Graduation date: 2006 / Canopy gap formation is a major factor contributing to maintenance of overstory species diversity and stand structure in forests and may be integral to development of understory shrub and herb layers as well. Acknowledgement of gap formation as a fundamental feature of natural forests has led to consideration of gaps as an option in forest management regimes. This study examined understory vegetation communities across canopy gaps created as a part of the Density Management Study (DMS), which investigates the effectiveness of a thinning regime in promoting late-successional habitat development in young Douglas-fir forests of western Oregon. Patterns in understory vegetation community composition in and around 0.1 and 0.4ha gaps created as a part of the DMS treatment were investigated. The primary goal of this research was to investigate the potential role of canopy gap creation in fostering heterogeneity in understory vegetation communities, and to examine the extent of gap influence on the surrounding thinned forest matrix. Tree species distributions have been shown to partition across gaps in tropical forest systems through differential responses of species to gradients in resource availability, a pattern known as gap partitioning. In temperate forests, understory vegetation communities are much more diverse than the overstories, and display a greater array of habitat requirements. Therefore, understory communities may be more likely than overstories to exhibit gap partitioning in these forests. Patterns in understory community composition across gaps suggest that gap partitioning has occurred. The strength of this partitioning effect appears to differ between gap sizes, as smaller gaps showed a less powerful effect. Abundance of ruderal species was strongly related to gap partitioning in larger gaps, while smaller gaps were dominated by competitor species. Partitioning may be related to an interactive relationship between harvest-related ground disturbance and resource gradients. Therefore, considerations of gap partitioning processes should take into account intensity and spatial distribution of ground disturbance in relation to resource gradients. In addition, conditions necessary for the expression of gap partitioning in understory vegetation communities may be rare in natural gaps in this region. The influence of gaps on understory vegetation communities in the surrounding forest appears to be relatively small. This small influence extent may help explain the lack of a stand level response to gap formation in these stands. Larger gaps exhibit a slight influence on the understory plant community in the surrounding forest to the north of the gap. In small gaps, there seemed to be an influence of the surrounding forest on gap interiors, resulting in an area of influence smaller than the physical gap area. This relationship may indicate that the area of gap influence on understory vegetation may not scale linearly with physical gap size. Species diversity was higher in gap interiors than in surrounding thinned forests. However this effect was partially due to the presence of exotic species, which showed an affinity for gap interiors. Late successional associated species were negatively related to gap interiors, but only in the larger gap size. Gap creation appears to be promoting small scale species diversity in these stands, but creation of large gaps may also promote the establishment of exotic species and may have a negative effect on late successional associated species. However, any and all of these effects may be transient, as understory communities will be strongly affected by overstory re-establishment, and related changes in resource availability. In general, gap formation may influence small-scale stand heterogeneity as evidenced by understory plant communities, but this effect may rely strongly on the nature of gap formation and intensity of disturbance related to this formation.

One of the human activities impacting biodiversity is the cutting of old-growth forests. In response to the controversy surrounding the cutting of old-growth in the coastal rainforest of BC, the Ministries of the Environment and Forests have produced biodiversity guidelines that are to be applied when manipulating stands in the provincial forest. This study augments these guidelines by investigating the diversity differences between second-growth and old-growth forests in relation to site quality. We demonstrate how standlevel plant diversity differs between 40-year-old and old-growth stands in the Very Wet Coastal Western Hemlock subzone (CWHvm) on Vancouver Island. This information is intended to provide foresters with an understanding of the effects of age, disturbance and site quality on stand-level plant diversity, thereby allowing for informed professional management decisions.

Central to the issue of harvest feasibility on the outer BC coast (CWHvh) is the question of whether sites, once harvested, can be regenerated, and whether the time period for replacement and subsequent growth is economically and environmentally acceptable. Since low productivity sites have not been harvested in the past, there is a lack of data to answer this question. We tried to provide an answer by assessing regeneration following natural disturbances. Small scale gap disturbances are the norm within old-growth stands. If regeneration is not a problem in gaps, then we have some evidence that regeneration should not be a problem upon implementation of our management practices. The objectives of this study were: (1) to develop baseline information on the mechanisms and the patterns of regeneration across a sequence of forest types; (2) to assess regeneration success with respect to productivity; and (3) to estimate future growth and productivity.

