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Theoretical positions (Sloman & Gilbert, 2000), current research (Robbins & Tanck, 1997) and clinical observations (APA, 1994) have generally concluded that depressed populations tend to demonstrate an elevated level of hostility. Based on the premises of the Social Rank Theory (SRT; Sloman & Gilbert, 2000), the current study explores the purported etiological underpinnings of the co-occurrence between depression and hostility. The SRT regards depression as a state of inferiority resulting from a drop in social rank and hostility as stemming from a sense of injustice over this inferiority. To test this idea, measures of perceived social rank, depression, trait anger, anger expression and perceived injustice were administered to 97 university students at two time points, one month apart. Long-term rank change was measured retrospectively at Time 1 and short-term rank change was measured prospectively by sampling at Time 1 and Time 2. Three hypotheses were advanced: 1) social rank would be negatively associated with depression; 2) unfavourable rank change would predict greater levels of depression; and 3) unfavourable rank change from an initially superior rank would predict greater levels of anger and perceived injustice. Results were partially supportive of the hypotheses. As expected, social rank was negatively associated with depression. As well, a long-term change in social rank predicted greater levels of anger suppression. Results were discussed with respect to their consistency with the SRT. Potential weaknesses of the methodology and future directions of this line of inquiry were also presented.

Theoretical positions (Sloman & Gilbert, 2000), current research (Robbins & Tanck, 1997) and clinical observations (APA, 1994) have generally concluded that depressed populations tend to demonstrate an elevated level of hostility. Based on the premises of the Social Rank Theory (SRT; Sloman & Gilbert, 2000), the current study explores the purported etiological underpinnings of the co-occurrence between depression and hostility. The SRT regards depression as a state of inferiority resulting from a drop in social rank and hostility as stemming from a sense of injustice over this inferiority. To test this idea, measures of perceived social rank, depression, trait anger, anger expression and perceived injustice were administered to 97 university students at two time points, one month apart. Long-term rank change was measured retrospectively at Time 1 and short-term rank change was measured prospectively by sampling at Time 1 and Time 2. Three hypotheses were advanced: 1) social rank would be negatively associated with depression; 2) unfavourable rank change would predict greater levels of depression; and 3) unfavourable rank change from an initially superior rank would predict greater levels of anger and perceived injustice. Results were partially supportive of the hypotheses. As expected, social rank was negatively associated with depression. As well, a long-term change in social rank predicted greater levels of anger suppression. Results were discussed with respect to their consistency with the SRT. Potential weaknesses of the methodology and future directions of this line of inquiry were also presented.

Learning allows animals to adapt to new and changing environments. Animals can learn through their own personal experiences, known as asocial or individual learning. Asocial learning produces reliable information, but it can be energetically costly and risky for the learner. So instead of learning on their own, animals can choose to learn by observing and copying the behaviours and choices of others, known as social learning. However, because individuals that socially learn are gaining second-hand information, this form of learning is often less reliable. Animals are expected to be flexible in their use of individual versus social information, and to use whatever strategy provides the greatest benefits. Not all animals or individuals have been found to employ a flexible strategy, and research shows that many have a clear preference for one type of learning over the other. This preference for social or individual learning can be influenced by their personality, their sex and even an individual’s reproductive status. Another factor that could influence learning is an individual’s rank, but this topic has received little attention. In my M.SC. research, I studied how social rank influences an individual’s performance in an asocial and in a social learning task using the cichlid fish Neolamprologus pulcher. Using this species I also tested if social rank influences information use, by providing conflicting individual and social information to the fish. I found that subordinate N. pulcher were faster at a reversal learning (suppressing a previously learned rule and learning a new one), but there were no clear differences between the social ranks in associative learning or in social learning. When presented with conflicting individual and social information, both subordinate and dominant N. pulcher relied on individual information first. However, dominant N. pulcher were more likely to also use the conflicting social information in addition to their individual information. Taken together these results help us better understand cognitive differences between social ranks, and shed light on how information and behaviours in social groups can be learned and spread. / Thesis / Master of Science (MSc)

