Submitted by Rosangela Silva (rosangela.silva3@unioeste.br) on 2017-08-11T17:24:39Z No. of bitstreams: 2 Taina de Souza 2017.pdf: 1552829 bytes, checksum: 891fdf36e760cfb11f85562d5918a74e (MD5) license_rdf: 0 bytes, checksum: d41d8cd98f00b204e9800998ecf8427e (MD5) / Made available in DSpace on 2017-08-11T17:24:39Z (GMT). No. of bitstreams: 2 Taina de Souza 2017.pdf: 1552829 bytes, checksum: 891fdf36e760cfb11f85562d5918a74e (MD5) license_rdf: 0 bytes, checksum: d41d8cd98f00b204e9800998ecf8427e (MD5) Previous issue date: 2017-02-16 / (Poaceae of the campestre formations from Guartelá State Park, Paraná, Brazil). Considered one of the greatest families among the angiosperms and with great ecological amplitude, Poaceae species can be found both in forest as well as in campestre formations. The goal of the present study was to perform the first floristic survey of Poaceae in one of the last remnants native vegetation of campestre formations in Paraná, the Guartelá State Park (GSP), besides elaborate dichotomous keys in order to identify the genera and species. The survey was carried out based on field collecting and database search. The dichotomous keys were elaborated using qualitative and quantitative characters of the genera and species. In the GSP, the family is represented by 54 species belonging to 29 genera, Paspalum L. (8 spp.), Andropogon L. (5 spp.) and Eragrostis Wolf (4 spp.) being the ones that presented the greater number of species. The number of species cited previously to the GSP increased in 29 names and from the total (54) five are considered exotic, five endemics to Brazil and two are categorized as least concern to conservation, occurring both in dry and wet areas, highlighting the importance of the GSP microenvironments conservation. Information regarding the distribution, habitat and species conservation are presented. Keywords: Campos Gerais, floristic, grasses, grassland vegetation, Poales. / (Sinopse de Poaceae das formações campestres do Parque Estadual do Guartelá, Paraná, Brasil). Considerada uma das maiores famílias entre as Angiospermas e com grande amplitude ecológica, Poaceae tem suas espécies encontradas tanto em formações florestais como em campestres. O presente estudo teve como objetivo realizar o primeiro levantamento florístico de Poaceae em uma das últimas vegetações nativas remanescentes de formações campestres no Paraná, o Parque Estadual do Guartelá (PEG), além de elaborar chaves dicotômicas para identificação dos gêneros e espécies. O levantamento foi realizado com base em coletas a campo e busca em banco de dados. As chaves dicotômicas foram elaboradas utilizando caracteres qualitativos e quantitativos dos gêneros e espécies. No PEG, a família esta representada por 54 espécies pertencentes a 29 gêneros, sendo Paspalum L. (8 spp.), Andropogon L. (5 spp.) e Eragrostis Wolf (4 spp.) os que apresentaram maior número de espécies. Ampliou-se o número de espécies citadas anteriormente para o PEG em 29 nomes e do total (54) cinco são consideradas exóticas, cinco endêmicas para o Brasil e duas estão categorizadas como pouco preocupantes para conservação, tendo ocorrência tanto em áreas drenadas quanto úmidas, destacando a importância da conservação dos microambientes do PEG. São apresentadas informações acerca da distribuição, hábitat e conservação das espécies.

