European forest ecosystems are changing drastically. The accelerating speed of current climate warming, further paired with drought events, storms and insect calamities, already results in reduced fitness, changes in forest health, species composition and productivity. Forest management is committed to develop suitable adaptation strategies to maintain the multitude of ecosystem services. In this context, the choice of seed origin for forest conversion and reforestation represents a key question to their success. Provenance trials, that test a wide range of provenances planted across multiple test sites, provide the urgently required evidence of population specific response to variable environmental conditions and may be used to derive recommendations under future conditions. This thesis concentrates on two economically and ecologically important native tree species, European beech and Norway spruce, as well as one of the most common introduced broadleaf species, Northern red oak. It provides a comprehensive analysis of their adaptive population differentiation and ability of phenotypically plastic response. For beech, 85 range-wide provenances were observed for 25 years in western, northern and eastern Germany. Considerable provenance differentiation exists, explaining 21% (stem form) to 45% (basal area) of the variance at individual sites and 20 to 39% across sites, while interactions between provenance and environment were absent. At the landscape level, spatially neighboring provenances showed similar trait expressions. These patterns were similar for height and basal area, but different for stem form. They could be directly linked to geographical variables using a multivariate regression tree analysis, which captured 58% of the phenotypic variation, whereby eleven ‘ecotypes’ characterized by local adaptation were delineated. A selection by two multi-trait indices gave highly concordant results and further underlined the trade-off between growth and stem form. Overall, the intermediate elevations of the central highlands in western Germany host highly suitable provenances. Lower elevation provenances from the southeast adapted to continental climate with harsh and cold winters, but warm and dry summers profit from the transfer to the favorable growing conditions in Germany and represent potential candidates for assisted migration. For red oak, each six provenances sourced from German stands and from North America were examined at age 33 on sites in northern, eastern and central Germany. In contrast to beech, provenance by environment interactions were significant and the thorough investigation of their nature provided highly consistent results. German provenances were mostly superior. This was particularly prevalent at the Atlantic site in Schleswig-Holstein, where the most outstanding provenance yields a basal area sum per hectare of +130% compared to the mean across six American provenances. Four of six German provenances were categorized as productive generalists, being suitable for planting across environments, while the other two are productive specialists for humid sites. Provenances from the native range had a lower performance, despite for the driest environment in eastern Germany, where one Canadian provenance sourced north of the Great Lakes emerged as a suitable specialist. Provenances from higher elevations in Georgia and Tennessee grew poorly and had low survival rates, especially in Schleswig-Holstein. Growth performance and survival of spruce was investigated based on the most extensive provenance trial series worldwide, established to test 1,100 provenances in 13 European countries. The analysis of 33 German sites of 100 provenances each, summarized to ten aggregated provenance groups, revealed relatively consistent ranks between groups up to age 51. Provenances from the Hercyno-Carpathian domain had superior short-term and positive long-term growth, while those towards the northern and southern range limits performed poorly. To develop a universal response function, the data set was extended to a total of 97 sites with 100 provenances each. Increasing summer heat paired with decreasing water availability was identified as the main climatic variable that restricts growth, while a prolonged frost-free period enables a longer period of active growth and therefore increasing growth potential within the restricted, remaining area. Clear signals of local adaptation to climatic clines spanning the entire range are barely detectable, as they are disguised by a latitudinal cline. This cline strongly reflects population differentiation for the Baltic domain, but fails to capture the high phenotypic variation associated to the geographic heterogeneity in the Central European Mountain ranges paired with the species history of postglacial migration. Finally, growth predictions of the trait-based model were compared to the predictions of a classic occurrence-based species distribution model. Both models showed a substantial