Student Number : 0115350V - MSc dissertation - School of Plant, Animal and Environmental Science - Faculty of Science / The effect of between-tree competition in separate even aged stands of a Eucalyptus grandis seedling and a single Eucalyptus grandis x Eucalyptus urophylla hybrid clone was assessed. This was done by comparing treatments within an experiment, that was designed to test the effect planting density and genotype had on timber production and stand dynamics through time. This experiment is located on an escarpment in the Summer rainfall region of South Africa, approximately 7 km North West of Albert Falls dam in the Kwa-Zulu Natal Midlands. The terrain is flat with deep apedal soils, and the site has an estimated mean annual precipitation of 1143 mm. The simplest method of evaluating competition effects in plantation stands is through the use of spacing trials. Spacing trials create stands/plots in which all silvicultural and climatic variables are constant. Differences between treatments can then be attributed to competition alone. The planting densities tested were 6667 stems.ha-1, 3333 stems.ha-1, 2222 stems.ha-1, 1587 stems.ha-1 and 1111 stems.ha-1 respectively. Competition was measured at three levels, namely; (i)individual tree, (ii)size-class and (iii)stand level. Vigour and its distribution within the population were used to explain competition processes. Basal area was chosen as the size variable for which vigour would be analysed. Relative Growth Rate (RGR) and Relative Production Rate (RPR) were chosen as the measures of vigour for analysis. Morphological changes in the trees as a result of competition were also used to understand the competitive mechanisms. It was found that resources are not shared amongst individuals in proportion to their size, but rather the larger individuals use disproportionately more of the resources than the smaller individuals, and therefore competition is asymmetrical. The dominant form of competition was found to be resource pre-emption where larger individuals passively prevent solar resources from reaching smaller competitors. Using RPR as the measure of vigour, competition was found to be one-sided, meaning that smaller individuals do not reduce the vigour of larger ones. If RGR is accepted as the measure of vigour then competition is two-sided meaning that the smaller individuals do reduce the vigour of the larger ones to a certain extent. There was no difference in the degree to which the smaller competitors in the clonal and seedling treatments reduced the growth of the larger individuals. The self-thinning rate in the seedling treatments was significantly higher than in the clonal treatments. Increases in mean tree size and production per hectare were accompanied by increases in mortality in the seedling treatments. In the clonal treatments it was found that even once the mean tree size approached the self thinning line, very few or no trees died, however there was also very little or no increase in mean tree size. When the relationship between number of live trees and production per hectare was analysed it was found that the clone could increase production per hectare without any change in survivorship, which is contrary to popular theory. As time progressed the skewness of the population progressed from negative to positive. This increase was the most pronounced in the higher planting density treatments, and the clonal treatments. The size variability in the population first decreased, then steadily increased. This increase was more pronounced in the high planting density treatments, and the clonal treatments. Clones and seedlings react in very different ways to competition. The large differences in population dynamics observed, indicate that we cannot just apply knowledge built up over the years regarding the management of Eucalyptus grandis and other plantation species to Eucalyptus hybrid clones.