Livestock grazing is a common and extensive land use practice in the United States occurring in a wide range of habitat types. As such, livestock grazing has the potential to alter ecosystem structure, function and community composition. The primary component (Chapter 1) of this thesis examined the effects of cattle grazing in a San Joaquin Valley grassland on two target taxa: reptiles and small mammals. The study took place on the Chimineas Unit of the Carrizo Ecological Reserve, San Luis Obispo County, California during Fall 2009 and Spring 2010. These taxa were sampled on matched pairs of two grazed and two ungrazed sites. Live trapping methods appropriate to sampling reptiles (coverboards and pit fall traps) and small mammals (Sherman live traps) were used to determine species richness and abundance of these taxa. Uta stansburiana (side-blotched lizard) abundances were greatest on Ungrazed1 (Fall: 1.75±0.49 captures/night, Spring: 3.58±0.35 captures/night). In addition, Chaetodipus californicus (Spiny pocket mouse) abundance was also greatest on Ungrazed1. Population estimates generated by Program CAPTURE suggest C. californicus could be 6.27 times more abundant on Ungrazed1 than Grazed1. However, sample size issues and site level effects confounded and made it difficult to determine significant differences between the grazed and ungrazed treatments for both reptile and small mammal taxa. Habitat structure at these sites was also evaluated including small mammal burrow abundance and vegetation structure. Again, differences between the grazed and ungrazed pasture could not be discerned because of site level effects occurring among matched pairs. As such, this thesis illustrates the difficulty in studying grazing and its potential effects on biotic systems because an array of variables can make unclear the differences between grazed and ungrazed areas. As a follow up study (Chapter 2) I used logistic regression to model U. stansburiana presence to examine possible sources of variation observed at trapping array locations utilized in the main study (Chapter 1). Shrubs exhibited a quasi-complete separation of data points and the three best models included: 1) Bare soil cover (AICc = 28.12), 2) Holes (AICc = 29.76), and 3) Bare soil cover + Holes (AICc = 29.90). Shrubs, bare soil cover and small mammal burrow density were all positively associated with U. stansburiana presence at array locations. Although species have general habitat requirements based upon their ecology and evolution, variations in habitat utilization exists depending upon the resources present at a specific location. Quantifying basic ecological information on a site specific basis is important to managing populations by identifying important resources and habitat components utilized by a given species on a given site.

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Spatial heterogeneity of the environment, as measured by floral diversity, composition and structure, is known to influence the distribution and diversity of invertebrates. Heterogeneity brought about by anthropogenic disturbance may be a threat to invertebrate diversity. This thesis investigates the impacts of vegetation heterogeneity at a range of scales on the diversity of invertebrate populations in modified high-altitude indigenous grasslands on the Rock and Pillar Range, Central Otago. Invertebrates were sampled in and on the edges of snow tussock fragments to assess whether species richness increased systematically with fragment area. Invertebrate composition was poorly related to fragment area, plant composition and environmental variables. Taxon richness, abundance and/or diversity for three invertebrate groups increased as fragment area decreased, perhaps reflecting an influx of species from the surrounding matrix. For snow tussock leaf invertebrates in autumn, richness and abundance were at least two times lower in tussocks exposed to the wind than those in the centre of fragments, suggesting selection of habitat may be based on microclimatic characteristics. Invertebrates were sampled from the bases of tussocks after they were clipped to simulate three levels of vertebrate grazing. Invertebrate community composition differed between sites and sampling dates but was unaffected by clipping treatment. At the higher altitude site invertebrate abundance was 1.45 times greater and Shannon-Wiener diversity (H�) 1.22 times lower than at the lower altitude site. The latter sampling date had higher abundance (2.12 times) and taxon richness (1.14 times) than the earlier date. Pitfall-trapped invertebrates in cushionfield, herbfield and snow tussock differed in community composition and often by taxon richness, abundance and diversity. Across habitats, plant composition, plant diversity and some environmental variables were correlated with invertebrate variables, but could not be separated from vegetation type. The invertebrates collected in the course of the study are listed. Four Phyla, eight Classes, 24 orders and over 300 taxa were recorded. Almost all taxa are endemic and many have limited distributions and/or are undescribed. A species list is provided with collection altitude, method and habitat type. Invertebrate assemblages from sites differing in altitude, vegetation type and level of habitat modification on the Rock and Pillar Range are compared. Sites differed in species composition and rank orders of abundance and richness. At lower elevations, invertebrate richness was at least 25% less, and standardised trap abundance at least 44% less, than that at the highest elevation. Richness and abundance of exotic invertebrates decreased with increasing altitude. This thesis highlights several points concerning the study of grassland invertebrates and heterogeneity on the Rock and Pillar Range. First, there are differences in invertebrate assemblages at a range of scales. Conserving invertebrate diversity will therefore require altitudinal sequences and different habitat types, including disturbed areas. At high elevations, tussock habitat may be disproportionately important due to its relative rarity. Second, the effects of disturbance on invertebrates were only visible at large spatial scales. Third, there is a paucity of research on New Zealand invertebrates, especially in regard to terrestrial disturbance, which has resulted in a shortfall of biological, distributional, taxonomic and ecological knowledge.