retreat towards the northern latitudes and higher elevations (-55% and -43% by the 2080s). However, thanks to the species’ particularly high phenotypic plasticity the decline is delayed. Model-based prescriptions for assisted migration may not decrease the predicted range decline for Norway spruce, but may help to capitalize on potential opportunities associated with warming climate in the remaining area.:1 General introduction 1.1 European forests under climate pressure 1.2 The capacity of natural populations to cope with changing conditions and its limits 1.3 Strategies of adaptive forest management 1.4 Provenance trials to study population response 1.5 The theory of local optimality and its significance for seed deployment 1.6 Thesis outline and objectives 1.7 Species of interest and experimental data basis 1.8 The analytical tool set 1.9 Author contributions 2 Ecotypic variation in multiple traits of European beech: selection of suitable provenances based on performance and stability 2.1 Abstract 2.2 Introduction 2.3 Material and Methods 2.3.1 Experimental design 2.3.2 Measurement of traits 2.3.3 Quantification of provenance and site effects 2.3.4 Spatial interpolation of trait variation 2.3.5 Multi-variate grouping of provenances 2.3.6 Calculation of multi-trait indices for provenance selection 2.4 Results 2.4.1 Differentiation among provenances at individual sites 2.4.2 Provenance by environment interactions 2.4.3 Spatial patterns in provenance performance 2.4.4 Selection of suitable provenances 2.5 Discussion 2.5.1 Absence of provenance by environment interaction 2.5.2 Ecotypic variation across species range 2.5.3 Trade-off between growth and stem form 2.5.4 Concluding remarks 2.6 Supplementary information 3 Provenances from introduced stands of Northern Red Oak (Quercus rubra L.) outperform those from the natural distribution 3.1 Abstract 3.2 Introduction 3.3 Material and methods 3.3.1 Provenances and experimental design 3.3.2 Measurements 3.3.3 Data analysis 3.4 Results 3.4.1 Percent gain in productivity 3.4.2 Provenance-environment interaction 3.4.3 Generalists vs. specialists 3.4.4 Differences in stem form 3.5 Discussion 3.5.1 Superiority of German provenances 3.5.2 Suitable P×E approaches for provenance trials 3.5.3 Use of P×E to improve provenance recommendations 3.6 Conclusion 3.7 Supplementary information 4 The IUFRO Norway spruce provenance test from 1964/68: a first conclusive evaluation of the German trial sites based on provenance groups 4.1 Abstract 4.2 Zusammenfassung 4.3 Einleitung 4.4 Material und Methoden 4.5 Ergebnisse 4.6 Diskussion 4.7 Schlussfolgerung und Ausblick 5 High phenotypic plasticity, but low signals of local adaptation to climate in a large-scale transplant experiment of Picea abies (L.) Karst. in Europe 81 5.1 Abstract 5.2 Introduction 5.3 Material and Methods 5.3.1 Common gardens 5.3.2 National forest inventories 5.3.3 Climatic and geographic data 5.3.4 OccurrenceSDM 5.3.5 TraitSDM 5.3.6 Species range change 5.4 Results 5.4.1 Habitat projections by OccurrenceSDM 5.4.2 Growth projections by TraitSDM 5.4.3 Comparison of future species range 5.5 Discussion 5.5.1 Phenotypic plasticity largely determines height growth 5.5.2 OccurrenceSDM underestimates species range 5.5.3 Limitations of the TraitSDM 5.5.4 Implications for forest management 5.6 Supplementary information 6 General discussion and synthesis 6.1 Geographic variation within species 6.2 Plasticity as a main driver of juvenile growth 6.3 Absence vs. presence of provenance by environment interactions 6.4 Evidence-based seed transfer 6.5 Implications for assisted migration 6.6 Outlook Cumulative Bibliography

Wood products play a key role in the transformation to a more sustainable society based on renewable bio-based resources, together with the positive effects on climate mitigation by replacing fossil fuels. However, to increase the use of forest fuel in practice it is important to understand the effects of handling and storage on its quality and removal of nutrients from the forest. This thesis addresses these effects with special focus on a comparative evaluation of the traditional dried-stacked with “new” and to some extent more controversial fresh-stacked methods for extraction of logging residues from Norway spruce (Picea abies (L.) Karst). The results indicate that a normal extraction of logging residues will leave at least 20% of logging residues at the clear-felled area, in accordance with Swedish Forest Agency recommendations. However, the results also indicate that the ambition of the dried-stacked method to leave the majority of the needles well spread over the clear-felled area does not meet these recommendations. In fact, the harvesting operation is more important than the extraction method, with respect to how much logging residues (nutrients) being left in the forest. The results also show that the quality of fuel yielded by the two handling methods differs only to minor extent, indicating that other factors have stronger effects, where “dried-stacked” and “fresh-stacked” logging residues from different clear-felling areas is often similar. Generally, logging residues stored over summer (regardless method), seem to provide sufficiently dry forest fuel, with a needle content of about 5–10%. There is a clear correlation between drying and effective loss of needles from twigs, but the loss does not necessarily mean that the needles will remain in the forest.  