Le bilan de masse en surface (noté BMS ; l'accumulation de neige diminuée de l'ablation) de la calotte glaciaire antarctique est sensible aux paramètres climatiques et contribue directement aux variations du niveau moyen des mers. Il est donc important, dans le cadre de la prévision du changement climatique, de développer des outils capables de simuler les processus physiques régissant le bilan de masse en surface antarctique. L'approche développée dans cette thèse consiste à utiliser une cascade de modèles atmosphériques allant de la grande échelle vers l'échelle locale. Ainsi, un modèle climatique régional (Modèle atmosphérique régional, MAR), forcé par des réanalyses du Centre européen pour les prévisions météorologiques à moyen terme (CEPMMT), fournit à un modèle diagnostique de désagrégation physique des précipitations les champs météorologiques nécessaires calculés à l'échelle régionale (typiquement, de résolution 40 km). Dans un premier temps, il est montré que le bilan de masse en surface généré par le MAR est conforme aux observations dans la plupart des régions. Toutefois, le ruissellement est surestimé ; ce problème disparaît en introduisant une dépendance de l'albédo avec la distance zénithale de l'astre solaire. Dans un second temps, il est montré que malgré la relative simplicité des paramétrisations physiques du désagrégateur, la connaissance du relief de fine échelle (de résolution 5 km) permet d'améliorer la variabilité spatiale de la précipitation, et, par conséquent, du BMS, sur les régions côtières de l'Antarctique. La validation est menée à l'aide, notamment, de mesures de hauteurs de neige délivrées par des stations météorologiques automatiques. Sur le site côtier de Law Dome, le gradient d'accumulation nette est davantage dû au forage orographique subi par la précipitation qu'au processus de chasse-neige. Le modèle de désagrégation sous-estime fortement la précipitation sur le plateau Antarctique, où les nuages stratosphériques polaires associés au refroidissement radiatif pourraient jouer un rôle dans la génération de la précipitation pendant la nuit polaire.

Les séismes sont à l'origine d'évènements sédimentaires gravitaires (turbidites) dont l'étude permet de reconstituer l'histoire mal connue de la paléosismicité des marges continentales. L'analyse d'une série de carottes de sédiments, collectées stratégiquement dans trois systèmes turbiditiques de la marge en subduction Hikurangi de Nouvelle-Zélande, permet d'établir les caractéristiques, les facteurs de contrôle et les mécanismes déclencheurs de la sédimentation gravitaire des derniers 18,000 ans. La succession sédimentaire comprend quatre lithofaciès et modes de dépôt : hémipélagite (sédimentation marine), turbidites (courants de turbidité), débrites (débris flows) et tephra (retombée de cendres volcaniques), dont l'organisation générale dépend de la morphologie de la marge et des fluctuations glacio-eustatiques. Les crues, les éruptions volcaniques et les " slope failures " sont les trois mécanismes déclencheurs des turbidites. Plus de 90% sont déclenchées par des " slope failures " en haut de pente (150 - 1,200 m) à la suite de séismes. L'adaptation d'un modèle empirique de stabilité de pente suggère que ces turbidites représentent l'enregistrement sédimentaire des ruptures répétées de trois failles actives, dont l'interplaque, et correspondent à des séismes de Mw≥7.3 avec un temps de retour de 150±50 ans. Parmi ces turbidites co-sismiques, 20 montrent une synchronicité de déclenchement sur l'ensemble de la marge et un volume important. Elles correspondent à des ruptures de la zone interplaque de Mw 7.5 - 8.4, dont les temps de retour montrent une phénomène de clustering où alternent les périodes actives à faible temps de retour (305 - 610 ans), et les périodes de quiescence à temps de retour élevé (1480 - 2650 ans). Ce calendrier paléosismique intégré aux modélisations en cours devrait permettre de mieux appréhender l'aléa sismique et les risques pour la population.