Stoats are introduced mammalian carnivores implicated in the decline of several of New Zealand�s endemic species. Most research into stoats in New Zealand has focused on beech forest habitat, especially in years of peak stoat abundance following heavy beech seedfall and peak cohorts of mice. In New Zealand, alpine grasslands occur above the altitudinal limit of beech forest (900-1000 m a.s.l.). Although previous research has shown stoats to be present there, little is known about the ecology of stoats in alpine grasslands. This research aimed to test whether alpine grasslands were a marginal habitat occupied by surplus stoats that had spilled over from beech forest populations, i.e. a sink habitat. The alternative is that alpine grasslands are a desirable habitat deliberately exploited by stoats. This question was answered using mark-recapture, radio-tracking, diet analysis and a food addition experiment. Another objective was to determine whether nest survival is higher in alpine grassland compared to beech forest and whether stoats are likely to be a frequent predator of ground nests in alpine grasslands relative to other introduced mammals that inhabit them. If nest survival is higher in alpine grassland then alpine grasslands may be a refuge from predation. However, if it is not then it is important for management to know the relative risk posed by stoats compared with other predators. An artificial nest experiment was used to answer these questions. This research was undertaken during two years of low to intermediate beech seedfall and therefore provided an opportunity to look at the ecology of stoats in a New Zealand National Park outside years of peak abundance. The principal study site for this research was the Borland Valley, Fiordland National Park. Compositional analysis showed that stoats in alpine grassland selected for it over adjacent beech forest. The range cores of these stoats were high up in alpine grassland away from the ecotone with beech forest. Stoats occurred at similar densities in alpine grasslands as they did in beech forest and observed survival was similar between the two habitats (with the exception of 2004 when it may have been higher in alpine grassland). The most frequent prey of stoats inhabiting beech forest were birds and mice. Although stoats in alpine grasslands also ate birds and mice their most frequent prey were ground weta and hare. Food addition appeared to cause diet switching but did not reduce the distances moved by stoats, suggesting that other factors may be more important in regulating their summer home range size in alpine grasslands. All of these factors lead to the conclusion that alpine grasslands in the Borland are not a marginal habitat for stoats, but may instead be a desirable one. Artificial nests had a higher probability of survival in alpine grassland compared to adjacent beech forest, but survival was too low to support the idea that alpine grasslands are a refuge. Stoats were the most frequent predator of artificial nests in both habitats, but 95 % confidence intervals overlapped the predation rate by possums, which was also high. These findings illustrate the need for a comprehensive landscape approach to stoat control in montane National Parks, for two reasons: 1) endemic biodiversity in alpine grasslands may be under threat from stoat predation, 2) alpine grasslands may act as a source for dispersing stoats that reinvade lowland stoat control areas. In the absence of heavy beech seedfall and peak mouse abundance, stoats occurred at densities of around 1 km⁻� in both habitats and there was recruitment into these populations. This raises the important question: What regulates the distribution and abundance of stoats in years of low beech seedfall and low mouse abundance? In these years birds, ground weta and hares may be as important as mice are in years of peak abundance following heavy beech seedfall.

Grassland birds have declined more than any other guild of birds in North America, largely due to loss and degradation of native grasslands. The Conservation Reserve Program (CRP) has restored some native warm-season grasses (NWSG), but grassland birds continued to decline (-1.1% annually) partly due to the limited acreage converted (1% of southeastern US). Using NWSG in production settings provides profit incentive to landowners while reducing dependency on government programs. Studies examining these production practices and their effect on grassland birds east of the Great Plains are limited. During 2009 – 2010, I surveyed 102 NWSG fields in Kentucky and Tennessee being used for production purposes (control, biofuel, seed, hay, and pasture treatments) to assess bird use and vegetation characteristics. Landscape cover around each field (250, 500, and 1000 m) was digitized from aerial photography. Using analysis of variance (ANOVA), I compared bird (relative abundance, species diversity, and species richness) and vegetation (average height, litter depth, vertical cover, litter cover, and vegetation cover) metrics across the five treatments. Relative abundance for all species, species diversity, and species richness were all greater for seed production fields (P <0.05); other treatments did not differ. Field sparrows (Spizella pusilla) were less abundant (P <0.05) in biofuel than control, hay and graze treatments, whereas eastern meadowlarks (Sturnella magna) and dickcissels (Spiza americana) were more abundant in seed fields. Average vegetation height, vertical cover, percent litter, percent forbs and percent woody plants differed (P <0.05) among treatments. Using Program Mark, I modeled occupancy for field sparrow, red-winged blackbird (Agelaius phoeniceus), eastern meadowlark, and northern bobwhite (Colinus virginianus) using vegetation and landscape cover as covariates. Treatment was influential in field sparrow and eastern meadowlark models, but not those for red-winged blackbird and northern bobwhite. Occupancy for field sparrow and northern bobwhite were affected by woody cover (+), for red-winged blackbird by vegetation height (-), and for eastern meadowlark by litter depth (+) or percent NWSG (+). All four species were negatively affected by forest within 250-m. Use of NWSG in production could increase the amount of available habitat and thus, help conservation efforts for grassland birds.