Coordenação de Aperfeiçoamento de Pessoal de Nível Superior / This work aimed at building a management dendrogram and determining the appropriate density for management of stands of Pinus elliottii in central region of the Rio Grande do Sul. To do that, stands in maximum stock, in different ages, located in the campus of Universidade Federal de Santa Maria and neighborhoods, were used. The dendrometric variables used in the calculation process were obtained from sample units with temporary characteristic, considering the method of determination of the punctual density proposed by Spurr, and isolated trees, with crowns in free growth. The maximum population density was estimated by the Reineke method, with an angular coefficient of -1,92009. The density in number of trees for the beginning of the total occupation of the dossal was estimated taking into account the crown areas, which were obtained from a lineal model estimator of crown diameter in function of the dap, fit for variables from trees in free growth. The construction of the dendrogram was in function of the basal area, the number of trees per hectare and the tree diameter with medium basal area. The density levels were determined proportionally to the maximum density for stands density index (IDP), in clusters of index 200, from the maximum IDP, 1200, to the minimum, 400. The appropriate density for management was found by calculation process and adjusted by the observation of two stands. Results indicate the maintenance of the density between IDP 600 and 840 for the production of wood of great dimensions. Plans of management with different initial densities and number of self-thinning interventions were elaborated, showing that the dendrogram is an efficient tool, able to assist the elaboration of the management plan, with production estimates for self-thinning and final cut. / Este trabalho teve como objetivos construir um dendrograma de manejo e determinar a densidade adequada para manejo de povoamentos de Pinus elliottii para a região Central do Rio Grande do Sul. Para tanto, foram utilizados povoamentos em estoque máximo, em diferentes idades, localizados no campus da Universidade Federal de Santa Maria e proximidades. As variáveis dendrométricas foram obtidas de unidades amostrais de caráter temporário pelo método de determinação da densidade pontual proposto por Spurr, e de árvores isoladas, com copa em crescimento livre. A máxima densidade populacional foi estimada pelo modelo de Reineke, com um coeficiente angular de -1,92009. A densidade em número de árvores para o início da ocupação total do dossel foi obtida através de um modelo linear estimador de diâmetro de copas em função do dap, ajustado com variáveis obtidas de árvores em crescimento livre. O dendrograma foi construído em função da área basal, do número de árvores por hectare e do diâmetro da árvore de área basal média. Os níveis de densidade foram determinados, proporcionalmente, à máxima densidade por índice de densidade do povoamento (IDP), em classes de índice 200, desde o IDP máximo, 1200, até o mínimo, 400. A densidade adequada para manejo foi ajustada por processo de cálculo e pela observação de dois povoamentos. Os resultados obtidos indicam a manutenção da densidade entre o IDP 600 e 840 para a produção de madeira de grandes dimensões. Foram elaborados planos de manejo com diferentes densidades iniciais e número de intervenções de desbaste, demonstrando que o dendrograma é uma ferramenta eficiente, capaz de auxiliar na elaboração de planos de manejo, com estimativas de produção para desbastes e corte final.

Maximum size-density relationships (MSDR) are used to quantify differences across sites in the number of trees of a given size and species that can be supported per hectare. These relationships are important to managers who are trying to maximize basal area and wood volume. In my study, I examined MSDR across Alabama, Georgia, Louisiana, and Mississippi using US Forest Service, Forest Inventory and Analysis (FIA) data. I determined the impact of species-specific, specific gravity, functional traits, and environmental factors on MSDR using a quantile regression approach. Overall, I found that climatic factors had the greatest influence on MSDR, and that species shade and drought tolerance were more influential than specific gravity across the southeastern US.

Self-thinning quantifies the reduction in tree numbers due to density-dependent mortality. Maximum size-density relationships (MSDRs) are a component of self-thinning that describe the maximum tree density per unit area obtainable for a given average tree size, often quadratic mean diameter (D). An MSDR species boundary line has been defined as a static upper limit of maximum tree density -- D relationships that applies to all stands of a certain species within a particular geographical area. MSDR dynamic thinning lines have been defined as the maximum tree density obtainable within an individual stand for a particular D which have been shown to vary relative to planting density. Results from this study show that differences in boundary levels of individual stands cause the MSDR species boundary line slope estimate to be sensitive to the range of planting densities within the model fitting dataset. Thus, a second MSDR species boundary line was defined whose slope is the average slope of all MSDR dynamic thinning lines. Mixed-models are presented as a statistical method to obtain an estimate of the population average MSDR dynamic thinning line slope. A common problem when modeling self-thinning is to determine what observations are within generally accepted stages of stand development. Segmented regression is presented as a statistical and less subjective method to determine what observations are within various stages of stand development. Estimates of D and trees per acre (N) where MSDR dynamic thinning lines begin and end on the logarithmic scale were used as response variables and predicted as a function of planting density. Predictions of MSDR dynamic thinning line beginning and ending D and N are used in an alternative MSDR dynamic thinning line slope estimation method. These models show that the maximum value of Reineke's Stand Density Index (SDI) varies relative to planting density. By relating planting density specific Zone of Imminent Competition Mortality boundaries to a MSDR species boundary line, self-thinning was found not to begin at a constant relative SDI. Thus, planting density specific Density Management Diagrams (DMD) showed that self-thinning began at 40 to 72% for planting densities of 605 and 2722 seedlings per acre, respectively. / Ph. D.

