grazing. Season of grazing was shown to significantly influence the abundance of H. scordifolia over time. The 'shrublherb complex', which constitutes the 'key resource' at the Nortier Experimental Farm displayed an increase in absolute abundance over the duration of the trial. This increase in absolute abundance was accompanied by an increase in the relative abundance of the palatable component of this resource. The application of medium to heavy stocking rates during spring, summer and autumn and low stocking rates during winter resulted in elevated absolute abundances of palatable plants. Furthermore, low stocking rates, when averaged across all season of grazing treatments, resulted in a significantly higher absolute abundance of unpalatable plants. These findings provide the basis for the development of management principles for the Strandveld Vegetation Type. The application of medium to heavy stocking rates within a rotational grazing system, as recommended by the literature dealing with grazing systems in the Karoo, is supported by the results of the Nortier grazing trial. Medium to heavy stocking rates should be applied during spring, summer and autumn and low stocking rates during the winter months. Furthermore, it is recommended that rests of between 12 and 14 months should be afforded to portions of the veld periodically due to the variability in growth, flowering and fruiting times ofdifferent plants in the Karoo. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2005.

Long-term trends in rangeland sward dynamics (species composition, structure, productivity) were examined on three trials established between 1989 and 1996 at Kokstad Research Station in the Highland Sourveld, while animal performance (average daily gain and gain per hectare) was examined on two of the trials. The region enjoys moderate rainfall of 782mm per annum, with hilly topography, and soil depths ranging from >1m to <20cm. The first trial was labelled the simulation trial, as it simulated a four-paddock rotational grazing system, in which animals spent two weeks in each of three paddocks while the fourth was rested for the entire season. The rested paddock was rotated each year. The trial tested two stocking rates (0.5 and 1.0 AU.ha-1) at five ratios of cattle to sheep, ranging from cattle only to sheep only. The trial was unreplicated, and was established in 1989 on flat topography with deep soils. The second trial (labelled the flat two-paddock trial) was established in 1992 adjacent to the simulation trial. The trial examined two stocking rates of sheep weaners (0.5 and 1.0 AU.ha-1 seasonally) in a continuous grazing two-paddock system, in which one paddock of each treatment was burned and grazed continuously while the second paddock was rested, to be burned and grazed in the following season. The trial was replicated twice. The third trial (labelled the steep two-paddock trial) mimicked the grazing system of the flat trial, but was located on a steep (c. 20%) West-facing slope with shallow soils. The trial incorporated two additional treatments: an intermediate stocking rate of 0.7 AU.ha-1 and an ungrazed treatment. Species composition of the sward was recorded biennially on all trials using the nearest plant-point technique with between 200 and 800 points per paddock. Sward standing crop was measured in the rested seasons of the simulation trial and at the beginning, middle and end of each season in one paddock of each two-paddock treatment of the two-paddock trials. In the two-paddock trials, sward standing crop was measured within and outside permanently placed exclosure cages. Animals were weighed fortnightly. The response of species to grazing pressure or animal type was mediated by soil depth and slope, as well as the grazing system. Tristachya leucothrix declined on all grazed treatments. The ungrazed treatments remained relatively stable over ten years. On the low stocking rate treatments of the steep trial, unpalatable species increased, but so did Themeda triandra. The heavily grazed treatment of the steep trial was surprisingly stable, with little significant change in relative abundance of key species other than an increase in the unpalatable Alloteropsis semialata and decline in T. leucothrix. The medium stocking rate treatment on the steep trial showed significant shifts in relative abundance of key species, with declines in T. triandra and T. leucothrix and increases in A. semialata and the unpalatable wiregrass D. filifolius. These trends were not repeated on the flat trial, however, with T. triandra and A. semialata increasing and all other key species declining or remaining stable. On the simulation trial, species responded largely unpredictably with species abundances often fluctuating considerably over time. Microchloa caffra and A. semialata increased substantially in both the low and high stocking rate sheep-only treatments, with a concurrent decline in T. triandra in the high stocking rate but not the low. Changes in composition over time, as measured by Euclidean distance, showed that shallow soils, high stocking rates and a high proportion of sheep caused greater shifts in species composition over time than deep soils, low stocking rates or more cattle. Three treatments, the sheep-only treatments on the simulation trial and the high stocking rate on the steep trial, showed an initial rapid shift in composition over about 6 years, before stabilising in subsequent seasons. The flat trial showed no substantial shift in composition over time. This general pattern of change was confirmed by Non-Metric Multidimensional Scaling. On the simulation trial, total standing crop was influenced by stocking rate and by the proportion of sheep in most seasons. On the two-paddock trials, increasing stocking rate significantly reduced sward vigour, and vigour