De par sa position, le Golfe du Lion est un site idéal pour l’investigation des changements paléo-environnementaux et des processus affectant le dépôt sédimentaire. Les travaux antérieurs ont permis de mettre en évidence les impacts de la variabilité climatique et glacioeustatique sur l’organisation stratigraphique de la marge, mais également sur les exports terrestres de matière organique.L’étude isotopique du carbone organique et de l’azote de la matière organique dans les sédiments du forage PRGL1-4 nous a permis de mettre en évidence de forts exports fluviaux lors des interstades survenus au cours des 200 000 derniers milles ans. La mise en regard de cette découverte avec les enregistrements paléo-climatologiques terrestre et marin disponibles dans la région indique que ces forts exports fluviaux résultent d’une augmentation des précipitations le long de la bordure Nord Méditerranéenne. Grâce à la position dePRGL1-4, nous proposons que ces pluies soient le résultat d’une augmentation du passage de dépressions Nord Atlantique dans le bassin Ouest Méditerranéen.Une caractérisation des isotopes du soufre préservés dans la pyrite sédimentaire a été réalisée. Les résultats obtenus ont permis de mettre en évidence une variation isotopique insoupçonnée, l’une des plus grandes observées de nos jours, dont la cyclicité semble indiquer un fort contrôle climatique. Nous proposons deux mécanismes influençant le fractionnement isotopique: une modulation de l’activité bactérienne par le climat, et/ou (ii) une modulation locale liée la nature des sédiments impliqués dans la formation des pyrites en lien avec les variations eustatiques. / By its position, the Gulf of Lion is an ideal location for investigation of past ecological changes and processes affecting the sedimentary deposition. Previous work has highlighted the impacts of climatic and glacio-eustatic changes on the GoL stratigraphic organization, but also on terrestrial exports of organic matter.This isotopic study based on the organic carbon and nitrogen preserved in PRGL1-4 sediments highlights important rivers runoff during warm periods of the last 200 000 years.Regional intercomparison with terrestrial and marine records indicates that these river exports resulting from an increase of precipitation over the North Mediterranean borderland.Using PRGL1-4 location, out of Mediterranean cyclogenetic area, we suggest that these pluvial events occurred in response to enhance passage of North Atlantic atmospheric perturbation into the Western Mediterranean basin.Pyrite sulfur isotopes investigations over the last 500 kyr have also been done. The stratigraphic variations (up to 76‰) in the isotopic data reported here are among the largest ever observed in pyrite, and are in phase with glacial-interglacial sea level. These results suggest that there exist important but previously overlooked depositional controls on sedimentary sulfur isotope records. Two different mechanisms influencing the isotopic fractionation can explain the observed dataset: a climatic modulation of the bacterial activity, and / or (ii) a local sedimentary modulation involve during early diagenetic formation of pyrite in relation with the eustatic variations.

L’introduction de petites faunes au sein du régime alimentaire humain dans le passé constitue un sujet de recherche essentiel dans la compréhension des modes de subsistance des chasseurs-cueilleurs, notamment au Paléolithique supérieur. Durant cette période, et notamment au Dernier Maximum Glaciaire, le lapin de garenne (Oryctolagus cuniculus) domine les assemblages fauniques de nombreux sites archéologiques de la Péninsule Ibérique. Plusieurs hypothèses ont été avancées afin de comprendre le rôle que ce petit gibier a pu jouer dans la subsistance des groupes humains. Néanmoins, la multitude et la variété de ces hypothèses reflètent, d’un côté, le manque de consensus quant à la portée de l’exploitation du lapin dans la subsistance durant cette période dans la Péninsule Ibérique, et elles soulignent d’un autre côté que l’appréhension des implications économiques, technologiques, environnementales et sociales de son exploitation est une tâche complexe. A partir de ce double constat, j’ai dès lors établi comme problématique d’explorer les potentielles motivations liées à la chasse du lapin et d'appréhender le rapport entre les humains et cette espèce de petite taille, et ce dans l’objectif d’obtenir une meilleure compréhension des modes de subsistance adoptés pendant cette période en Ibérie. L’approche développée dans cette thèse repose sur la proposition que l’explication du pourquoi les humains exploitaient le lapin doit être traitée par une compréhension du comment les humains chassaient ce petit gibier. Si nous voulons analyser le régime alimentaire humain dans le passé, nous devons effectivement comprendre comment les humains prenaient leurs décisions cynégétiques. A cette fin, un des cadres théoriques principalement utilisé en archéologie, la Optimal Foraging Theory, a été employé, en complément de la modélisation à base d’agents. La problématique a été répartie en trois questions de recherche distinctes et complémentaires, chacune faisant l’objet d’une publication. Dans le premier article, l’application du Diet Breadth Model issu de la Optimal Foraging Theory a permis d’observer que l’exploitation