Coordenação de Aperfeiçoamento de Pessoal de Nível Superior / Density management diagrams (DMD) is a tool used to carry out the prognosis of production in density, basal area and volume in order to establish management strategies for forest planning. The aim was to build a density management diagram for Acacia mearnsii De Wild. Data from 12 cities in the northeastern region of Rio Grande do Sul were used for the DMD construction. Since the Tang model had the best evaluation, it was used to select the maximum population density estimative, thus the density 1x1 was used to construction of the DMD. With the same model, self-thinning graphs and Stand Density Index (SDI) were generated; adopting a standard quadratic mean diameter of 10cm. Density levels were determined proportionally to SDI maximum density in 200 class intervals, with higher 2400 and lower 1400 SDI. The DMD were produced with an average diameter and density control, and volume and green rind bark weight per hectare. A second DMD model was developed in order to insert a site production control through the dominant height of the stand. These DMD models allowed estimating the volume and green rind bark with great precision, from the control of diameter, density and dominant height. / O Diagrama de Manejo da Densidade (DMD) é uma ferramenta utilizada para realizar a prognose da produção em densidade, área basal e volume a fim de estabelecer estratégias de manejo para o planejamento florestal. O objetivo do presente foi construir um diagrama de manejo da densidade para Acacia mearnsii De Wild., com dados dendrométricos provenientes de 12 municípios da região nordeste do Rio Grande do Sul. A máxima densidade populacional foi estimada pelo modelo de Tang, ajustado com dados de parcelas em espaçamento 1x1m. Com este modelo foram gerados os gráficos de autodesbaste e Índices de Densidade de Povoamento (IDP) para um diâmetro padrão de 10cm. Os níveis de densidade foram determinados, proporcionalmente, a partir da máxima densidade, por índice de densidade de povoamento (IDP), em intervalos de classe de 200, com IDP máximo 2400 e mínimo 1400. Os DMDs foram elaborados com controle de diâmetro médio e densidade populacional, e como variável de resposta o volume por hectare e o peso de casca verde por hectare. Um segundo modelo de DMD foi elaborado com o objetivo de inserir um controle produtivo dos sítios, através da altura dominante do povoamento. Estes modelos de DMD permitiram estimar o volume e peso de casca verde com ótima precisão, a partir do controle do desenvolvimento do diâmetro, densidade e altura dominante.

Conselho Nacional de Desenvolvimento Científico e Tecnológico / The goal of this study is to develop Density Management Diagrams for Eucalyptus grandis W. Hill, which will serve as auxiliary models to establish the variation of appropriate density, and the management group, to conduct settlements with the economic evaluation of management employees schemes. To study area were used stands in full density in reduced spacing, located in Rio-grandense Eastern center and Porto Alegre metropolitan in Rio Grande do Su states mesoregions. Of the density models tested, Tang showed good accuracy, with adjusted coefficient of determination of 0,74 and 3,17% coefficient of variation, which served as the base model for determining the population density rates, competition areas, construction of Management Density Diagrams and the size of the average diameter at the time that starts self-thinning for trees planted per hectare density. It was shown that the value of the slope calculated for the model of Tang, was different from that proposed by the author, worth -3/2 of self-thinning law. It was observed that the average diameter size at the time of the start self-thinning occurs with the removal of the population individuals varies considerably with the tree planting density. Preferably, you should lead the development trajectory of the density of trees per hectare below the line of maximum density, where the line of imminent mortality occurs. With reference to the standard diameter of 25 cm was obtained 5 Settlement density index curves describing curves between them proportional exponentially decreasing trends of frequency individuals per hectare with the diameter increment. Management density diagrams with the variables mean diameter, basal area, volume stand density index provided good volumetric estimates per hectare in relation to actual volumes with efficiency of 0,95, indicating good precision. The tested probability distributions, the Weibull function 2 parameters described accurately the frequency of diameters in time, and the re-estimation of its parameters, allowed frequency distributions by