Está demonstrado em plantas atuais que níveis atmosféricos de CO2 e frequência estomática estão inversamente correlacionados. A utilização de dados estomáticos na calibração das curvas de CO2 paleoatmosférico do Fanerozoico tem sido eficiente não apenas para o Cenozoico, mas também apresenta excelente correspondência desde o Eodevoniano. Cutículas de glossopterídeas provenientes das jazidas de carvão de Faxinal (RS) e Figueira (PR), incluídas em níveis estratigráficos distintos da Formação Rio Bonito (Sakmariano e Artinskiano respectivamente), viabilizaram análises para floras desenvolvidas sob a vigência de megaciclo icehouse. Cabe ressaltar que o uso de glossopterídeas, classificadas dentro do grupo extinto das Pteridospermophyta, esbarra na impossibilidade de se estabelecer um equivalente ecológico atual, dada a ausência de relações taxonômicas, estruturais e ecológicas com grupos viventes. Portanto, optou-se por uma abordagem proposta por outros autores como alternativa para diferentes grupos de pteridospermas, que consistiu em comparar entre si resultados de análises estomáticas na morfo-espécie Glossopteris communis Feistmantel procedente de distintos níveis associados a carvões na Bacia do Paraná. O objetivo geral do presente estudo foi estabelecer possíveis relações entre os padrões estomáticos calculados e flutuações nos teores de CO2 paleoatmosférico. A metodologia consistiu em resgatar mecanica e quimicamente as cutículas da rocha matriz, clareá-las em solução de Schulze e montá-las em lâminas de gelatina glicerinada para observação em microscopia de luz transmitida. A observação deu-se com auxílio de filtro de contraste por interferência diferencial e as contagens foram feitas com o auxílio de programa para análise de imagens. As técnicas de estudo compreenderam cálculos de densidade estomática (DE- número de estômatos por unidade de área da folha) e índice estomático (IE- percentual de estômatos sobre o total de células epidérmicas). Os resultados obtidos foram de DE média= 234,73 e IE médio= 15,7 para a jazida de Faxinal (Sakmariano) e de DE média= 284,14 e IE= 18,9 para a jazida de Figueira (Artinskiano). Esses valores enquadram-se dentro da curva global de CO2 atmosférico para o Fanerozoico (modelo GEOCARB). As frequências estomáticas mais baixas das folhas do Faxinal com relação àquelas de Figueira foram relacionadas a processo de reversão temporária da tendência global de baixos teores de CO2 atmosférico para a base do Permiano. Essa reversão é atribuída à provável influência de fatores paleoecológicos locais relativos à grande extensão das turfeiras na parte sul da bacia, responsável pela emissão em larga escala de gases-estufa. Além disso, a flora de Faxinal está preservada em uma camada de tonstein, registro de atividade vulcânica que poderia ter afetado os níveis de CO2. Por outro lado, as turfeiras de registro muito esparso ocorrentes no nordeste da bacia, em intervalo mais jovem, por sua pequena extensão e ausência de indícios de vulcanismo, não alteraram o padrão paleoatmosférico. Estudos focados no final do Paleozoico têm especial relevância porque, nesse intervalo vigoravam, nas diferentes paleolatitudes, condições ambientais análogas àquelas ocorrentes na atualidade, como a existência de calotas de gelo nos pólos e períodos de aquecimento global. / In extant plants stomatal frequency and the concentration of the atmospheric CO2 have been shown to correlate inversely. The use of stomatal data to calibrate phanerozoic paleoatmospheric CO2 curves has been considered a reliable technique not only for the Cenozoic estimates but also for results obtained since the Early Devonian. Glossopterid cuticles from Faxinal and Figueira coalfields (respectively in Rio Grande do Sul and Paraná States) from distinct stratigraphic levels in the Rio Bonito Formation allowed stomatal counting for floras developed under icehouse megacycle. However, it is important to highlight that the efficiency of Glossopteris, classified in the extinct group of Pteridospermophyta, is restricted as paleo-CO2 proxy for the Paleozoic due to the difficulty of establishing a nearest living equivalent for it, given the lack of taxonomic, structural and ecological relationships with extant groups. Thus it was here adopted an alternative approach proposed by different authors analyzing other groups of pteridosperms, which consists in comparing results obtained from the morphospecies Glossopteris communis Feistmantel between two different coal levels in the Paraná Basin. The general aim of this study was to establish possible relations between the calculated stomatal patterns and the fluctuation in the paleoatmospheric CO2 levels. The methodology consisted in mechanically and chemically isolating the cuticles from the matrix rock, bleaching them with Schulze solution and then mounting glycerin jelly slides for observation in transmitted light microscopy. The microscopic observation was made using a differential interference contrast filter and the counting was carried out with the help of software for image analysis. The study techniques included the calculation of stomatal densities (SD- number of stomata per foliar area unit) and of