Thesis (M. En.)--Miami University, Institute of Environmental Sciences, 2008. / Title from first page of PDF document. Includes bibliographical references (p. 16-21-Xx).

The risk of spread and establishment of invasive species to interior habitat within urban parks is of great concern to park managers and ecologists. Informal trails as a vector for this transmission are not well understood. To characterize effects of informal trails on understory plant communities, I conducted a study of the informal trail network in Forest Park, Portland, Oregon. The system of 382 informal trails was mapped and evaluated qualitatively, and from this population a systematic sample was selected for analysis. To identify hotspots of informal trail activity, showing the relationship of informal trails to formal trails, other park features, and trail use level, I evaluated all mapped trails using line density spatial analysis tools. To characterize understory communities, thirty transects were placed along informal trails, with paired transects along nearby formal trails for comparison. I measured percent cover by species for non-graminoid understory plants, and percent total plant cover at different structural layers, for quadrats at regular intervals from the trail edge. I calculated richness and Shannon-Weaver diversity for non-graminoid understory plants. For community analysis, species were grouped by dispersal strategy, native status, and growth form. Observations from system mapping suggest that "hidden" behaviors drive many informal trails: bathroom stops, party spots, waste dumping, and camps make up 28% of all informal trails. Trails to private property are few but represent over 29% of total trail length. Informal trail density is highest along Balch Creek. Hotspots of informal trail presence are associated with trailheads, trail intersections, and water access. Quadrats located within one meter of informal trails showed higher richness and diversity due to increased number of introduced and ruderal species. Formal trails exhibit these same patterns to a stronger degree and over a greater distance (two meters) from the trail edge. Distance from trail edge explained variation in plant communities when grouped by dispersal type, but not by growth form. This study shows that although informal trails are widely distributed throughout the park, they are concentrated in high use areas. The presence of informal trails leads to significant changes in Forest Park plant communities that favor invasive and ruderal species, but these effects appear limited to two meters from the trail edge.

A three dimensional sampling technique was used to compare field understory conditions in Southeastern Louisiana using a laser range finder at three height levels (0.5m, 1.0m, and 1.5m) to LiDAR generated understory conditions to determine if a relationship existed. A similar comparison was made between densitometer crown closure measurements and understory LiDAR vegetation counts. A comparison between overstory LiDAR counts and understory LiDAR counts was also performed. LiDAR and understory counts exhibited a significant linear relationship but were poorly correlated at each sample level (Level-1 R2 = 0.34 ? 0.38, Level-2 R2 = 0.36 ? 0.43). The Level-3 LiDAR slope coefficient was non-significant. The crown closure versus understory linear model did not produce any significant results. The overstory LiDAR versus understory LiDAR model produced a moderate correlation (R2 = 0.5226) and was significant. The process of relating LiDAR points to understory conditions was not repeatable, even in the same geographic region.

Graduation date: 2006 / Best scan available. Ink on original is smeared.