Conceptual and theoretical advancements have been developed in recent years to break down the assumptions and traditional boundaries that establish seemingly independent disciplines, and the research outlined in this dissertation aspires to build on these advancements to provide innovative solutions to a broad array of modern problems in social-ecological. I used a variety of techniques to address challenges ranging from disconnections between theory and application, perceived versus realized roles of prescribed fire in resprouting shrublands, and the need for broader participation in research as part of undergraduate education. The chapters in this dissertation serve as a case-study approach across multiple scientific disciplines that overcome the traditions and assumptions that conflict with our ability to develop innovative solutions to modern social-ecological problems. First, I bridge theoretical and applied concepts by showing how recent theoretical advancements in resilience can be integrated into a predictive framework for environmental managers. Second, experimental data from multiple experiments were collected in two ecological regions of Texas to assess the potential for using extreme fire, in isolation and in combination with herbicide, as a novel intervention approach in resprouting shrublands of the southern Great Plains. The findings from these experiments demonstrate the importance of moving past traditional assumptions of when prescribed fire should be applied to demonstrate new patterns of woody plant responses to the applications of “more extreme” prescribed fires while not causing undesirable invasions by exotic grasses and exotic insects. Finally, I initiated a PhD instructed course on undergraduate research that sought to increase undergraduate participation while lowering the costs of conducting research. This chapter shows how traditional approaches of supporting undergraduate research are incapable of meeting the broader goals established by society and reveal a novel approach that can provide an additional pathway for supporting undergraduate student participation at large, research-based universities. Ultimately, this research suggests that our capacity to enhance services in social-ecological systems ultimately hinges upon the integration of theoretical and applied concepts that drive policy and governance and overcoming the assumptions and traditions that limit their integration.

Although semi-natural grasslands in Europe are declining there is often a time delay in the local extinction of grassland species due to development of remnant populations, i.e., populations with an extended persistence despite a negative growth rate. The objectives of this study were to examine the occurrence of remnant populations after abandonment of semi-natural grasslands and to examine functional traits of plants associated with the development of remnant populations. We surveyed six managed semi-natural grasslands and 20 former semi-natural grasslands where management ceased 60-100 years ago, and assessed species response to abandonment, assuming a space-for-time substitution. The response of species was related to nine traits representing life cycle, clonality, leaf traits, seed dispersal and seed mass. Of the 67 species for which data allowed analysis, 44 species declined after grassland abandonment but still occurred at the sites, probably as remnant populations. Five traits were associated with the response to abandonment. The declining but still occurring species were characterized by high plant height, a perennial life form, possession of a perennial bud bank, high clonal ability, and lack of dispersal attributes promoting long-distance dispersal. Traits allowing plants to maintain populations by utilizing only a part of their life cycle, such as clonal propagation, are most important for the capacity to develop remnant populations and delay local extinction. A considerable fraction of the species inhabiting semi-natural grasslands maintain what is most likely remnant populations after more than 60 years of spontaneous succession from managed semi-natural grasslands to forest. / <p>authorCount :3</p>

The structural integrity of fire-dependent ecosystems, such as ponderosa pine (<i>Pinus ponderosa </i> Dougl.) and Interior Douglas-fir (<i>Pseudotsuga menziesii </i> (Mirb.) Franco) biogeoclimatic zones in Interior British Columbia (BC) is changing. The problems within these ecosystems include decreased rangeland area, reduced carrying capacity and loss of biodiversity due to tree encroachment and forest ingrowth caused mainly by fire suppression. The goal of this study was to determine the effect of burning and thinning on understory vegetation of grassland and forested sites. The burning experiment took place at Dew Drop (Tranquille Ecological Reserve) located 20 km northwest of Kamloops, BC. Thinning was done at two upper grassland sites near Cache Creek, BC; Coal Mine Pasture and Gladys Lake Pasture. Species evenness and values of the Shannon-Weiner Diversity Index (H) were reduced (13 and 27%, respectively) within three years following burning (P = 0.014 and P = 0.038, respectively). Burning reduced canopy cover of shrubs on grassland sites (P = 0.005) and it reduced graminoid cover on forest sites (P = 0.014) immediately after the treatment (1999) but both functional groups had recovered by 2002. Litter depth and total canopy cover of plants were reduced in grasslands and forests immediately following burning (1999) but litter depth and canopy cover had recovered by 2002. Litter cover and litter depth beneath the tree canopy were reduced by burning (P = 0.037 and P = 0.009, respectively). Trends in the data indicate forb standing crop increased and total understory standing crop increased following burning in the grassland compared to the control. Graminoid standing crop was reduced 47% by burning in the forests (P = 0.049). Thinning reduced species richness in the first (P = 0.033) and fourth (P = 0.030) years, and H in the first year (P = 0.037) following the thinning at Coal Mine Pasture. Trends in the data suggest understory standing crop increased at Coal Mine and Gladys Lake Pastures following thinning. At both locations, thinning reduced litter depth. Therefore, burning and thinning kills trees, reduces fuel loads, and increases standing crop of the understory.