stomatal indices (SI- a ratio of the number of the stomata to the total number of epidermal cells). The results were mean SD= 234.73 and mean SI= 15.7 in Faxinal coalfield (Sakmarian) and mean SD= 284.14 and SI= 18.9 in Figueira coalfield (Artinskian). These values agree with the curve of global atmospheric CO2 for the Phanerozoic (GEOCARB model). The lower stomatal frequencies detected at the climax of the coal interval (Faxinal coalfield, Sakmarian) when compared to the higher ones obtained in leaves from a younger interval (Figueira coalfield, Artinskian) could be attributed to temporarily high levels of atmospheric CO2. Therefore, the occurrence of an extensive peat generating event at the southern part of the basin and the consequent greenhouse gases emissions from this environment may have been enough to reverse regionally and temporarily the reduction trend in atmospheric CO2. Additionally, the Faxinal flora is preserved in a tonstein layer, which is a record of a volcanic activity that could also imply a rise in atmospheric CO2. During the Artinskian, the scarce generation of peat mires, as revealed by the occurrence of thin and discontinuous coal layers, and the lack of volcanism evidences would be insufficient to affect the general low CO2 trend. Studies focused in the Late Paleozoic are especially relevant because of the presently shared icehouse climate with glacial-interglacial cyclicity which includes times of global warming.

Doctorat en philosophie et lettres / info:eu-repo/semantics/nonPublished

For two centuries, questions about the origin of terrestrial plants and their impacts on the Earth systemhave occupied palaeobotanists. This essay attempts to synthesise the state of research to date, and outlineareas where major questions remain. Fossil evidence for land plants first appears in rocks of MiddleOrdovician age (~ 470 Ma), but it was not until the Devonian that vascular plants became the dominantgeobiological agents on the continents. Plants began as small organisms, lacking any vascular tissues,and essentially confined to wetlands. Key developments in their reproductive biology and the evolutionof mycorrhizal symbiosis subsequently enabled early plants to exploit a broader range of environments,enhancing water uptake and absorption of nutrients. In turn, the evolution of plant roots has significantlyimpacted terrestrial landscapes. The Devonian rise of plants led to a modification of the weathering rateand a sharp increase in the rate of mudrock production. This was driven by the impacts that plants exerted on watercourses, with the creation of meandering rivers and deltas that retained fine siliciclasticmaterials. This increase in weathering rate, combined with the development of leaves and the intensification of photosynthesis also had consequences for the carbon cycle and atmosphere, reducing the levelof CO2 and increasing that of O2 in the atmosphere. The increased proportion of oxygen and creation ofcombustible material is also thought to have led to the planets first wildfires, whilst the decrease in CO2lowered global temperatures. Via a complex set of feedbacks, these modifications may even have drivena series of anoxic events in the oceans, generating one of the five major mass extinctions at the end ofthe Devonian. / Paleobotanik är studien om fossiliserade växter. Då växter ursprungligen först saknades från jordens ytahar deras utveckling och kolonisering av kontinenterna (för 450 miljoner år sedan) haft många effekter.Ursprungligen var de första landväxter små och liknande modernt gräs. Deras små rötter begränsadedem till närvaron av vattenkällor (hav, flod, träsk…). En symbios med svampen (kallad mykorrhizalsymbios) underlättade landväxtens upptag av näring samt utvecklingen av djupare rötter, vilket ocksåunderlättade deras upptag av vatten. Då landväxter nu kunde migrera till torrare landskap har antaletlandväxter ökat med tiden. Det växande antalet landväxter var viktigt för bildningen av exempelvis kol,vilket är slutprodukten av dött växtmaterial som begravts och utsatts för höga tryck och temperaturförhållanden. Den geologiska perioden Karbon (359–299 miljoner år sedan) namngavs tack vare av denenorma andelen kol som kan dateras tillbaka till denna tid. Genom fotosyntes kan växter även absorbera CO2 och frigöra syre, och på så vis förändra atmosfärenssammansättning. På grund av högre syrehalter i atmosfären blev bränder ett allt vanligare fenomen eftersom det försågs med växter och syre som bränsle, som i sin tur även förbättrade bevarandet av frönoch växtvävnader i form av träkol. Oavsett om det är växtens frigöring av syre, minskning av CO2 (som är en växthusgas) eller genombränder, har uppkomst av landväxter haft en stor inverkan på atmosfärens sammansättning, och därföräven klimatet. På grund av en hastig nedgång av temperatur och syrehalt i haven mellan 450–375 miljoner år sedan (från 40 till 25 °C), utrotades nästan 75% av allt djurliv. Upphovet av denna massutrotning, genom försämringen av vattenkvalité, och uppkomsten av syrefritt vattenmiljöer tros bero på näringsläckage av jordar orsakat av ett ökat tillstånd av alger som spreds i samband med landväxters rotutveckling.