d’une garenne est optimale lorsque plusieurs individus capturent un minimum de 7 lapins à l’aide de filets au cours du Dernier Maximum Glaciaire dans la Péninsule Ibérique. Dans le second article, nous avons mené des expériences de simulation à travers le développement d’un modèle à base d’agents pour évaluer les implications de ce type d’exploitation dans un cadre plus réaliste au sein duquel les humains interagissent, avant de l'intégrer, dans le troisième article, dans différents contextes socio-économiques qui lui sont associés et observés dans le registre ethno-historique. Nos résultats montrent que la composition du régime alimentaire humain et, par-là, le retour énergétique quotidien, sont influencés par l'organisation sociale entre les membres d'un groupe, la stratégie de chasse et la technologie associée afin d’exploiter une garenne. Particulièrement, la rentabilité de l’exploitation des ressources est encore plus grande si les femmes, les enfants et les ainés sont impliqués dans l’exploitation des garennes à travers des battues et/ou l’aide de filets. Par conséquent, cette thèse expose que le développement de formations socio-culturelles optimales basées sur une division des tâches de subsistance selon l’âge et/ou le sexe (et de la technologie qui leur est associée) a pu permettre aux groupes humains de s'adapter aux environnements de la Péninsule Ibérique au Dernier Maximum Glaciaire. Finalement, le lapin ayant probablement été un complément alimentaire important pendant cette période, cette thèse souligne la nécessité de considérer son exploitation (et celle de petites faunes en général) lors de l’interprétation du registre zooarchéologique car des changements d’un point de vue économique, technologique mais aussi socio-culturel dans les modes de subsistance adoptés dans le passé peuvent être ainsi appréhendés. / The introduction of small fauna into the human diet in the past is an essential research topic in the understanding of hunter-gatherer subsistence patterns, particularly in the Upper Paleolithic. During this period, and particularly during the Last Glacial Maximum, the European rabbit (Oryctolagus cuniculus) dominates the faunal assemblages of many archaeological sites in the Iberian Peninsula. Several hypotheses have been proposed to understand the role played by this small game in the subsistence of past human groups. Nevertheless, the number and diversity of these hypotheses first reflect a lack of agreement as to the extent to which rabbits played a role in subsistence during this period in the Iberian Peninsula. Secondly, they underline the complexity of understanding the economic, technological, environmental, and social implications of rabbit exploitation. From these observations, I established as the problematic to explore the potential motivations behind rabbit hunting, and to apprehend the relationship between humans and this small game, with the aim of providing a better understanding of the subsistence patterns adopted during this period in Iberia. The approach developed in this thesis is based on that the explanation of why humans exploited rabbits must be addressed by an understanding of how humans hunted them. If we are to analyze human diet in the past, we must indeed understand how humans made their hunting decisions. To this end, one of the theoretical frameworks mainly used in archaeology, the Optimal Foraging Theory, has been adopted, complemented by Agent-Based Modeling. I divided the problematic into three distinct and complementary research questions, each the subject of a publication. In the first article, the application of the Diet Breadth Model derived from the Optimal Foraging Theory enabled us to observe that the exploitation of a warren is optimal when several individuals capture a minimum of 7 rabbits using nets during the Las Glacial Maximum in the Iberian Peninsula. In the second paper, we carried out simulation experiments through the development of an Agent-Based Model to assess the implications of this type of exploitation in a more realistic environment in which humans interact, before integrating it, in the third paper, into different socio-economic contexts related to it and observed in the ethno-historical record. Our results show that the composition of the human diet and, consequently, the daily energetic return are influenced by the social organization between group members, the hunting strategy and the associated technology employed for exploiting a warren. In particular, the energetic efficiency of resource exploitation is even greater if women, children, and elders are involved in exploiting warrens through net hunting and/or drives. Consequently, this thesis argues that the development of optimal socio-cultural group configurations based on an age and/or gender-based division of labor (and their associated technology) may have enabled human groups to successfully adapt to the environments of the Iberian Peninsula during the Last Glacial Maximum. Finally, as rabbit was probably an important energetic source during this period, this thesis highlights the need to consider its exploitation (and that of small game in general) when interpreting the zooarchaeological record, as not only economic and technological, but also socio-cultural changes in subsistence patterns adopted in the past can be apprehended.