diameter class and therefore real development of the stand. The selected trunk shape function was to Hradetzky with determination coefficient of 0,98 and 8.43% coefficient of variation, generating through its integration the estimated volume of the whole stem and parts of it. The Net Present Value of R$ 12.372,35 ha-1 was determined for the management regime with three thinning, at 52,8; 85,4; 121,8 months and last cut at 185,1, higher than the other simulated systems. The Equivalent Annual Value was good and viable economically for all tested systems, the most attractive was the regime where there were two thinnings to 52,8; 85,4 months and last cut at 121,8 months with Equivalent Annual Value of R$1.550,78 ha-1. The internal rate of return was attractive in any of the simulated management regimes, ranging from 6,48% by year in stands without thinning and final cut at 52,8 months at 25,31% by year in stands managed with thinnings at 52,8; 85,4 months and last cut at 121,8 months. The biggest reason benefit on costs was 3.15, obtained in the management regime with two thinning, at 52,8; 85,4 months and last cut at 121,8 confirming the Equivalent Annual Value and Internal rate or return criteria. / O objetivo do presente estudo é elaborar Diagramas de Manejo da Densidade para Eucalyptus grandis W. Hill, que servirão de modelos auxiliares para estabelecer a variação de densidade adequada, e a faixa de manejo, para condução de povoamentos juntamente com a avaliação econômica dos regimes de manejo empregados. Para área de estudo foram utilizados povoamentos em densidade completa em espaçamentos reduzidos, localizados nas mesorregiões Centro Oriental Rio-grandense e Metropolitana de Porto Alegre do Estado do Rio Grande do Sul. Dos modelos de densidade testados, o de Tang apresentou boa precisão, com coeficiente de determinação ajustado de 0,74 e coeficiente de variação de 3,17%, que serviu de modelo base para determinação dos índices de densidade do povoamento, zonas de concorrência, construção dos Diagramas de Manejo da Densidade e a dimensão do diâmetro médio no momento que inicia o autodesbaste para uma densidade de árvores implantadas por hectare. Comprovou-se que o valor do coeficiente angular calculado para o modelo de Tang, foi diferente do proposto por este autor, com valor de -3/2 da lei de autodesbaste. Observou-se que a dimensão de diâmetro médio no momento em que ocorre o início do autodesbaste, com a supressão de indivíduos da população, varia, consideravelmente com a densidade de árvores de plantio. Preferencialmente, deve-se conduzir a trajetória de desenvolvimento da densidade de árvores por hectare logo abaixo a linha de máxima densidade, onde ocorre a linha de iminente mortalidade. Tendo como referência o diâmetro padrão de 25 cm obtiveram-se 5 curvas de Índices de Densidade do Povoamento que descreveram curvas proporcionais entre si com tendências de decréscimo exponencial das frequências de indivíduos por hectare com o incremento diamétrico. Os diagramas de manejo da densidade com as variáveis diâmetro médio, área basal, volume por índice de densidade do povoamento permitiram boas estimativas volumétricas por hectare em relação aos volumes reais com eficiência de 0,95, indicando boa precisão. Das distribuições de probabilidade testadas, a função de Weibull 2 parâmetros descreveu com acurácia a frequência dos diâmetros no tempo, e a reestimativa dos seus parâmetros, permitiu as distribuições de frequência por classe diamétrica e por conseguinte desenvolvimento real do povoamento. A função de forma de tronco selecionada foi a de Hradetzky com coeficiente de determinação ajustado de 0,98 e coeficiente de variação de 8,43%, gerando por meio de sua integração os volume estimados de todo o fuste e de partes dele. O Valor Presente Líquido de R$ 12.372,35 ha-1 foi determinado para o regime de manejo com três desbastes, aos 52,8; 85,4 e 121,8 meses e corte raso aos 185,1 meses, superior aos demais regimes simulados. O Valor Anual Equivalente foi positivo e viável economicamente para todos os regimes testados, o mais atrativo foi o regime onde ocorreram dois desbastes aos 52,8; 85,4 meses e corte raso aos 121,8 meses com Valor Anual Equivalente de R$1.550,78 ha-1. A taxa interna de retorno foi atrativa em qualquer dos regimes de manejo simulados, variando de 6,48 % a.a. em povoamentos sem desbaste e corte raso aos 52,8 meses a 25,31 % a.a. em povoamentos manejados com desbastes aos 52,8; 85,4 meses e corte raso aos 121,8 meses. A maior razão benefício sobre os custos foi de 3,15, obtida no regime de manejo com dois desbastes, aos 52,8; 85,4 meses e corte final aos 121,8 corroborando com os critérios Valor Anual Equivalente e Taxa Interna de Retorno.