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Ce projet porte sur la reconstitution paléo-écologique d'un environnement forestier fossile retrouvé sur un plateau au sud-ouest de l'île Bylot. Il a comme objectifs 1) de préciser la chrono-stratigraphie du site; 2) d’établir une liste et une succession des différents taxons polliniques retrouvés dans les différentes unités stratigraphiques du site; 3) d’estimer leur âge et 4) d’en inférer des conditions climatiques (température et précipitations). Plusieurs coupes stratigraphiques ont été excavées puis échantillonnées afin de réaliser des analyses stratigraphiques, paléomagnétiques et polliniques. Un GPS différentiel fut également utilisé afin de caractériser à petite et grande échelle les unités stratigraphiques associées aux unités organiques fossiles. Les résultats des analyses granulométriques indiquent la séquence de dépôt suivante au sein d’une dépression dans la roche en place (schiste tertiaire) : 1) un diamicton glaciaire local ; 2) un sédiment limoneux d’origine glacio-lacustre ; 3) une unité organique tourbeuse; 4) une unité de type alluvial ; 5) un sédiment fluvio-glaciaire et 6) un diamicton glaciaire d’origine allochtone. Les analyses polliniques suggèrent une végétation similaire à celle présente près de la limite des arbres actuelle, environ 2000 km plus au sud. Les conditions climatiques plus humides et plus chaudes permettaient notamment la croissance du pin (Pinus type strobus et banksiana), de l’épinette (Picea cf. mariana), de l’aulne (type crispa et incana) et du mélèze (Larix, indifférencié). Enfin, les études paléomagnétiques et la présence d’espèces éteintes suggèrent un âge pour les dépôts organiques fossiles situé entre 2,14 et 2,15 Ma ou entre 2,581 et 3,040 Ma. / The project focuses on the paleo-ecological reconstitution of a fossil forest environment found on a plateau on the southwest part of Bylot Island, in the Canadian Arctic. The objectives were 1) to describe the chrono-stratigraphy of the site; 2) to establish a list and the succession of different pollen taxa found in organic and inorganic fossil units of the site; 3) to estimate their age and 4) to infer the past climatic conditions of the site. Several stratigraphic cuts were excavated and sampled for stratigraphic, paleomagnetic and pollen analyses. Also, a differential GPS was used to characterise the stratigraphic units associated with the organic fossil units at a small and large scale. Results of the granulometric analysis suggest the following deposition sequence starting from a depression in the bedrock (tertiary shale): 1) a glacial diamicton with local clasts; 2) a silty sediment of glacio-lacustrine origin; 3) an organic peat unit; 4) an alluvial type unit; 5) a fluvio-glacial sediment and 6) a glacial diamicton of allochtone origin. The pollen analyses suggest vegetation similar to that of the current tree-line limit, situated about 2000 km south of the site. The climatic conditions were warmer and more humid, allowing the local growth of pines (Pinus type strobus and banksiana), spruce (Picea cf. mariana), alder (Alnus type crispa and incana) and larch (Larix, undifferentiated). Paleomagnetic analyses and the presence of extinct species finally suggest an age for the organic fossil deposits of between 2,14 and 2,15 My or most likely between 2,581 and 3,040 My.