declined over time. Stocking rate reduced total standing crop on both trials at the end of the 2004/05 seasons and the crop of unpalatable species on the steep trial. Total palatable plants were unaffected by stocking rate on both trials. The classic Jones-Sandland model of animal performance as influenced solely by stocking rate was not supported. Sheep performance was influenced by stocking rate and the interaction of stocking rate and seasonal rainfall. There was no difference in average daily gain between treatments over time, and hence cumulative animal production per hectare increased with increasing stocking rate. Animal performance was possibly influenced by many factors beyond the scope of this study, including the effect of predator attacks on surviving animals, and resource availability such as shade and shelter and high-production patches in some paddocks and not others. Scale effects on ecology are being increasingly investigated and a meta-analysis of this type shows that, even in one research farm, slight differences in management and environment can have significant effects on plant and animal responses to grazing. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2010.

A series of trials and investigations were implemented to address concerns surrounding the dynamics of the fire-climax wooded/edaphic grasslands within the iSimangaliso Wetland Park, northern KwaZulu-Natal, South Africa. The research problem surrounded inadequate historical evaluations of changes in vegetation structure, grasslands progressing to a woody dominated composition, and increases in Helichrysum kraussii (Curry bush). These were addressed as follows: Firstly, the recovery of vegetation in response to different periods of fire exclusion in different communities along a topographical gradient of a coastal dune area, was assessed over a two year period. Secondly, the regeneration after wildfire of the persistent, stress tolerant shrub H. kraussii, was studied on different catenal positions with differing fire exclusion periods and with and without defoliation of surrounding plant biomass in the coastal edaphic grasslands north of Manzengwenya, South Africa. Thirdly, aerial photography from 1937, 1975, and 2000 was georectified, digitised and analysed using a Geographic Information System to examine broad vegetation changes in response to different management regimes for a site on the Eastern Shores of Lake St Lucia and a site within the Tewate Wilderness Area. In the absence of fire, the coastal edaphic grasslands progressed to a closed canopied scrub forest within six years. An increase in fire exclusion period resulted in a decrease in species abundance, an increase in woody height, and a decrease in plant density. Richness increased initially but declined marginally with increased fire exclusion period. Higher lying east and west facing sites had a better veld condition index compared with bottom sites and had an increased response (vigour) to defoliation but were far more likely to succeed through to woody scrub forest. Woody plant biomass vigour was greater for west facing sites. Ordination of species composition across sites in response to fire exclusion and catenal position revealed greater similarities within exclusion periods than between. Bottom sites were more similar with similarity decreasing for east and west facing sites. Fire exclusion resulted in an initial increase in woody species and a subsequent increase in herbaceous species. iii Growth response of H. kraussii was unaffected by catenal position and fire exclusion period, whereas defoliation of surrounding grass tended to increase in size (P<0.05). Density and height for this species however increased with increasing fire exclusion. An increase in soil moisture negatively affected H. kraussii growth indicating susceptibility to high water tables. The number of other woody species establishing beneath H. kraussii may be due to changes in the transmission of light through the canopy where an increase in canopy diameter resulted in an increase of photosynthetically active radiation at the soil surface. The effects of fire on landscape change were investigated for the Eastern Shores and Tewate Wilderness Area, iSimangaliso Wetland Park, South Africa using aerial photography. Changes to historical disturbance regimes largely through active exclusion of fire resulted in the majority of the higher lying coastal grasslands changing to savanna scrub or closed canopied forest within 63 years on the Eastern Shores. The degree of fragmentation of these grasslands was greatly reduced within the Tewate Wilderness Area where disturbance regimes included greater frequencies of fire. Hygrophilous grasslands remained largely unaffected by woody encroachment but did not preclude woody species establishment indicating possible susceptibility during long drier periods. Frequent fires result in the maintained distribution of the higher grasslands. This vegetation type is a system which becomes resilient in response to fire, whereas in the absence of fire readily progresses to Dune Forest. The coastal grasslands above the high water table are therefore highly unstable and transformed easily in the absence of regular disturbance. It would appear that a threshold of approximately six years exists, after which substantial management intervention may be required to reverse the succession back to grassland. The growth of H. kraussii was unaffected by fire and remained persistent irrespective of fire exclusion period. An ability to attain size (height and canopy diameter) was limited with increased soil moisture but density was reduced through regular burning. Frequent fires are necessary to reduce density of H. kraussii and reduce the competitive advantage gained with age. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2007.