Conselho Nacional de Desenvolvimento Científico e Tecnológico / This thesis aims at studying the efficiency of models which express the size-density relationship in even-aged stands of Pinus taeda L., handled in complete and full stock densities. For that, Reineke, Yoda, Zeide and Tang models were tested using data obtained from 50 permanent sample plots placed in stands implanted in spacing of 1.5x1.0 m, 2.5x1.0 m, 1.5x2.0 m, 2.0x2.0 m, 1.5x3.0 m, 2.5x2.0 m, 2.0x3.0 m, 2.5x3.0 m, measured every year until the age of 18 and maintained in full stocks. Results indicated that all of the size-density relationship models presented good statistical precision. However, the no-linear model of Tang was more efficient than the others, presenting excellent coefficient of determination (0.99), low standard error of estimate (0.0948), low coefficient of variation (1.17%), low bias (0.0086) and high efficiency (0,8976), in the estimate of the values of density of trees per hectare for average diameter, and the smallest considered value of the scores of the statistical criteria, equal to 6 points, with good distribution of residues of the number of trees per hectare. The self-tinning occurs in a dimension of average diameter directly proportional to the density of trees per hectare of planting. As bigger is the initial spacing, bigger is the average diameter in the moment of the beginning in the stand the self-thinning. The slope of the size-density and diameter equation varied per planting spacing, not allowing proving the universality of the self-tinning law, with value of this slope is equal -3/2. With help of the size-density model, 5 Indexes of Density of the Stand (IDS) were generated, with variation from the maximum 1400 to the minimum 600, with interval of class of IDS equal to 200, having as reference the standard diameter of 25 cm. The stand s Density Management Diagram (DMD), elaborated for the dendrometric variables average diameter, basal area and volume for hectare by IDS presented good efficiency with low mistake in the estimate of the values of these variables. Real volume and estimate volume per hectare in DMD presented an absolute difference of just -7.39 m3 ha-1 and a relative difference of -1.79 %, showing a great precision of the DMD model. The estimates volume of DMD per hectare, in relation to the real values, presented a value of efficiency equal to 0.99, what indicates a high precision, and a χ 2 value equal to 0.00034, not significant to 0.01% of probability, demonstrating that the real and the estimate values don't differ statistical amongst themselves. / A presente dissertação foi desenvolvida com o objetivo de estudar a eficiência dos modelos que expressam a relação de densidade e diâmetro em povoamentos equiâneos de Pinus taeda L., manejados em densidade completa e altamente estocados. Para isto, foram testados os modelos de Reineke, Yoda, Zeide e Tang, através de dados originados de 50 parcelas permanentes, medidas anualmente até os 18 anos, alocados em povoamentos implantadas em espaçamentos de 1,5x1,0 m, 2,5x1,0 m, 1,5x2,0 m, 2,0x2,0 m, 1,5x3,0 m, 2,5x2,0 m, 2,0x3,0 m, 2,5x3,0 m, e mantidas em densidade completa. Os resultados indicaram que todos os modelos de densidade e diâmetro apresentaram boa precisão estatística, porém o modelo não linear de Tang foi mais eficiente que os demais, tendo apresentado ótimo coeficiente de determinação, igual a 0,99, baixo erro padrão da estimativa igual a 0,0948, baixo coeficiente de variação, igual a 1,17 %, mínima tendência, com valor igual a 0,0086, e elevada eficiência, com valor igual a 0,8976. Na estimativa dos valores de densidade de árvores por hectare por diâmetro médio, e a menor soma de escores dos critérios estatísticos, foi igual a 6 pontos, com boa distribuição dos