Cette recherche a été effectuée dans la baie de Kuuvik (Nunavik, Canada) pour mieux comprendre les occupations dorsétiennes et thuléennes-inuit dans un contexte biophysique postglaciaire en évolution. Des analyses géomorphologiques, stratigraphiques, micromorphologiques, macrofossiles et géochimiques (FTIR et ICP-AES) ont été effectuées sur une maison semi-souterraine unifamiliale hivernale thuléenne-inuit du site Paaliup Qarmangit 1 et dans la vallée dans laquelle elle s’inscrit afin de documenter les processus de formation et la taphonomie de la structure étudiée. Les données extra-sites ont permis de reconstituer l’évolution des environnements sédimentaires dans la vallée depuis la dernière glaciation. L’approche intra-site a révélé des occupations dorsétienne (après 143-327cal A.D.) puis thuléennes-inuit (entre 1317-1413 cal A.D. et1466-1642 cal A.D) du site. Le caractère stratégique du lieu d’implantation (accessibilité aux matériaux de construction, aux ressources alimentaires et hydriques) pourrait expliquer son utilisation par deux cultures différentes successives. Les résultats montrent la prédominance des processus nivéo-éoliens et de nivation dans la formation de l’unité stratigraphique contenant les restes archéologiques et attestent d’un remaniement post-dépositionnels naturels et/ou anthropiques des artefacts dorsétiens. Des signatures chimiques anthropiques modérées mais significativesont été détectées dans les sols de la maison semi-souterraine (e.g., Mg, Fe, S). Elles pourraient avoir été atténuées par des processus pédologiques, tels que le lessivage et la remobilisation des éléments par les organismes du sol, et/ou des processus anthropiques, tels qu’un nettoyage régulier de la structure par ses habitants. / This research was carried out in Kuuvik Bay (Nunavik, Canada) to document Dorset and Thule-Inuit occupations in an active periglacial context. Geomorphological, stratigraphical, micromorphological, macrofossil and geochemical (FTIR and ICP-AES) analyses were performed over a single-family semi-subterranean Thule-Inuit house of Paaliup Qarmangit 1 site and the surrounding periglacial valley to document site formation processes of the studied structure and to identify anthropogenic chemical signatures in the soil of the house. Off-site data enabled to reconstruct the sedimentary environments in the valley since the last glaciation. In-site approach revealed a Dorset occupation of the site (after 143-327cal A.D.) prior Thule-Inuit settlement (between 1317-1413 cal A.D. and 1466-1642 cal A.D.). Strategic features of the site (such as availability of building material, food, and water resources) may explain its use by two different cultures. Results showed the predominance of niveo-aeolian and nivation processes in the formation of the unit containing archaeological remains and provided evidence of post-depositional natural and/or anthropogenic reworking of Dorset artefacts. Soil of the house recorded moderate but significant anthropogenic chemical signatures (e.g., Mg, Fe, S), which may have been buffered by pedological processes, such as leaching and biological remobilization, and/or anthropogenic processes, such as a regular cleaning (maintenance) of the structure

I. An original methodological discussion is proposed on the problem of the typology of tropical rain forest’s plant communities, based on the study of forest types across gradients of continentality and elevation, within Atlantic central Africa. These investigations were based on the statement that the main problems in forest typology are related to the non-zonal or zonal character of the different vegetation types and to non considering the relations and differences between forest strata. II. Field data consisted in phytosociological homogeneous sample plots localized within different recognized phytogeographical entities, in a region of tropical Africa where these entities are known to be well conserved. A total of 37 such plots were inventoried in the region extending from the littoral forests of Ndoté, Equatorial Guinea, which are wet evergreen forests, to the continental forests of the Dja, Cameroon, known as evergreen seasonal forests. The studied region also included the oriental Atlantic forests of Equatorial Guinea, known as moist evergreen forests or caesalp forests. In various parts of this continentality gradient, some plots were localized within climax non-zonal formations, namely the submontane rain forests. The emphasis was put on the vegetation of the Monte Alén National Park. The sampling methodology was willing to be as "complete ", including all strata, "quantitative ", enumerating all individuals, and "representative ", within each stratum, as possible. These multi-layers plots were realised using nested sub-plots, with a sampling size of 100 individuals for every ligneous stratum recognized (dominant trees, dominated trees and shrubs) and a sampling size of 200m² for the herbaceous and suffrutex stratum. Forest types were defined independently for each stratum and the differences were analysed. A method was proposed for the simultaneous analysis of all floristic data, converting and standardizing the values from ligneous strata, on the one hand, and from understorey strata, on the other hand. III. Ten forest types were described using IndVal and discussed in the general context of the guineo-congolian region, from a syntaxonomic view point (agglomerative classification) and from a phytogeographical view point (divisive classification). Homologies between these two approaches are described. The proposed phytogeographical system is based on an "open " conception of hierarchical classifications, combining advantages of agglomerative and divisive classifications. In concrete terms, the non-zonal criteria, for example the submontane variants, are categorised separately and in analogy with the zonal criteria, related to the usual phytochoria. Analysis of ecological relationships for the 10 communities showed that the main variables related to the floristic variability in our mainland rain forests are elevation, rainfall, hygrometry (estimated using bryophytes cover levels) and distance to the ocean. The two extremes on the vertical microclimatic gradient, dominant trees stratum and herbaceous stratum, give similar typologies, however canonical analysis showed that for the herbaceous layer, non-zonal variables (hygrometry and elevation) were gaining more importance when the influence of the two zonal variables was attenuated. In every case, spatial autocorrelation was less important than the environment in explaining floristic variability but its role increased in the spatial arrangement of understorey species, whose dispersal capacity is generally lower than canopy trees. The phytosociological, phytogeographical and ecological description of forest types is accompanied by a physiognomical description using biological types spectrum, as well as architectural models, leaf sizes, etc. With regard to diversity, we have demonstrated that species richness was higher from upper to lower strata because of the accumulation in lower strata of species from various strata. On the other hand, the proper stratum diversity, i.e. the structural set, decreased from dominant trees to shrubs. The proper diversity of the herb layer showed relatively high figures mainly due to the higher individual density in relation to the existence of microstrata. Within the 37 sample plots, 1,050 taxa have been identified to species or morpho-species levels, for a total of 25,750 individuals. These taxa represent 442 genus among 104 families. The richest forest type is found on the foothills of the Niefang range, on the windward side. This forest type is also characterised by a high number of oligotypic genus and by species belonging to functional types indicators of glacial refuges. These functional types are defined on the basis of the dispersal capacity and on kind of stand needed for effective germination. We formulated the hypothesis that this kind of "foothills refuge ", characterised by his zonal nature, could have been one of the rare refuges for species from mainland rain forests, while montane and fluvial refuges would mainly have preserved species from non-zonal forest types: (sub)montane and riverine. Based on indicator species of submontane forests, a potential distribution map of this forest type has been realised at the Atlantic central African scale. More than 400 submontane forest localities have been mapped. These forests begin at 400m of altitude near the ocean, and progressively at higher altitude for increasing distance to the ocean. Many lowland localities also comprised submontane species, which could indicate the existence of ecological transgressions. These transgressions would allow migratory tracks for submontane species between isolated mountain ranges, not only during glacial periods, through heights at the northern and southern borders of the congo basin, but also contemporarily through the lowland riverine forest network, in the centre of this basin. Finally, a special attention has been attributed to littoral forests and to some cases of choroecological transgressions, coupled to the ecological equalization phenomenon.