Water provision is an important tool in the management of savanna ecosystems. Artificial water sources are a potential focus for degradation (biodiversity and loss of ecosystem resilience at a range of spatio-temporal scales), because they alter plant-animal interactions and soil function and stability, through the creation of piospheres. This study was undertaken as part of a drive by the Kruger National Park (KNP) to enable managers to integrate artificial waterhole management (e.g. waterhole closure or rotation) when setting goals for heterogeneity and biodiversity conservation in the park. The over-arching goal was to quantify the relationship between water provision and different attributes of heterogeneity, as part of a broader initiative to place water provision and piospheres within an ecosystem threshold framework. Herbivore utilisation gradients (piospheres) around artificial waterholes in the KNP, described in 1990, were resurveyed in 2006-2007, against a backdrop of waterhole closure in the KNP, to contribute to an understanding of the factors governing recovery and resilience in grazing systems. The responses of the plant community and soil parameters to a relaxation of herbivore utilisation pressure around closed waterholes (recovering piospheres) were examined, as were changes in the same parameters at sites that have remained open (active piospheres). These ecosystem properties were considered in relation to structural and functional ecosystem thresholds, and the piospheres surveyed incorporated a range of rainfall and edaphic gradients in the KNP. Herbaceous basal cover and soil infiltration capacity both increased significantly between 1990 and 2006/7, regardless of waterhole status. This was linked to higher rainfall in 2006/7, compared to 1990. The only vegetation variables to respond consistently to distance from water were the remote-sensed Normalized Difference Vegetation Index (NDVI) and herbaceous species composition. NDVI increased with distance from water, and annual grasses and forbs were most abundant close to water. Perennial, disturbance-sensitive climax species increased in abundance further from water. Soil analyses (N, P, pH, organic matter, and texture) and field measurements (infiltration, compaction) revealed no systematic piosphere patterns. Waterhole closure did not result in soil or vegetation recovery, but piosphere intermittency and the increases of basal cover and infiltration rate indicated that ecosystem resilience has not been compromised vii by long-term artificial water provision in the KNP. This study has shown that the traditional piosphere model is of limited use in sub-humid savanna ecosystems like the KNP during above-average rainfall periods. / Thesis (Ph.D.)-University of KwaZulu-Natal, Pietermaritzburg, 2010.