resíduos do número de árvores por hectare. O autodesbaste ocorre numa dimensão de diâmetro médio diretamente proporcional a densidade de árvores por hectare de plantio. Quanto maior o espaçamento inicial maior é o diâmetro médio no momento do início do autodesbaste da população. O coeficiente angular da equação de densidade e diâmetro variou por espaçamento de plantio, não permitindo comprovar a universalidade da lei do autodesbaste, cujo valor do coeficiente angular é estipulado em -3/2. Com o modelo de densidade e diâmetro foram gerados 5 Índices de Densidade do Povoamento - IDP, que variaram de um valor máximo de 1400 até o mínimo de 600, com intervalo de classe de IDP de 200, tendo com referência o diâmetro padrão de 25 cm. Os Diagramas de Manejo da Densidade - DMD da população, elaborados para as variáveis dendrométricas: diâmetro médio, área basal e volume por hectare, por IDP, apresentaram uma boa eficiência com baixo erro na estimativa dos valores destas variáveis. Os volumes reais por hectare e os estimados no DMD apresentaram uma diferença absoluta de apenas -7,39 m3ha-1 e relativa de -1,79 %, demonstrando elevada precisão do modelo de DMD. As estimativas volumétricas por hectare do DMD em relação aos valores reais, apresentaram valor de eficiência igual a 0,99, o que indica uma alta precisão, e um valor de χ 2 igual a 0,00034, não significativo a 0,01 % de probabilidade, demonstrando que os valores reais e os estimados não diferem estatisticamente entre si.

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Submitted by Automa??o e Estat?stica (sst@bczm.ufrn.br) on 2017-06-02T23:08:22Z No. of bitstreams: 1 JeanneRaquelDeAndradeFranco_DISSERT.pdf: 3675339 bytes, checksum: 97a2dabc11a0348c6acf6db2de294e5d (MD5) / Approved for entry into archive by Arlan Eloi Leite Silva (eloihistoriador@yahoo.com.br) on 2017-06-12T19:27:50Z (GMT) No. of bitstreams: 1 JeanneRaquelDeAndradeFranco_DISSERT.pdf: 3675339 bytes, checksum: 97a2dabc11a0348c6acf6db2de294e5d (MD5) / Made available in DSpace on 2017-06-12T19:27:50Z (GMT). No. of bitstreams: 1 JeanneRaquelDeAndradeFranco_DISSERT.pdf: 3675339 bytes, checksum: 97a2dabc11a0348c6acf6db2de294e5d (MD5) Previous issue date: 2017-02-14 / Coordena??o de Aperfei?oamento de Pessoal de N?vel Superior (CAPES) / Manguezais s?o ecossistemas costeiros e estuarinos que pertencem a climas quentes e temperados ou tropicais ?midos, s?o dominados por esp?cies lenhosas, fornecem importantes bens e servi?os ecossist?micos e possuem valor socioecon?mico. Os manguezais s?o sens?veis a dist?rbios e est?o em constante processo de degrada??o devido ? carcinicultura e ao desenvolvimento urbano. Intera??es entre indiv?duos vegetais podem resultar em competi??o e facilita??o interferindo no crescimento, no padr?o de distribui??o e na sobreviv?ncia das plantas, al?m de modificar caracter?sticas f?sico-qu?micas do ambiente. Vari?veis abi?ticas e bi?ticas interagem permitindo an?lise da adapta??o das esp?cies em diferentes condi??es ambientais, podendo tamb?m inferir sobre a intensidade de degrada??o. O objetivo geral do estudo foi identificar os principais fatores limitantes no crescimento e na sobreviv?ncia de plantas de Rhizophora mangle L. (Rhizophoraceae) em dois ambientes dominados pela esp?cie e localizados em ?rea de carcinicultura, sendo um com presen?a de ?rvores e copa fechada, e outro aberto com plantas jovens e subarbustivas. Foram realizados dois monitoramentos com sele??o de parcelas e plantas em cada ambiente, totalizando 90 indiv?duos, para quantificar o crescimento ao longo do tempo atrav?s de medidas da altura, n?mero de ramos e di?metro a 30 cm do solo. Para as vari?veis abi?ticas foram feitas medidas da temperatura e pH do solo, luminosidade, salinidade, velocidade do vento, temperatura do ar e penetrabilidade