Grassland bird populations in North America are in steady decline. Despite declining faster and more consistently than any other group of birds, grassland songbird populations are relatively understudied and little is known about factors driving breeding-site philopatry and dispersal. Landscape and habitat composition may influence fidelity of grassland songbirds to a breeding area. As predicted by the theory of the <i>Ideal Free Distribution</i>, high-quality sites are likely to have a higher percentage of return breeders than low-quality sites because higher quality sites should have more or better-quality resources birds need for improved fitness. Using stable hydrogen isotope (äD) analysis, I approximated minimum fidelity rates of two grassland songbirds to two landscape (grass- vs crop-dominated landscapes) and two habitat (native grass vs planted grass) types. I hypothesized that grassland songbirds would return more readily to higher quality sites. For Spragues Pipit (<i>Anthus spragueii</i>), a habitat specialist, this would mean returning more readily to native grass habitat in grass dominated landscapes. I expected no difference in return rate of Savannah Sparrow (<i>Passerculus sandwichensis</i>), a habitat generalist, to either habitat or landscape. However, I found that the proportion of non-returning breeders was not influenced by landscape or habitat for either species. Furthermore, I examined attributes (distance from capture point to nearest crop and to the nearest road, as well as the percentage of native grass, planted grass, water and woody vegetation within landscape and territory buffers around the capture point) of the landscape and territory of each individual to determine if specific landscape or territory characteristics influenced their return rate to a breeding area. Neither species showed an affinity or aversion to any of the landscape or territory characteristics considered.<p> At a larger scale, geographical position within the breeding range may influence dispersal rates of migratory songbirds. Given that environmental factors often change in a clinal manner, central populations presumably experience the most favorable environmental conditions and peripheral populations the least favorable. Therefore, geographically peripheral locations likely occur in ecologically marginal or stressful conditions, resulting in higher dispersal rates of migratory birds. I examined the differences in dispersal rates of two grassland songbirds at two geographically distinct locations; one centrally located in the Spragues Pipit breeding range (Last Mountain Lake area, SK, Canada) and one at the periphery (Bowdoin National Wildlife Refuge, MT, USA). As expected, Spragues Pipits at the Bowdoin National Wildlife Refuge had a significantly larger dispersal rate than at the Last Mountain Lake area. Savannah Sparrow dispersal rates did not differ between locations at the more conservative outlier classification. These results provide some limited evidence that geographical position within the breeding range can influence dispersal rates.<p> Using äD analysis, I found that local amount-weighted growing-season deuterium in precipitation (äDp) at locations within grassland ecosystems differed from those long-term (45+ year) models described by stable hydrogen isotope ratio basemaps, illustrated in Hobson and Wassenaar (1997), Meehan et al. (2004) and Bowen et al. (2005). Therefore, I describe how äDp values were corrected from the long-term isoscape value predicted by Bowen et al. (2005). This method of determining year-specific local weighted growing-season äDp is an improvement upon the currently used Bowen et al. (2005) isoscape that is based on long-term precipitation patterns. To improve assignment of individuals to origins based on their äDf values, future research should incorporate year-to-year variation by applying year-specific corrections to the Bowen et al. (2005) isoscape.<p> More research is needed to determine the factors affecting the philopatry and dispersal of grassland songbirds in order to conserve them.