Perennial ryegrass is an intensive, temperate pasture grass that responds well to applied fertiliser. The purpose of this project was to study the effects of fertiliser on the productivity and quality of perennial ryegrass in KwaZulu-Natal and how this impacts on animal intake. It was hypothesised that over-application of fertiliser to a perennial ryegrass pasture would negatively affect the quality of the herbage for grazing by dairy cattle and that intake would be affected. Thus the project aimed to assess the effects of applied fertiliser on yield, quality and intake of an established perennial ryegrass pasture. The trial consisted of a set of six separate experiments. Each experiment focused on the interaction between two of the major nutrient elements nitrogen (N), phosphorus (P), potassium (K) and sulphur (S). The experiments (NxP, NxK, NxS, PxK, PxS and KxS) were managed separately to avoid possible transfer of nutrients during grazing, which would result in the contamination of treatments. Each factor had three levels (low, medium and high), giving a total of nine treatments per experiment. Each of the experiments was replicated three times in a randomised block design. Increased fertiliser N application rates increased perennial ryegrass yield with a pattern of diminishing return, where split applications above 40 kg N ha-1 produced smaller increases in yield when compared with the response at lower applications of N. Applied P, K and S did not affect yield, suggesting that even the lowest application levels were sufficient to not limit production. Nitrogen application affected apparent intake, but it is suggested that this is due to the yield effect rather than a direct effect of N on apparent intake. The application of P, K and S did not affect apparent intake. Results from this study showed that the quality of perennial ryegrass herbage, especially in terms of feed value to dairy cows, can be significantly affected by applied fertiliser. The extent of the response was affected by sampling date (i.e. time of year) and this must be taken into account when planning a fertiliser management strategy. This is particularly so with respect to N fertiliser recommendations. Crude protein (CP) content of herbage increased with increasing levels of applied N and the extent of the response was influenced by season. P, K and S did not affect CP concentration in herbage, except in the PxK experiment where increased levels of K lowered herbage CP. Applied N considerably increased the concentration of non-protein nitrogen (NPN) in perennial ryegrass herbage. P and S did not affect NPN levels, whereas applied K decreased NPN content in the iv NxK and PxK experiments. Non-structural carbohydrate (NSC) content of herbage was decreased by applied N but was unaffected by applications of P, K and S. Neither neutral detergent fibre (NDF) nor acid detergent fibre (ADF) was affected by applied fertiliser. In this study herbage P declined and herbage Ca increased with increasing levels of applied N. The addition of fertiliser K resulted in lower herbage Ca values. There was no herbage S response to applied fertiliser in this study. Classification and regression tree (CART) analysis identified the primary determinant of apparent intake in experiments containing N as a factor as the amount of material available to be grazed and that NSC, NPN and ADF are also determinants of apparent intake. Cows do not regulate diet choice within the short-term time frame of a meal. Thus intake is determined by short-term needs rather than by meeting long-term nutrient requirements. Fibre creates physical fill within the rumen, thus restricting intake. High NPN content is associated with high nitrate values. The reduction in intake of herbage with high nitrate content may be due to reduced palatability or to a negative feedback system limiting the intake of nitrate and ammonium. Increased NSC content is associated with increased intake, possibly through the effect of sugar on herbage palatability.

Pine plantations have been established on secondary grassland on the dune systems of the Eastern Shores of Lake St Lucia, KwaZulu-Natal, South Africa for the past 40 years. These plantations have been progressively felled for the past six years, and will continue to be felled until the year 2011, by which time they will be eliminated. Space-for-time substitution was used to determine the direction of both woody and herbaceous vegetation succession and to predict possible future management implications for the Eastern Shores. Soil samples were taken from undisturbed grassland, grassland with trees, dune forest, pine plantations, and clearfelled areas at various successional stages. To determine the effects of the pine plantations on the soils of the area, soils were subjected to particle size analysis, and determination of pH, organic carbon, phosphorus, exchangeable bases, iron and aluminium. Minimal modification of the sandy soils by the pine plantations was found to have occurred. That which did occur was shown to be short-term, and to be ameliorated by the establishment of an indigenous woody understorey, resembling pioneer dune forest. Soil under plantations was shown to have a lower pH and cation exchange capacity than under opposite indigenous vegetation but no other direct effects were observed. The direction of succession was determined by the nature of the indigenous vegetation adjacent to the plantation. Pine plantations were shown to facilitate succession towards dune forest by the exclusion of fire, provision of perches and refugia for forest-dwelling animals, and creation of a forest environment for the establishment of trees. However, the extent of re-establishment of indigenous dune forest under pine plantations was shown to be directly related to the nature of the adjacent indigenous vegetation, be it grassland, grassland with trees or dune forest. / Thesis (M.Sc.)-University of Natal, Pietermaritzburg, 1998.