I. An original methodological discussion is proposed on the problem of the typology of tropical rain forest’s plant communities, based on the study of forest types across gradients of continentality and elevation, within Atlantic central Africa. These investigations were based on the statement that the main problems in forest typology are related to the non-zonal or zonal character of the different vegetation types and to non considering the relations and differences between forest strata.<p><p>II. Field data consisted in phytosociological homogeneous sample plots localized within different recognized phytogeographical entities, in a region of tropical Africa where these entities are known to be well conserved. A total of 37 such plots were inventoried in the region extending from the littoral forests of Ndoté, Equatorial Guinea, which are wet evergreen forests, to the continental forests of the Dja, Cameroon, known as evergreen seasonal forests. The studied region also included the oriental Atlantic forests of Equatorial Guinea, known as moist evergreen forests or caesalp forests. In various parts of this continentality gradient, some plots were localized within climax non-zonal formations, namely the submontane rain forests. The emphasis was put on the vegetation of the Monte Alén National Park.<p><p>The sampling methodology was willing to be as "complete ", including all strata, "quantitative ", enumerating all individuals, and "representative ", within each stratum, as possible. These multi-layers plots were realised using nested sub-plots, with a sampling size of 100 individuals for every ligneous stratum recognized (dominant trees, dominated trees and shrubs) and a sampling size of 200m² for the herbaceous and suffrutex stratum.<p><p>Forest types were defined independently for each stratum and the differences were analysed. A method was proposed for the simultaneous analysis of all floristic data, converting and standardizing the values from ligneous strata, on the one hand, and from understorey strata, on the other hand.<p><p>III. Ten forest types were described using IndVal and discussed in the general context of the guineo-congolian region, from a syntaxonomic view point (agglomerative classification) and from a phytogeographical view point (divisive classification). Homologies between these two approaches are described. The proposed phytogeographical system is based on an "open " conception of hierarchical classifications, combining advantages of agglomerative and divisive classifications. In concrete terms, the non-zonal criteria, for example the submontane variants, are categorised separately and in analogy with the zonal criteria, related to the usual phytochoria.<p><p>Analysis of ecological relationships for the 10 communities showed that the main variables related to the floristic variability in our mainland rain forests are elevation, rainfall, hygrometry (estimated using bryophytes cover levels) and distance to the ocean. The two extremes on the vertical microclimatic gradient, dominant trees stratum and herbaceous stratum, give similar typologies, however canonical analysis showed that for the herbaceous layer, non-zonal variables (hygrometry and elevation) were gaining more importance when the influence of the two zonal variables was attenuated. In every case, spatial autocorrelation was less important than the environment in explaining floristic variability but its role increased in the spatial arrangement of understorey species, whose dispersal capacity is generally lower than canopy trees. The phytosociological, phytogeographical and ecological description of forest types is accompanied by a physiognomical description using biological types spectrum, as well as architectural models, leaf sizes, etc.<p><p>With regard to diversity, we have demonstrated that species richness was higher from upper to lower strata because of the accumulation in lower strata of species from various strata. On the other hand, the proper stratum diversity, i.e. the structural set, decreased from dominant trees to shrubs. The proper diversity of the herb layer showed relatively high figures mainly due to the higher individual density in relation to the existence of microstrata. Within the 37 sample plots, 1,050 taxa have been identified to species or morpho-species levels, for a total of 25,750 individuals. These taxa represent 442 genus among 104 families. The richest forest type is found on the foothills of the Niefang range, on the windward side. This forest type is also characterised by a high number of oligotypic genus and by species belonging to functional types indicators of glacial refuges. These functional types are defined on the basis of the dispersal capacity and on kind of stand needed for effective germination. We formulated the hypothesis that this kind of "foothills refuge ", characterised by his zonal nature, could have been one of the rare refuges for species from mainland rain forests, while montane and fluvial refuges would mainly have preserved species from non-zonal forest types: (sub)montane and riverine.<p><p>Based on indicator species of submontane forests, a potential distribution map of this forest type has been realised at the Atlantic central African scale. More than 400 submontane forest localities have been mapped. These forests begin at 400m of altitude near the ocean, and progressively at higher altitude for increasing distance to the ocean. Many lowland localities also comprised submontane species, which could indicate the existence of ecological transgressions. These transgressions would allow migratory tracks for submontane species between isolated mountain ranges, not only during glacial periods, through heights at the northern and southern borders of the congo basin, but also contemporarily through the lowland riverine forest network, in the centre of this basin. Finally, a special attention has been attributed to littoral forests and to some cases of choroecological transgressions, coupled to the ecological equalization phenomenon.<p> / Doctorat en sciences agronomiques et ingénierie biologique / info:eu-repo/semantics/nonPublished