do solo nas parcelas de cada ambiente aberto e fechado por 12 meses. Indiv?duos adultos de R. mangle foram competidores interferindo no crescimento de plantas jovens da mesma esp?cie atrav?s da limita??o de luz e do estresse advindo do ac?mulo de sedimentos. Em ambiente aberto, com plantas jovens apresentando mesma idade e distribu?das de forma agregada, ocorreu facilita??o com redu??o de condi??es estressantes devido ? maior incid?ncia de radia??o solar e temperatura desse ambiente, e tamb?m competi??o atrav?s de alta densidade de plantas. Devido a essas intera??es positivas e negativas agindo simultaneamente, o efeito final da densidade-depend?ncia no crescimento de plantas do ambiente aberto foi neutro. Al?m disso, temperaturas m?dias de 30?C, pH do solo com valores entre 5,5 e 6,5 e salinidade com cerca de 35 ppm foram fatores que beneficiaram o crescimento das plantas ao longo do tempo em ambiente aberto atrav?s da fotoss?ntese, de adapta??es fisiol?gicas e poss?vel melhor assimila??o de nutrientes do solo. Esses resultados ajudaram a entender as caracter?sticas ambientais que facilitam e interferem na recupera??o de ?reas degradadas pela carcinicultura. / Mangroves are ecosystems coastal and estuarine belonging to warm and temperate climates and wet tropics, mangroves are dominated by woody species, provide important ecosystem goods and services and have socioeconomic value. Mangroves are sensitive to disturbances and are constantly being degraded due to shrimp farming and to urban development. Interactions between plant individuals may result in competition and facilitation by interfering on the plant growth, pattern of distribution and plant survival, besides modifying physical-chemical characteristics of the environment. Abiotic and biotic variables interact allowing the analysis of the adaptation of the species in different environmental conditions and also about the intensity of degradation. The main objective of the study was to identify the main limiting factors on the growth and survival of Rhizophora mangle L. (Rhizophoraceae) plants in two environments dominated by this specie and located in the shrimp farming area, one with trees and closed canopy, and another open with young and sub-shrub plants. Two monitoring were carried out with selection of plots and plants in each environment, totaling 90 individuals, to quantify the growth over time by means of measures of height, number of branches and diameter at 30 cm. For the abiotic variables, measurements of soil temperature and pH, luminosity, salinity, wind velocity, air temperature and soil penetrability were taken in the plots of each open and closed environment for 12 months. Adult individuals of R. mangle were competitors interfering in the growth of young plants of the same species through the limitation of light and of the stress coming from the accumulation of sediments. In the open environment, with young plants presenting the same age and distributed in an aggregated way, facilitation occurred with reduction of stressful conditions due to the higher incidence of solar radiation and temperature of this environment, and also competition through high density of plants. Due to these positive and negative interactions acting simultaneously, the final effect of density dependence on plant growth of the open environment was neutral. However, mean temperatures of 30?C, soil pH values between 5.5 and 6.5 and salinity with about 35 ppm were factors that benefited the growth of the plants over time in open environment through photosynthesis, of physiological adaptations and possible better assimilation of soil nutrients. These results helped to understand the environmental characteristics that facilitate and interfere the recovery of areas degraded by shrimp farming.