The KwaZulu-Natal Drakensberg, South Africa, is predominantly a grassland ecosystem maintained by fire. The effect of the current burning regime on invertebrate and flowering forb diversity in this ecosystem is poorly understood. The overall aim ofthis study was to contribute towards the development of an effective burning regime for the KwaZulu-Natal Drakensberg that will conserve invertebrates and indigenous forbs, two major components of biodiversity. The objectives were to examine the effect of fire and fire frequency on flowering forb and invertebrate species diversity, to determine whether fire frequency, time since last burn or locality were influencing species composition, and to identify potential biodiversity indicators that reflect overall species richness for use in monitoring of invertebrates and forbs. Sampling took place in March, September and November of 2002 at Giants Castle Game Reserve. Invertebrates were sampled using sweep netting and targeted netting along transects, yellow pan traps and soil quadrats. Invertebrate taxa sampled were ants (Formicidae), butterflies (Lepidoptera), grasshoppers (Orthoptera), leafboppers (Cicadellidae), bees (Apoidea), bee flies (Bombyliidae), hover flies (Syrphidae), robber flies (Asilidae), spiders (Araneae), earthworms (Oligochaeta) and millipedes (Diploda). These were identified to species level with the assistance of taxon experts. Flowering forbs were sampled using five replicates of five by five metre quadrats randomly placed in each site. Overall flowering forb and invertebrate species diversity was higher in grasslands that were burnt for two consecutive years in 2001 and 2002 than in grasslands that were not burnt during those two years. Frequently (annual) and intermediately (biennial) burnt grasslands had significantly higher invertebrate and flowering forb diversity than infrequently (five years without burning) burnt grasslands. This, together with the fact that grasslands burnt during the year of sampling had higher species richness than grasslands burnt two and five years previously suggests that invertebrates and forbs are generally resilient to fire and many forb species appear to be stimulated by fire. However, each burn frequency had its own suite of unique flowering forb and invertebrate species. Invertebrate communities were influenced mostly by locality and the length of time past since the last fire and flowering forb communities were influenced mostly by the length oftime past since the last fire. Fire frequency had the least influence on both invertebrate and forb communities. Ecological succession occurred after each fire in the invertebrate communities but forb communities appear to need more than five years without fire for ecological succession to occur. The findings of this study therefore suggest that using a combination of three fire frequencies would result in patches of grassland in various stages of ecological succession, and would conserve species unique to each burning frequency, and would therefore conserve maximum diversity. Flowering forb species richness and certain invertebrate taxa (ants, leafboppers, spiders and bees) have the potential to act as indicators of overall invertebrate species richness for use in monitoring programmes. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2006.

The Okavango Delta is an extremely dynamic system with variable vegetation comprised of permanent swamps, seasonal swamps, dry islands, floodplains and dry grassland, savanna and woodland. The system is largely driven by the interaction between fire and the annual flood, which filters down from the Okavango River catchments in Angola. While extensive research has been conducted on the flood-driven vegetation little is known about the dry woodland and savanna regions bordering these flood-driven habitats. A taxonomic classification of woody species composition resulted in eleven vegetation types. These data were then reanalyzed in terms of woody species morphology allowing these eleven vegetation types to be grouped into four functional response groups in order to provide a platform for improving the understanding of how dry woodland and savannas interact with the environment. These four groups were the savanna group mixed thornveld and the three woodland groups; mixed broadleaf woodland, shrub mopane woodland and tall mopane woodland. Burning in mixed thornveld and mixed broadleaf woodland was found to decrease woody species density and grass fuel loads and could be used for grazing management to remove unpalatable growth and improve grass species composition, while burning in shrub mopane woodland and mixed mopane woodland merely decreased the woody understory and is not recommended. Utilization dominated by grazing livestock resulted in overutilization of the grass sward leading to bush encroachment in both mixed thornveld and shrub mopane woodland, while utilization by goats alone resulted in underutilization of the grass sward and a dominance of herbaceous annuals. Livestock utilization had no effect on the occurrence of Pecheul-loeschea leubnitziae, a shrubby pioneer previously thought to be an indicator of overgrazing, however extensive P. leubnitziae cover was associated with a sward dominated by shade-tolerant grasses with low forage quality. Shrub mopane woodland and tall mopane woodland appear to be more stable vegetation states than mixed broadleaf woodland and mixed thornveld being less vulnerable to colonization by pioneer species and alteration as a result of utilization or environmental factors. For this reason management and monitoring of mixed thornveld and mixed broadleaf woodland is essential to prevent vegetation degradation and to ensure optimal forage availability for both livestock and wildlife. / Thesis (Ph.D.)-University of KwaZulu-Natal, Pietermaritzburg, 2012.