Ce projet porte sur la reconstitution paléo-écologique d'un environnement forestier fossile retrouvé sur un plateau au sud-ouest de l'île Bylot. Il a comme objectifs 1) de préciser la chrono-stratigraphie du site; 2) d’établir une liste et une succession des différents taxons polliniques retrouvés dans les différentes unités stratigraphiques du site; 3) d’estimer leur âge et 4) d’en inférer des conditions climatiques (température et précipitations). Plusieurs coupes stratigraphiques ont été excavées puis échantillonnées afin de réaliser des analyses stratigraphiques, paléomagnétiques et polliniques. Un GPS différentiel fut également utilisé afin de caractériser à petite et grande échelle les unités stratigraphiques associées aux unités organiques fossiles. Les résultats des analyses granulométriques indiquent la séquence de dépôt suivante au sein d’une dépression dans la roche en place (schiste tertiaire) : 1) un diamicton glaciaire local ; 2) un sédiment limoneux d’origine glacio-lacustre ; 3) une unité organique tourbeuse; 4) une unité de type alluvial ; 5) un sédiment fluvio-glaciaire et 6) un diamicton glaciaire d’origine allochtone. Les analyses polliniques suggèrent une végétation similaire à celle présente près de la limite des arbres actuelle, environ 2000 km plus au sud. Les conditions climatiques plus humides et plus chaudes permettaient notamment la croissance du pin (Pinus type strobus et banksiana), de l’épinette (Picea cf. mariana), de l’aulne (type crispa et incana) et du mélèze (Larix, indifférencié). Enfin, les études paléomagnétiques et la présence d’espèces éteintes suggèrent un âge pour les dépôts organiques fossiles situé entre 2,14 et 2,15 Ma ou entre 2,581 et 3,040 Ma. / The project focuses on the paleo-ecological reconstitution of a fossil forest environment found on a plateau on the southwest part of Bylot Island, in the Canadian Arctic. The objectives were 1) to describe the chrono-stratigraphy of the site; 2) to establish a list and the succession of different pollen taxa found in organic and inorganic fossil units of the site; 3) to estimate their age and 4) to infer the past climatic conditions of the site. Several stratigraphic cuts were excavated and sampled for stratigraphic, paleomagnetic and pollen analyses. Also, a differential GPS was used to characterise the stratigraphic units associated with the organic fossil units at a small and large scale. Results of the granulometric analysis suggest the following deposition sequence starting from a depression in the bedrock (tertiary shale): 1) a glacial diamicton with local clasts; 2) a silty sediment of glacio-lacustrine origin; 3) an organic peat unit; 4) an alluvial type unit; 5) a fluvio-glacial sediment and 6) a glacial diamicton of allochtone origin. The pollen analyses suggest vegetation similar to that of the current tree-line limit, situated about 2000 km south of the site. The climatic conditions were warmer and more humid, allowing the local growth of pines (Pinus type strobus and banksiana), spruce (Picea cf. mariana), alder (Alnus type crispa and incana) and larch (Larix, undifferentiated). Paleomagnetic analyses and the presence of extinct species finally suggest an age for the organic fossil deposits of between 2,14 and 2,15 My or most likely between 2,581 and 3,040 My.

Extreme haustorial parasites have long captured the interest of naturalists and scientists with their greatly reduced and highly specialized morphology. Along with the reduction or loss of photosynthesis, the plastid genome often decays as photosynthetic genes are released from selective constraint. This makes it challenging to use traditional plastid genes for parasitic plant phylogenetics, and has driven the search for alternative phylogenetic and molecular evolutionary markers. Thus, evolutionary studies, such as molecular clock-based age estimates, are not yet available for all parasitic lineages. In the present study, we extracted 14 nuclear single copy genes (nSCG) from Illumina transcriptome data from one of the “strangest plants in the world”, Hydnora visseri (Hydnoraceae). A ~15,000 character molecular dataset, based on all three genomic compartments, shows the utility of nSCG for reconstructing phylogenetic relationships in parasitic lineages. A relaxed molecular clock approach with the same multi-locus dataset, revealed an ancient age of ~91 MYA for Hydnoraceae. We then estimated the stem ages of all independently originated parasitic angiosperm lineages using a published dataset, which also revealed a Cretaceous origin for Balanophoraceae, Cynomoriaceae and Apodanthaceae. With the exception of Santalales, older parasite lineages tend to be more specialized with respect to trophic level and have lower species diversity. We thus propose the “temporal specialization hypothesis” (TSH) implementing multiple independent specialization processes over time during parasitic angiosperm evolution.