Conselho Nacional de Desenvolvimento Científico e Tecnológico / The objective of the work was to study the relationship between the density of trees per hectare and the average diameter to a Deciduous Forest, as well as adjust the model to describe this behavior Reineke. The study area is located in the town of Silveira Martins, in the central region of Rio Grande do Sul and is in early stages of succession after agriculture. The information about the number of the trees per hectare and the average diameter were obtained by the method of density-off proposed by Spurr. These plots were sampled in the early stages of a secondary forest, picking up areas where the predominant Camboatá-vermelho (Cupania vernalis). As natural forests have irregular spacing, density is highly variable, therefore, to select only high-density plots were chosen in areas that there was the occurrence of dead individuals. Were tested different methods to estimate the upper limit of the self thinning line: regression analysis (for all data and relative density greater than 60%), correcting the intercept so that the wastes were negative, the manual adjustment, the relative density (DR> 90%) and stochastic frontier analysis. The method that estimated the maximum density was regression analysis with data from at least 60% of maximum density, obtaining a slope of -1.563 for the model of Reineke. There was no significant difference between the powers provided by the different methods. The maximum Stand Density Index was 1779 trees per hectare, to a dg of 25 cm. The density management diagram was constructed on the basis of basal area, number of trees per hectare and diameter of the tree of average basal area. Were used the densities of 15% and 60%, to close the canopy, and the induction of mortality, respectively. The densities were determined proportionally to the maximum density by stand density index (PDI) for a reference diameter of 25 cm in different classes of index 200, since the IDP 1700, to a minimum of 300. Populations whose density is greater than 60% of the maximum were considered overstocked, between 60 and 15% fully stocked, and below 15% under stocked. They recommended a combination of Dendrogram generated in this study with the method of Spurr to guide interventions in the Deciduous Forest. / O objetivo deste trabalho foi estudar a relação entre a densidade de árvores por hectare e o diâmetro médio para uma Floresta Estacional Decidual, bem como, ajustar o modelo Reineke para descrever este comportamento. A área de estudo localiza-se no município de Silveira Martins, na região central do Rio Grande do Sul e encontra-se em estágio inicial de sucessão, após uso agrícola. As informações referentes ao número de árvores por hectare e o diâmetro médio foram obtidas por meio do método de densidade pontual proposto por Spurr. Estas parcelas foram amostradas nos estágios iniciais de uma floresta secundária, escolhendo-se áreas onde predominava o camboatá-vermelho (Cupania vernalis). Como as florestas naturais apresentam espaçamento irregular, a densidade é bastante variável, assim sendo, para selecionar apenas parcelas em alta densidade, foram escolhidas áreas em que se verificava a ocorrência de indivíduos mortos. Foram testados diferentes métodos para estimar o limite superior da linha de autodesbaste: a análise de regressão (para todos os dados e densidade relativa maior que 60%), corrigindo o intercepto para que os resíduos fossem negativos; o ajuste manual; o de densidade relativa (DR>90%) e a análise de fronteira estocástica. O método que melhor estimou a máxima densidade foi a análise de regressão com dados de no mínimo 60% da densidade máxima, obtendo um coeficiente angular de -1,563 para o modelo de Reineke. Não houve diferença significativa entre as potências fornecidas pelos diferentes métodos. O Índice de Densidade de Povoamento máximo foi de 1779 árvores por hectare, para o dg de 25 cm. O diagrama de manejo da densidade foi construído em função da área basal, do número de árvores por hectare e do diâmetro da árvore de área basal média. Foram utilizadas as densidades de 15% e 60%, para o fechamento das copas, e a indução da mortalidade, respectivamente. Os níveis de densidade foram determinados, proporcionalmente, à máxima densidade por índice de densidade do povoamento (IDP), para um diâmetro de referência de 25 cm, em classes de índice de 200, desde o IDP de 1700, até o mínimo de 300. Populações cuja densidade for maior que 60% da máxima foram consideradas superestocadas, entre 60 e 15% estocadas, e abaixo de 15% subestocadas. Recomendou-se a combinação do Dendrograma gerado neste estudo com o método de Spurr para guiar as intervenções na Floresta Estacional Decidual.