The grazing management project in the Okhombe ward of the Amazizi Tribal Authority formed part of the National Department of Agriculture's LandCare program to address communal natural resource management issues. Okhombe land is communal whereby every member of the community is the legal owner of the rangeland with individual ownership of stock with the chieftaincy playing a major role in land allocation. In order to avoid critics of the past and address the top-down approach of the past interventions, a participatory approach was conducted in the planning and implementation of the grazing system. The service providers held a series of visioning workshops with the community in an effort to better understand community resource use patterns, needs, constraints and opportunities as part of the participatory approach. Issues identified by the community were the need for fencing grazing camps, animal health improvement, subdivision of rangeland and crop fields and the development of a rotational grazing system. The main aims of this study were to develop a participatory grazing plan with the community, develop and support institutional structures governing range management, and build capacity of the community in range management. The effect of the current grazing system on species composition was determined. In addition to these, the project investigated the potential different fodder trees has on alleviating feed and nutritional deficit, particularly during the dry winter months of the Upper Thukela. Among the main achievements of this study was the development and strengthening of local institutional structures and effective liaison by all structures with the Inkosi and the tribal council. The community developed a rotational grazing plan, marked the camp boundaries, produced digital maps and successfully built fence boundaries (approximately 20 kms of fencing) to divide their land. The fence boundaries separated the crop fields and rangeland, closed ward boundaries in the upland to prevent access by cattle from neighbouring wards, and divided the land into three camps. Six crush pens were constructed in each subward of the Okhombe ward. A communal herders fund opened and fence construction improved crop yields due to a decrease in crop damage by cattle. Okhombe ward, located in the Highland Sourveld region of KwaZulu-Natal, experiences feed and nutrition deficits to ruminants during winter. The prevailing species composition in Okhombe was investigated as part of the grazing plan. The veld condition of the sites ranged from poor (40.7%) in the bottomland to an averaged of 47.0% in upland sites. The most distinctive feature of the rangeland in this area was the loss of highly palatable Decreaser grass species (P <0.05), such as Themeda triandra in the bottom slopes « 1300 m) when compared to the upland (> 1800 m). The proportional abundance of Decreaser species accounted for an averaged of 1.02% of the bottomland and an averaged of 11.5% of the upland compared to the values of 49% in the benchmark (grassland in optimal condition). The composition of the less palatable Increaser Il species was very high at all elevations (1200 m -80.7%, 1400 m - 75.8% and 1700 m - 55.7%) when compared to the low benchmark composition of 19%. The dominant grasses of the bottom slopes were Increaser Il species, such as Eragrostis curvula, Eragrostis plana and Sporobolus africanus and unpalatable Increaser III species such as Aristida junciformis. A significant difference (P < 0.05) in the composition of Decreaser, Increaser I and Increaser Il species was found between the bottom and slopes compared to the upland region. However, the grass cover formed by these tufted species was generally high, making it more resistant to physical degradation. The bottom slope ranged from reasonable to excellent cover (16.9%), the middle slope ranged from reasonable (15.9%) to a good cover of 18.1%, averaging 16.7% and a range of 16.1% to 17.9% for the upland plateau. In the agroforestry trial the potential of different fodder species for supplementing fodder was examined. Leucaena leucocephala had the potential of being a suitable fodder tree species for use in alley cropping (P < 0.05) compared to Morus alba and Acacia karroo. Results from the partially intercropped treatments showed that L. leucocephala yield (665 kg ha-I) varied significantly (P < 0.05) from the A. karroo (378 kg ha-I) and M alba yield (345 kg ha-I). Treatments that were fully intercropped varied, but no significant difference (P > 0.05) were recorded. Morus alba produced the least yield of 345 kg ha-I, A. karroo yielded 378 kg ha-1 and 1. leucocephala recorded the high of 664 kg ha-I. Results from the second season showed similar trend in that 1. leucocephala yielded a significant (P < 0.05) fodder production of 1715 kg ha-I in comparison to M. alba (1101 kg ha-I) and A karroo (1140 kg ha-I). M alba yielded the least dry matter production (P < 0.05) but had high potential (P < 0.05) for addressing lack of firewood in rural areas. Morus alba yielded high fuel wood production from both two seasons. There were no significant differences in fuel