However, needle color (green or brown) is not a strong indicator for a reduction in needle content. Acceptance of the fresh-stacked method would provide opportunities for the development of new technologies, more efficient use of machinery throughout the whole year, reduced costs, shorter lead times and increased amounts of logging residues extracted from each clear-felled area. This is mainly because it would enable extraction at optimal times from a logistical, financial and/or forestry perspectives. Written in English with summary and conclusion in Swedish. / Skogen och dess produkter har en nyckelroll i omställningen till ett framtida hållbart samhälle eftersom användande av träråvara har en positiv effekt på klimatet, t.ex. genom träbyggande eller att fossila bränslen ersätts med bioenergi. Men för att kunna ta tillvara mer skogsbränsle i form av grenar och toppar (grot) från bestånd dominerade av gran (Picea abies (L.) Karst) är det viktigt att förstå hur hanteringen påverkar såväl bränslekvalitet som bortförsel av näringsämnen från skogen. Denna avhandling belyser hanteringens effekter på bränslekvalitet och näringsförluster genom att jämföra den traditionella metoden (skotning av torkad grot, ”brunrisskotning”), med den ”nya” och till viss del ifrågasatta skotningen av färsk grot (”grönrisskotning”). Resultatet visar att ett normalt grot-uttag kommer att lämna minst 20 % av groten kvar på hygget, helt i linje med Skogsstyrelsens rekommendationer. Däremot uppfylls inte rekommendationerna med avseende på att lämna merparten av barren väl spridda på hygget, detta trots att det är ambitionen med den traditionella hyggestorkningen av grot. Hur mycket grot (och således näringsämne) som lämnas kvar på hygget påverkas i själva verket mer av hur avverkningen utförs, än när groten skotas ihop. Resultatet visar också en skillnad i bränslekvalitet mellan de båda beskrivna hanteringsmetoderna. Denna skillnad är dock så liten att andra faktorer sannolikt påverkar mer än hanteringsmetoden. Hyggestorkad grot från ett enskilt hygge i södra Sverige, kan i verkligheten ofta vara ganska likt färskskotad grot från ett annat hygge. Grot som har lagrats över sommaren kan alltså förväntas ha torkat tillräckligt, samt ha en barrandel på ca 5–10 % oavsett hanteringsmetod. Det verkar också finnas ett klart samband mellan torkning och att barren släpper från kvisten, men det betyder inte nödvändigtvis att barren blir kvar i skogen. Hur som helst, barrens färg (gröna eller bruna) är inte en rättvisande indikator på avbarrning. Om skotning av färsk grot accepteras ger det möjligheter för utveckling av ny teknik, effektivare användning av maskinresurser över hela året, minskade kostnader, kortare ledtider, samt möjligt ökat grot-uttag från enskilda hyggen. Detta beror främst på att det skulle vara möjligt att utföra grot-uttaget när det passar bäst ur ett logistiskt, ekonomiskt och/eller skogligt perspektiv. Skriven på engelska, men med sammanfattning och slutsatser på svenska / <p><strong>Tidigare handledare har också varit:</strong></p><p>Thomas Thörnqvist, Professor, Linnéuniversitetet</p><p>Dick Dandberg, Professor, Linnéuniversitetet</p><p>Åsa Blom, Docent, Linnéuniversitetet</p><p><strong>Sammanläggningsavhandlingen består av totalt fyra Paper:</strong></p><p>Paper I</p><p>Nilsson, B., Blom, Å., Thörnqvist, T. 2013. The influence of two different handling methods on the moisture content and composition of logging residues. <em>Biomass and Bioenergy</em>, 52, 34–42.</p><p>Paper II</p><p>Nilsson, B., Nilsson, D., Thörnqvist, T. 2015. Distributions and losses of logging residues at clear-felled areas during extraction for bioenergy: Comparing dried- and fresh-stacked method. <em>Forests</em>, 6, 4212–4227.</p><p>Paper III</p><p>Nilsson, D., Nilsson, B., Thörnqvist, T., Bergh, J. 2016. Amount of nutrients extracted and left behind at the clear-felled area using the fresh- and dried-stacked method of logging residue extraction. Submitted to <em>Scandinavian Journal of Forest Research </em>(2016-11-01).</p><p>Paper IV</p><p>Nilsson, B., Lerman, P. 2016. Experimental study of relations between defoliation, moisture content and color change in logging residues. Department of Forestry and Wood Technology, Linnaeus University, Växjö, Sweden. <em>Manuscript</em>.</p>