Problematic weight regulation as found in obesity and Anorexia Nervosa (AN) are chronic conditions which require long-term management. In order to develop long-term strategies to manage these conditions, a clearer understanding of the factors that can contribute to the development and also recovery from these conditions are a necessity. Although obesity and AN are at the opposite end of the bodyweight spectrum, some shared psychological processes may drive these states. One factor that has been suggested to contribute to problematic weight regulation is psychosocial stress whilst positive affect systems and affect regulation processes are important for regulating stress-related experiences. Gilbert (2005) describes an affect regulation system which consists of two positive affect systems known as social rank and attachment. Whilst the latter affect system refers to the attachment bond that develops between an infant and its caregiver (which extends to adult relationships), the former is used to form relationships that allow us to compete for limited resources and maintain our status in the social environment. Affect regulation processes in the current research are self-criticism and self-reassurance. Whilst self-critical thoughts and feelings can be triggered by perceptions of being low rank, the idea that people can be self-reassuring or being able to self-reassure at times of difficulty is nested in the positive infant-caregiver attachment bond and a consequence of internalizing parental soothing (Gilbert, 2006). Hence, as stress is suggested to be an important factor in problematic weight regulation and affect systems and processes are central to the regulation of emotional responses to stress-related experiences, the current series of studies examined these factors in relation to problematic weight regulation. The current research consisted of four studies designed to examine the role of stress and affect regulation in relation to weight change, weight regain following weight loss and recovery versus symptom maintenance in AN in women. A longitudinal study (Study One) was conducted to examine the change trajectories of stress, eating pathology and bodyweight, how these changes influence each other and the role of affect regulation systems and processes on these changes in a community based sample (N = 1157). Study Two examined the role of stress and affect regulation as predictors of weight regain in those who have lost weight (N = 42) and Study Three used a measure of life events and difficulties to investigate the role of stressful life changes and affect systems on recovery and relapse following AN (N = 30). Finally, in Study four, an expressive writing task which has been demonstrated to have a positive impact on stress-related health outcomes was used to explore the role of stress, affect systems and processes on problematic weight regulation and eating at times of stress (N = 57). The findings of the research studies demonstrated that there is a concurrent link between stress and the regulation of bodyweight and eating in a community-based sample of women. However, the proposed relationship between stress, bodyweight and eating behaviours was not confirmed when examined longitudinally in a community-based sample, over a 7-month period in women who have lost weight or when examined retrospectively as contributing to symptom maintenance in women with AN. However, the main finding of the current series of studies suggested that affect systems and affect regulation processes do have important implications for regulating stress-related experiences, bodyweight and eating behaviours. Perceived low social status, greater insecurity of attachment, more self-critical and less self-reassuring thoughts and feelings were related to increases in stress levels, higher bodyweight and higher levels of dysfunctional eating patterns. In addition, whilst expressive writing did not reduce stress, influence bodyweight or improve affect regulation at times of difficulty, writing about positive experiences had a positive impact on reducing dietary restraint behaviours during a stressful period. In conclusion, these findings suggest that it may not be stress per se that contributes to unhealthy changes in bodyweight and eating behaviours but how we use our affect systems and processes to manage our emotions at times of difficulty. Consequently, these findings have important implications for practice as weight loss programmes, Eating Disorder prevention programmes and stress management interventions should address the issues of perceived low social status, self-criticism and attachment insecurities.