wood yield (P > 0.05) from the partially intercropped M alba (507.9 kg ha-I) and 1. leucocephala (455.0 kg ha-I) but the yield from both species varied significantly from the A. karroo yield (103.kg ha-I). With regard to fully intercropped plots, fuel wood yield from all tree species varied significantly, A karroo resulting in low yield (63 kg ha-I), 1. leucocephala recorded 243 kg ha-l and M alba the highest yield of 444 kg ha-I. In the second season, M. alba yielded an averaged fuel wood production of 728 kg ha-l and a low of 439 kg ha-I from 1. leucocephala. Acacia karroo, a slow growing indigenous tree, might be preferred by farmers due to its less branches resulting in minimal light competition with crops. Leucaena leucocephala tend to grow slowly in its initial establishment stage, but once roots become well established, it grows fast and produces high quantity of fodder. The effect all fodder trees had on crop yield was not negative during the trial period and further research on long term effects of alley cropping is recommended. The conclusions drawn here were based on tree growth and their likely impact in alley cropping. Leucaena leucocephala was also recommended as a preferred species for rural ruminants based on the forage quality study. The results showed high content of crude protein (19.27%), low NDF content (50.38%) and very low tannin content (1.19%) from 1. leucocephala compared to A. karroo with a high tannin content of 5.69%. Acacia karroo had a crude protein content of 13.60%, NDF percentages of 44.16 and 34.64% of ADF content. Morus alba also had a recommended chemical composition of 11.71% of CP, 42.86% of NDF, 36.96% of ADF and a low tannin content of 0.65%. L. leucocephala foliage proven is readily degradable under different diet ranging within 24 hrs of intake (P < 0.001) compared to other feeds. L. leucocephala had high dry matter loss degraded from the rumen under Eragrostis hay diet with poor nutrients to high protein concentrates diet. Under the Eragrostis hay diet for instance, L. leucocephala tend to degrade rapidly with values of dry matter loss ranging from 32.2% to 39% at 4 hrs to 16 hrs, when compared to low dry mater loss of 26% at 4 hrs to 31.33% at 16 hrs. Feeds such as M alba tend to degrade slowly within 24 hrs of intake and rapidly degrades after the stated period. The ep content of maize stover was very low ranging from 1.60% in maize stalks to 2.63% in maize leaves. The fibre content in maize stover was very high when compared to lower values in fodder samples. The NDF content ranged from 77.92% in maize leaves to 81.60% in maize stalks. Maize leaves when compared to a combination of maize leaves and maize stalks sole tend to degrade better within 24 hrs of intake. This was due to low (P < 0.05) degradability rate of maize stalks compared to a combination of maize stalks and leave and leaves sole and least NDF content in maize leaves might have attributed to these results. Due to poor chemical compositions of these roughage samples, the study recommended the establishment of fodder banks and agroforestry systems to curb the nutrients deficit during winter. In conclusion therefore, this study highlight that the sustainability of rural systems to manage communal grazing land should be further explored. Most of the challenging issues in communal range management are social in nature rather than technical concepts. These include ways of improving social contributions from cattle to the community while maintaining cultural values of the use of cattle. The interventions in communal range management by service providers should understand the institutional arrangements within a community and an attempt to strengthen such existing structures is recommended. Further interventions by service providers in Okhombe ward should bring in the planning discussions, experts from social sciences, to deal with understanding of community dynamics. Complexities in communal range management involve dealing with non-stock owners within project boundaries. Communities from neighbouring wards should not be ignored and ways of improving communications and updating project details to them should be formulated. Shortage of land and closing of ward boundaries to prevent access to land by neighbouring wards is among community complexities to be explored. Communities in rural lands do share land and in most cases boundaries are known but invisible by an outsider to identify. It is important to strengthen and maintain every success in communal lands as that may form core of the project. Successes on grazing management by locals is far from being the improvement in veld but there are rather various factors to the successes of grazing projects in rural areas. Examples of successes based on Okhombe project are reduction in stock theft, improved in relationship between community and locals institutions, a reduction in stock mortalities during winter and improved animal health. Veld improvement is among successes but there are accomplishments phases to fulfil before focusing on improvement of species composition. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2005.