This thesis describes aspects of durability of Swedish softwood in above ground applications. The thesis consists of a summary of nine papers, which all aim to explain the existing variation in above ground microbial durability and moisture sensitivity of Norway spruce (Picea abies) and Scots pine (Pinus sylvestris). The methods used were two field tests, one accelerated durability test, and three water permeability tests. The main conclusions made in the study were that Norway spruce sapwood is more susceptible to discolouring fungi than heartwood. With respect to Scots pine, the only important factor for its natural durability above ground was whether the samples consisted of heartwood or of sapwood. There was also no systematic variation between pine heartwood from different stand origins in above ground conditions. Origin had no influence on durability in practice for neither spruce nor pine. Furthermore, in contradiction with traditional thinking, annual ring width and density had no influence on durability in any of the tests. The material came from two different samplings. The ‘old’ material was sampled in the beginning of the 1980s and consisted of Scots pine and Norway spruce from three different origins: the north, the central area, and the south of Sweden. The material was subjected to different handling conditions and surface and end-grain treatments. The ‘old’ material was used in a large above ground field test, carried out between 1985 and 1994. Unfortunately, the spruce sapwood and heartwood was not separated in this field test. In the field trial with the ‘old’ material, the most important factor for the durability of Norway spruce was the samples’ surface- and end-grain treatment. It was also observed that untreated spruce showed better durability than samples that were painted but without proper end-grain treatment. Samples originating from one of the stands were more sensitive towards air-drying than samples from the other two, which was reflected in a higher average moisture content and mass loss for the untreated samples. With respect to Scots pine, the most important factor was whether the sample consisted of heartwood or of sapwood. The heartwood samples were durable irrespective of their previous treatment. The Scots pine sapwood samples, on the other hand, had a very fluctuating moisture content if they were not surface- and end-painted. This was also reflected in higher mass losses in these samples. Even when properly surface- and end grain treated, the sapwood samples did not perform as well as the heartwood samples. The ‘new’ material consisted of Scots pine taken from six different stands, and Norway spruce from five different stands, all from areas in southern Sweden. The sampling was performed in order to achieve a large variation in wood properties. Thus, logs from areas with different climate and growth conditions were collected. This material was used for laboratory tests, and also for the second field test, which was evaluated after a test period of two years. In the second field test untreated Scots pine and Norway spruce samples from the ‘new’ material were investigated. Effects of origin and different tree diameters were examined. Furthermore, Scots pine was separated to heartwood and sapwood, and Norway spruce to mature and juvenile wood. The Norway spruce samples were sawn with vertical or horizontal annual rings. For the Norway spruce in the second field test, vertical annual rings were shown to be very beneficial for avoiding crack formation, while samples with horizontal annual rings displayed a large number of cracks. Despite this, the larger number of cracks did not result in a higher moisture content or more fungal discoloration. Juvenile wood had a slightly higher average discolouring fungal growth grading than the other spruce samples. For Scots pine, the only factor of importance was whether the samples consisted of heartwood or of sapwood. Annual ring width, density and origin had no significant effect on either fungal growth or moisture uptake. This observation held for both Norway spruce samples and Scots pine samples. An accelerated test was run in order to investigate Norway spruce sapwood and heartwood with respect to their differences in durability towards discolouring fungi and moisture uptake. The Mycologg method was used to accelerate fungal growth during a number of forced moisture cycles. The results showed that sapwood was much more sensitive to discolouring fungi than heartwood. Sapwood also recorded a higher average moisture content than heartwood. The differences that were observed between the types of samples were not due to annual ring width, density or origin. It was observed that Norway spruce sapwood samples and heartwood samples showed differences in liquid water permeability. This observation was made especially evident in a droplet absorption test. Water droplets were absorbed much faster on the sapwood samples in comparison with the heartwood samples. This higher affinity to water suggested that a sapwood surface would reach a higher moisture content than a heartwood surface. This observation provides a major explanation of why the sapwood samples showed poorer durability towards discolouring fungi in the Mycologg trials.