Airborne discrete-return lidar is an active remote sensing technology capable of obtaining accurate, fine-resolution three-dimensional measurements over large areas. Discrete-return lidar data produce three-dimensional object characterizations in the form of point clouds defined by precise x, y and z coordinates. The data also provide intensity values for each point that help quantify the reflectance and surface properties of intersected objects. These data features have proven to be useful for the characterization of many important forest attributes, such as standing tree biomass, height, density, and canopy cover, with new applications for the data currently accelerating. This dissertation explores three new applications for airborne discrete-return lidar data. The first application uses lidar-derived metrics to predict understory vegetation cover, which has been a difficult metric to predict using traditional explanatory variables. A new airborne lidar-derived metric, understory lidar cover density, created by filtering understory lidar points using intensity values, increased the coefficient of variation (R²) from non-lidar understory vegetation cover estimation models from 0.2-0.45 to 0.7-0.8. The method presented in this chapter provides the ability to accurately quantify understory vegetation cover (± 22%) at fine spatial resolutions over entire landscapes within the interior ponderosa pine forest type. In the second application, a new method for quantifying and locating snags using airborne discrete-return lidar is presented. The importance of snags in forest ecosystems and the inherent difficulties associated with their quantification has been well documented. A new semi-automated method using both 2D and 3D local-area lidar point filters focused on individual point spatial location and intensity information is used to identify points associated with snags and eliminate points associated with live trees. The end result is a stem map of individual snags across the landscape with height estimates for each snag. The overall detection rate for snags DBH ≥ 38 cm was 70.6% (standard error: ± 2.7%), with low commission error rates. This information can be used to: analyze the spatial distribution of snags over entire landscapes, provide a better understanding of wildlife snag use dynamics, create accurate snag density estimates, and assess achievement and usefulness of snag stocking standard requirements. In the third application, live above-ground biomass prediction models are created using three separate sets of lidar-derived metrics. Models are then compared using both model selection statistics and cross-validation. The three sets of lidar-derived metrics used in the study were: 1) a 'traditional' set created using the entire plot point cloud, 2) a 'live-tree' set created using a plot point cloud where points associated with dead trees were removed, and 3) a 'vegetation-intensity' set created using a plot point cloud containing points meeting predetermined intensity value criteria. The models using live-tree lidar-derived metrics produced the best results, reducing prediction variability by 4.3% over the traditional set in plots containing filtered dead tree points. The methods developed and presented for all three applications displayed promise in prediction or identification of unique forest attributes, improving our ability to quantify and characterize understory vegetation cover, snags, and live above ground biomass. This information can be used to provide useful information for forest management decisions and improve our understanding of forest ecosystem dynamics. Intensity information was useful for filtering point clouds and identifying lidar points associated with unique forest attributes (e.g., understory components, live and dead trees). These intensity filtering methods provide an enhanced framework for analyzing airborne lidar data in forest ecosystem applications. / Graduation date: 2013

Old-growth stands are important for management, conservation, wildlife, recreation, and maintaining biological diversity in forested landscapes. However, we are lacking the information needed to adequately identify and characterize old-growth stands. This is especially true for high elevation, interior forests. The characterization of stand structure and regeneration pattern will help in the development of site-specific guidelines for identifying old growth stands and restoring some of the old-growth characteristics in managed stands. This pamphlet presents a synopsis of a study investigating stand structure and regeneration of old-growth stands in the Moist Cold Engelmann Spruce - Subalpine Fir (ESSFmc) Subzone near Smithers, B.C. The three stands selected for the study were located on zonal sites, each in different watersheds, and the stands were established after fire. The criteria used for selection were: i) absence of lodgepole pine, ii) presence of advanced regeneration, and iii) abundant snags and coarse woody debris. These stands were presumed to represent the old-growth stage of stand development or the final (climax) stage of secondary succession.

This research examines the relationship between understory plant diversity and logs in a Pacific Northwest (PNW) Sitka spruce (Picea sitchensis)-western hemlock (Tsuga heterophylla) old-growth, coastal forest. These forests are renowned for their high forest productivity, frequent wind storms, and slow log decomposition rates that produce unmatched accumulations of coarse woody debris (CWD) yet few studies have examined the relationship between CWD and understory vegetation ecology. My research addressed this topic by comparing understory plant census data between paired fallen log and forest floor sites (n=20 pairs). My objectives were to: 1) determine the influence of substrate type on community composition and diversity, and 2) examine successional pathways and species assemblage patterns on CWD in various stages of decomposition. To meet these objectives I employed non-metric multidimensional scaling (NMDS) ordinations and unsupervised cluster analyses to identify and compare community assemblages on both substrates. These methods revealed similar species diversity and evenness between log and forest floor sites with compositional differences within and between substrates corresponding to habitat availability for colonization and light and moisture gradients. My results also suggest understory successional pathways related to decay class and characterized by an initial abundance of bryophytes, forbs, and seedlings followed by woody shrubs. Understory communities developing on logs also experienced increasing diversity, evenness, and divergence from forest floor communities consistent with log decomposition. These results differ from findings for boreal forests that reveal increasing similarity between substrate communities with increasing decay class. Recommendations for future research include the employment of a more robust sample size and direct measurements of environmental variables. Additional comparator studies are also needed to confirm the effects of forest type and decomposition on the relationship between CWD and forest understory communities. This study demonstrates how fine-scale wind disturbance fosters biodiversity through the creation of CWD substrate. My results and future research are essential for the development of silvicultural models designed to promote biodiversity in PNW coastal forests.