Ma thèse porte sur une dimension fondamentale de la structure des groupes sociaux : la hiérarchie. Chez l'être humain, les hiérarchies sociales régissent en profondeur les interactions. Pour naviguer avec succès dans son environnement, il doit être en mesure de repérer précisément les positions hiérarchiques des autres membres de son groupe. Ce travail de thèse vise à caractériser certains mécanismes neuronaux, comportementaux et physiologiques impliqués dans l'analyse d'un indice hiérarchique. Pour préciser la nature du traitement de la hiérarchie, j'ai exploré son influence sur différentes étapes de la perception des visages. Je me suis tout d'abord intéressée au décours temporel du traitement neuronal des visages dans un contexte hiérarchique. Deux études menées en électroencéphalographie m'ont permis d'identifier les potentiels neuronaux et les composants oscillatoires évoqués par la perception de visages associés soit, à un rang hiérarchique établi à l'issue d'une compétition, soit à un statut social induit par la profession. Une étude réalisée ensuite en oculométrie avait pour but de capturer l'influence de la hiérarchie sur des mécanismes fins du contrôle de l'attention visuelle. J'ai étudié à la fois l'exploration visuelle de classements hiérarchiques incluant le participant, et celle de visages associés à des rangs hiérarchiques différents. Enfin, j'ai tenté de déterminer si un signal ou une situation d'asymétrie hiérarchique véhicule une valence émotionnelle et motivationnelle non neutre susceptible d'induire des variations de certains paramètres physiologiques, comme le rythme cardiaque ou la réponse électrodermale / Hierarchy is a key organizational feature of social groups. In order to successfully navigate their social environment, humans must precisely read the hierarchical position of other during social interaction. This present thesis intends to characterize the neural correlates as well as the early physiological and behavioral mechanisms involved in the processing of social rank. The influence of hierarchy was mainly investigated in the context of face perception. To begin, my focus was on the time course of neuronal processing of faces embedded in a hierarchical context. Using eletroencephalography in two studies, it has been possible to identify evoked neuronal potentials and oscillatory components in response to faces varying in hierarchical rank, established through competition or social status induced by profession. The next study used eye-tracking methodology to explore the influence of hierarchy on the subtle mechanisms of visual attention control. I aimed at characterizing the visual scanning pattern of hierarchical rankings (during a competition) and of faces associated with different hierarchical ranks. Finally, I tried to determine if a hierarchical signal or a social asymmetrical situation conveyed an emotional/motivational valence. During face perception and a minimal social interaction, I examined if this particular dimension of hierarchy generated variations of physiological activity, such as heart rate and skin conductance response

This is a linguistic study that will apply theories as a way of understanding the contexts of aspects of the play King Lear by William Shakespeare, as they relate to the possession, and exercise of power. It focuses on targeting and exploring the language of the play and how it impacts characters’ behaviour to gain or sustain power. To do this, specific theoretical frameworks have been applied, including semantics and pragmatics in the analysis of a passage. Examples from the opening scene of King Lear are displayed in order to answer three research questions. Among the findings are differences in the selection of nouns and pronouns with references to authority such as when females tend to overuse “I”, “love” and “lord” when conversing. This research discovered that semantic approaches therefore can be used to explain how Shakespeare portrays, for example, gender differences between the characters by his selection of words, metaphors, and metonymic expressions. Since Lear does not speak in the same manner as his later self as he would have done when at the heights of his power, his linguistic shift mirrors his shift in status following abdication. The analysis also draws certain conclusions with regard to implicatures that are derived from the use of vagueness and ambiguity as outlined in the field of pragmatics, including Speech Act Theory, Deixis and Grice´s Cooperative Principle. However, this essay argues that Grice´s Theory of Implicature and his Maxims can be insightful when analysing Shakespearean dramas, especially floutings and violations of the Maxim of Manner. By applying approaches from the fields of semantics and pragmatics this study concludes that the findings relate to Shakespearean works in general and other works from that period and genre.