Pine weevils cause major damage to newly planted conifer seedlings in reforestation areas. However, recent findings indicate that small (“mini”) seedlings, planted at the age of 7-10 weeks, are gnawed less by pine weevils than the larger, conventionally planted seedlings. Thus, it has been proposed that planting young conifer seedlings in clear-cut areas may reduce the damage caused by pine weevils. In attempts to determine why mini seedlings appear to be less damaged by pine weevils than “conventional” seedlings, the volatiles released by Norway spruce and Scots pine mini seedlings were investigated, since such chemicals are of great importance in herbivore-plant communication, inter alia acting as repellents, attractants or antifeedants. Volatiles from the seedlings were collected, separated and identified by solid phase microextraction (SPME) followed by gas chromatography-mass spectrometry. The results show that there are high levels of chemodiversity among both spruce and pine seedlings. Between-tissue and age-related variations in their emissions were also found. Norway spruce clones infested by mites were also examined to assess genotype- and pest-specific stress reactions of Norway spruce. Finally, the effects of certain spruce defense compounds on the behavior of the large pine weevil Hylobius abietis were examined. / QC 20100818

Wood is a major construction material that is used in many contexts, and for different purposes. Serious problems may arise, however, when moisture related deformations as twist occur in wood used in different types of building structures, joinery and furniture. Twist can be explained to a great degree by the helical deviation of the grain angle in relation to the longitudinal direction of the log or the sawn board. Wood fibres form a spiral within the tree, and this is a natural occurrence that is named spiral grain. The wood fibres close to the pith in Norway spruce form a left-handed spiral. In most trees the grain angle turns over to be right-handed with time. Sawn timber that exhibits large grain angles lead to problems of shape stability and stiffness in finished constructions. In this thesis the spiral grain in Norway spruce (Picea abies (L.) Karst.) was stated as well as the effect on sawn timber. The material was based on sample trees from Sweden and Finland. Samples were taken in twenty-two stands at different heights in tree. From six stands studs were sawn and dried for measuring twist and other deformations. The spiral grain was measured with the method scribe test on 390 log discs taken at the top-end of the logs. Account was given concerning changes in grain angle from pith to bark, regarding both increasing annual ring numbers and distance from pith. The development of grain angle over tree age was utilized to study whether annual growth, size of tree, height in tree as well as silvicultural treatments affected spiral grain. Moreover, the relation between grain angle and distance to pith (in mm) was used to forecast twist in sawn timber. The left-handed grain angle was at its greatest between the fourth and eighth annual rings. Thereafter for most trees the grain angle turned from left-handed to right-handed in a linear fashion, in a manner that was unique for each individual tree. The pattern of spiral grain differed significantly between different stands, regarding change of inclination with increasing age or distance from pith. The culmination of the grain angle close to the pith occurred at somewhat higher age higher up in the trunk. The grain angle decreased faster in top logs than it did in the butt logs. The largest trees within a stand had a grain angle that turned to right in a slower way than smaller ones. The thinning strength and type of thinning regime also affected the character of spiral grain in the remaining trees in a stand. There was an indication that strong thinnings, where fast growing trees are retained, may lead to more individuals in a stand that exhibit high grain angles under bark. With knowledge of the size and direction of the grain angle under bark, and the diameter of the log, calculations can be made that show how twisted the sawn timber will be after drying. This can be used for deciding whether an individual log can profitably be sawn and processed further or not. The grain angle under bark can be used to remove trees showing the greatest degree of spiral grain already in the first thinning. Silvicultural methods aiming at even and dense Norway spruce stands, which normally is practised in Scandinavia, will probably result in timber with relatively low risk concerning large grain angle and subsequent risk for twist in sawn wood.