Two riparian habitats in southeastern Arizona provide the setting for a study of 127 plants useful to human foragers. A view of plant part availability is based on annual phenological profiles, and on historic and prehistoric records of plant use. Food choice is limited in March and April, but high August through November. Riparian plants also offer numerous non-food resources. Trees and shrubs serve more needs in relation to number of available species than do perennial herbs (including grasses) and annuals. Southwestern ethnographic literature hints that certain native taxa (Panicum, Physalis, Populus, Salix, Typha and Vitis) might receive special care. Inherent qualities of parts, coupled with ethnographic records of preparation and use, provide a basis for speculation on which parts might survive in an ancient record. Most are expected to disintegrate in open sites. Parts sought for different needs can enter a dwelling via diverse routes that produce confusingly similar archaeological debris. Modern experiments to wash pollen from 14 separate harvests permit evaluation of plant fruit and leaves as pollen traps, to help interpret pollen recovered from ancient dwellings. High amounts of Berberis, Rumex and Ribes pollen, sometimes in clumps or as tetrads, travel on harvested fruit. Arctostaphylos, Monarda, Oxalis, Rhus, Rhamnus, Vitis and Juniperus parts carry lower amounts. Quercus and Gramineae pollen grains travel on parts of other taxa, as well as on their own fruit. The phenological profiles offer insight into group life-form activities in response to local temperature and precipitation trends. Rising and maximum temperatures coincide with intense vegetative and reproductive activity for trees, shrubs, herbaceous perennials, and annuals. Increased levels of precipitation coincide with maximum flowering and fruiting of herbaceous perennials and fall annuals. Limited data on six taxa from Utah generally agrees with observations in this study, suggesting strong genetic control in the phenology of some riparian taxa.

To assist in developing a more precise understanding of past climatic changes in southern Africa, further pollen analytical research is required. In the past, pollen sites in the subregion have been restricted to swampy areas such as permanent springs and peat deposits. While such sites are often rare as a consequence of the aridity of the country, rich polliniferous deposits can be found in the peatlands surrounding coastal lakes in the Maputaland Coastal Plain. The Mfabeni peatland, situated on the eastern shores of St. Lucia, contains relatively old sediments dating back to >45000 years bp at a depth of 7.80m. A multi-proxy approach, comprising radiocarbon, stable carbon isotope (513C) and palynological analysis, was applied in the investigation of Late Quaternary climatic conditions and vegetation changes along the Maputaland Coastal Plain. A single 10 m sediment core, dating back to >45000 years bp, was extracted from the Mfabeni Peatland. A detailed fossil pollen analysis of Mfabeni sediments indicated the existence of extensive Podocarpus-abundant coastal forests before ca. 44500 years bp. The onset of wetter local conditions after this time is inferred from forest retreat and the development of swampy conditions, which prevailed until ca. 25000 Cal years BP. Conditions during the Last Glacial Maximum (LGM; 18000 years BP) are inferred to have been generally colder and drier, as evidenced by forest retreat and replacement of swampy reed/sedge communities by dry grassland. A significant depletion in 813C values at ca. 18200 Cal years BP indicates the dominance of C3 vegetation during the LGM, reflecting considerably colder conditions. This is in agreement with palaeoenvironmental indications from elsewhere in the Transvaalian Ecozone, although conditions at Mfabeni were more moderated in their manifestation, which can be attributed to the proximity of this site to the ocean. Cool, relatively moist conditions are inferred for the Holocene Altithermal (ca. 8000-6000 years BP), as evidenced by forest growth and expansion during this time. Warm, dry conditions are inferred for the Late Holocene, with the establishment of grassland/savanna type vegetation in the area after ca. 2000 Cal years BP. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2005.