Social dominance relationships are formed within numerous animal species and reduce costly fights over resources. Dominant individuals often enjoy greater access to important resources such as food and mating partners, and are generally more aggressive, bold, active and explorative compared to subdominant individuals. These behavioural traits can differ among individuals, but they can also be consistent within the individual, thereby describing the individual’s personality type. However, the causal direction of the observed correlation between dominance and personality is not well studied. One possibility is that some personality types have higher chances of obtaining a dominant social position. This would suggest that personality has consequences for fitness. Another possible explanation is that possessing different social positions gives rise to consistent behavioural differences among individuals on various timescales. If social status has a lasting effect on behaviour, social status would constitute a ‘stable state’ that explains some of the diversity of personality types that has been observed in a multitude of animal species. Dominance and personality may also share underlying proximate factors. In this thesis, I investigate the relationship between social dominance and personality using male domestic fowl, Gallus gallus domesticus. The species is group-living with pronounced dominance hierarchies, and dominance increases male access to mating partners. I show that some aspects of personality, exploration, vigilance and in particular aggressiveness, increased a male’s chances of obtaining dominance (paper III, IV, V), and that aggressiveness can be even more important than body weight and ornament size (comb size, paper V) or recent experience of winning or losing (paper IV). Winning a social interaction resulted in an increase in aggressiveness, while a decrease was seen in males that experienced a loss (paper IV). By observing behaviour before and after changes in male dominance relationships, I further show that a recent (2 days earlier) change in social status induced behavioural adjustments to the new social status in activity, exploration and vigilance (paper I). By extending the time of the new social relationship to 3 weeks, I show that such behavioural changes did not continue (paper II). Finally, I show that the social environment during juvenile development had little impact on adult male competitiveness (paper V). Social interactions appear to have several short-term effects on behaviour, but did not contribute significantly to variation and long-term consistency of personality in male fowl. Instead, a male's personality, and in particular his aggressiveness, affected the outcome of dominance interactions. Overall, my studies reveal important consequences of individual differences in behaviour, and contribute to the highly sought-after empirical testing of hypotheses explaining variation in animal personality. / <p>At the time of the doctoral defense, the following papers were unpublished and had a status as follows: Paper 2: Manuscript. Paper 5: Manuscript.</p>

Identifying the causes and consequences of intra-specific variation in cognitive abilities is fundamental to our understanding of the evolution of cognition. The social environment and cognitive abilities appear inextricably linked, yet evidence for how the social environment affects cognitive performances and further, how cognitive performances influence the social environment, has seldom been explored. Using the pheasant, Phasianus colchicus, I explore the relationships between individual variation in cognitive performances in relation to broad and fine-scale structure of the social environment and endeavour to separate cause and consequence. I demonstrate a positive causal effect of the broad-scale social environment on cognitive performances by observing increases in the accuracy of spatial discrimination performances when individuals are in larger groups (Chapter Two and Chapter Four). I show that the positive effects of larger group size occur over a relatively short period (less than one week), suggesting that cognitive performances are flexible in response to the social environment and I suggest four potential mechanisms. I show that while males are part of a social hierarchy, spatial discrimination performances are related to this fine-scale social structure and higher-ranking males outperform lower ranking males (Chapter Three). When attempting to determine cause and consequence, I found that spatial learning performances early in life did not predict adult cognitive performances on the same task or predict their adult social rank (Chapter Four). Hence, my results do not support that social rank is a consequence of spatial learning abilities in male pheasants. The relationship between spatial learning performances and social rank was found in adult males that had their social rank artificially elevated, suggesting that cognitive performances were not simply the result of the current social environment but remain closely related to past agonistic relationships. I did not find a relationship between early life aggression with performances on either a spatial or a non-spatial task in females or males (Chapter Five). This highlights the importance of investigating early life relationships and suggests that the relationship between spatial learning and aggression in adult males may become associated over time as a consequence of further spatial learning experiences, and, or, aggressive interactions. I then demonstrate a consequence of individual variation in cognitive abilities and show that adult foraging associations in the wild disassort by early life cognitive performances (Chapter Six). Individuals with good inhibitory control performance and poor visual discrimination performances were more central in social networks. I propose that differences in cognitive abilities manifest in foraging strategy and influence the resulting social structure. The implications of this predictable social structure remain to be explored. Finally, I discuss these results and how they contribute to our understanding of how the social environment causes individual differences in cognitive performances, as well as how variation in cognitive performances may shape the social environment. I suggest the potential implications of these findings and ideas for future work.

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