From studies of the circadian clock in the plant model species Arabidopsis (Arabidopsis thaliana), a number of important properties and components have emerged. These include the genes CIRCADIAN CLOCK ASSOCIATED 1 (CCA1), GIGANTEA (GI), ZEITLUPE (ZTL) and TIMING OF CAB EXPRESSION 1 (TOC1 also known as PSEUDO-RESPONSE REGULATOR 1 (PRR1)) that via gene expression feedback loops participate in the circadian clock. Here, we present results from ectopic expression of four Norway spruce (Picea abies) putative homologs (PaCCA1, PaGI, PaZTL and PaPRR1) in Arabidopsis, their flowering time, circadian period length, red light response phenotypes and their effect on endogenous clock genes were assessed. For PaCCA1-ox and PaZTL-ox the results were consistent with Arabidopsis lines overexpressing the corresponding Arabidopsis genes. For PaGI consistent results were obtained when expressed in the gi2 mutant, while PaGI and PaPRR1 expressed in wild type did not display the expected phenotypes. These results suggest that protein function of PaCCA1, PaGI and PaZTL are at least partlyconserved compared to Arabidopsis homologs, however further studies are needed to reveal the protein function of PaPRR1. Our data suggest that components of thethree-loop network typical of the circadian clock in angiosperms were present beforethe split of gymnosperms and angiosperms.

This thesis presents a number of studies, where a multivariate approach was taken to construct models that predict wood species and thermo mechanical pulp brightness from roundwood of Norway spruce and Scots pine. The first and second studies produced multivariate prediction models for wood species from the bark of spruce and pine. These models can be used for wood species classification and would replace the manual log assessment that takes place today. Principal Component Analysis, PCA, and Partial least squares projections to Latent Structures, PLS, were used to predict the wood species from multivariate measurements recorded from the bark of spruce and pine. Two different kinds of measurements were employed, near-infrared spectroscopy and digital imaging. Both methods showed that it was possible to predict the wood species with a high accuracy. The third and fourth studies of the thesis are related to the wood storage of roundwood and the deterioration of wood that occurs during the storage. The third study used an experimental design with five storage factors that provided different conditions for the analysed wood. The experimental design made it possible to identify the factors and the interaction between factors, which were important for the ISO brightness of peroxide and dithionite bleached thermo mechanical pulp, TMP. The final study of the thesis used NIR spectroscopy for predicting the ISO brightness of bleached TMP. Spectra recorded from stored wood were used to construct PLS prediction models.

Growth responses to climatic variables of red pine (Pinus resinosa Aiton) and Norway spruce (Picea abies L. Karst) were studied at the Morgan Arboretum, near Montreal, in southern Quebec, Canada (45$ sp circ$ 25$ sp prime$ N, 73$ sp circ$ 57$ sp prime$ W; 15.2 m above sea level). The relationships between climatic variables and basal area growth were examined using linear and quadratic models. Current and previous year's climatic variables were tested separately and in combination using multiple regression models. The best models explained 82% and 85% of the total variance of the growth of Norway spruce and red pine, respectively. The growth of both species was more associated with evapotranspiration than precipitation. The growth of Norway spruce was best explained by the current year's annual evapotranspiration (43%), while the growth of red pine was more related to previous year's August evapotranspiration (33%) at our site. / The JABOWA model was used to predict tree growth in hypothetical climates which could result from global climate changes. Based on literature, five treatments were applied: normal, and increases of 1, 3, 5, and 10$ sp circ$C. Comparison between the last (1983-1992) and next (1993-2002) ten years growth showed no significant differences between species when temperature was normal or increased by 1 and 3$ sp circ$C, but significant differences between species were observed when the temperature was increased by 5$ sp circ$C. Both species declined when the temperature was increased by 10$ sp circ$C. We concluded that Norway spruce is more sensitive to increases in atmospheric temperatures than red pine at our site.

Resultat från 3 olika bestånd i Västra Götaland. Åldern på bestånden är 30, 50 och 50 år. Varje provyta var 100 m2. Bestånden har vuxit utan gallring och gödsling och skötts enligt en metod som kallas Hyssnametoden. Syftet med studien var att beskriva, följa upp och utvärdera effekterna av en alternativ skogsskötselmetod. Resultatet av mätningarna visar att volymproduktionen är högre än vad som förväntas i det aktuella området. / Results from three stands of Norway spruce in Vaestra Goetaland in Sweden. The age of the stands are 30, 50 and 50 years old. Each sample area was 100 m2. The stands have grown without any thinning and fertilizer. The maintenance method is called “Hyssnametoden”. The purpose of the study was to describe, follow-up and evaluate the impact of an alternative silvicultural method. The result shows that volume growth is higher than expected in this field.

Det svenska skogsbruket har med fokus på beståndsvis avverkning och återbeskogning genererat ett mosaikartat landskap med en mängd väl avgränsande kantzoner mellan yngre och äldre bestånd, och att uppkomna förändringar hos växtmiljön i anslutning till dessa beståndsgränser kan påverka såväl tillväxt som artsammansättning har konstaterats i ett flertal tidigare studier. Men även om dessa studier påvisat förekomsten av kanteffekter har resultaten varierat utifrån geografiskt undersökningsområde och studerade trädslag. Tidigare studier har utifrån ett nordiskt perspektiv dessutom främst fokuserat på tillväxtförändringar hos tall i Finland och norra Sverige, vilket föranlett att denna studie valt att undersöka förekomst och omfattning av tillväxteffekter inom kantzoner belägna mellan angränsande äldre och yngre granbestånd i västra Götaland. Studien har genomförts genom att inom ett antal representativa objekt samla in data utifrån utlagda provyor i terrängen och undersökningen har avgränsats till att omfatta trädens höjd- och diametertillväxt samt markvegetationens artsammansättning. Vidare jämförelser har därefter gjorts utifrån avstånd från aktuell beståndsgräns, kantzonernas väderstreck i nordlig respektive sydlig riktning samt de yngre beståndens skötselhistorik i form av röjt eller oröjt. För att undersöka förekomsten av signifikanta skillnader mellan grupperna har insamlad data analyserats med hjälp av variansanalys (ANOVA) samt Tukey’s test för skillnader mellan medelvärden. Resultatet visade på signifikanta kanteffekter hos yngre granbestånd där tillväxtnedsättningarna var som tydligast närmast beståndsgränsen, oavsett väderstreck och tidigare skötselhistorik. Dessa tillväxtnedsättningar var dock mindre än de som tidigare konstaterats hos tall, och till skillnad från tidigare studier kunde inga tillväxteffekter konstateras hos angränsande äldre bestånd. Inga tydliga skillnader kunde heller konstateras utifrån kantzonernas väderstreck eller de yngre beståndens skötselhistorik. Markvegetationens artsammansättning uppvisade i sin tur en ökad förekomst av lavar och bärris intill beståndsgränserna. Bärris, lavar och ljung förekommer frekvent inom sydligt exponerade kantzoner medan smal- och bredbladigt gräs i sin tur uppvisade en ökad förekomst inom nordligt exponerade kantzoner. Utifrån dessa resultat skulle en tänkbar rekommendation inför kommande skogsbruksåtgärder och återbeskogning kunna vara att utifrån konstaterade tillväxtnedsättningar ej prioritera plantering av gran inom ett avstånd av 0-2,5 m från beståndsgräns angränsande mot äldre granbestånd. Därmed skulle kantzonen kunna nyttjas för naturvårdande ändamål då dessa områden gynnar den biologiska mångfalden och utgör intressanta miljöer för arter som annars skulle riskera att bli utkonkurrerade i angränsande mer slutna granskogsbestånd. / The aim of this study was to evaluate edge effects along forest edges between young and old Norway spruce stands in south-western Sweden, and examine its influence on tree growth and field vegetation. Comparison between distance from stand edge, exposure (north or south facing) and young forest clearing (cleared or not cleared) showed a reduced growth among young trees near the stand edge, mainly at a distance of 0-2,5 m. This edge effect occurred regardless of exposure or forest clearing history. On the other hand, in contrary to earlier studies, no positive growth effect was found in older stands adjacent to younger stands. Regarding edge effects found on field vegetation, blueberries, lingonberries and lichens showed an increased incidence adjacent to stand edges. Berries, lichens and heather showed an increased incidence at south facing edges while grass showed an increased incidence at north facing edges.