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Die vorliegende Arbeit befasst sich mit dem großen Oberthema der Ikonizität und ihrer Verbreitung auf verschiedenen linguistischen Ebenen. Ikonizität bezeichnet die Ähnlichkeit zwischen der sprachlichen Form und ihrer Bedeutung (vgl. Perniss und Vigliocco, 2014). So wie eine Skulptur einem Objekt oder einer Person ähnelt, kann auch der Klang oder die Form von Wörtern der Sache ähneln, auf die sie verweisen. Frühere theoretische Ansätze betonen, dass die Arbitrarität von sprachlichen Zeichen und deren Bedeutung ein Hauptmerkmal menschlicher Sprache ist und Ikonizität für die Sprachevolution eine Rolle gespielt haben mag, jedoch in der heutigen Sprache zu vernachlässigen ist. Im Gegensatz dazu ist das Hauptanliegen dieser Arbeit, das Potenzial und die Bedeutung von Ikonizität in der heutigen Sprache zu untersuchen. Die einzelnen Kapitel der Dissertation können als separate Teile betrachtet werden, die in ihrer Gesamtheit das umfassende Spektrum der Ikonizität sichtbar machen. Von der sprachevolutionären Debatte ausgehend wird in den einzelnen Kapiteln auf die unterschiedlichen Ebenen der Ikonizität eingegangen. Es werden experimentelle Untersuchungen zur Lautsymbolik, am Beispiel der deutschen Pokémon-Namen, zur ikonischen Prosodie und zu ikonischen Wörtern, den sogenannten Ideophonen, vorgestellt. Die Ergebnisse der einzelnen Untersuchungen deuten auf die weite Verbreitung der Ikonizität im heutigen Deutsch hin. Darüber hinaus entschlüsselt diese Dissertation das kommunikative Potenzial der Ikonizität als eine Kraft, die nicht nur die Entstehung der Sprache ermöglichte, sondern auch nach Jahrtausenden bestehen bleibt, sich immer wieder neu entfaltet und uns tagtäglich in mündlicher, schriftlicher Form und in Gesten begegnet. / This dissertation is concerned with the major theme of iconicity and its prevalence on different linguistic levels. Iconicity refers to a resemblance between the linguistic form and the meaning of a referent (cf. Perniss and Vigliocco, 2014). Just like a sculpture resembles an object or a model, so can the sound or shape of words resemble the thing they refer to. Previous theoretical approaches emphasize that arbitrariness of the linguistic sign is one of the main features of human language; iconicity, however, may have played a role for language evolution, but is negligible in contemporary language. In contrast, the main point of this thesis is to explore the potential and the importance of iconicity in the language nowadays. The individual chapters of the dissertation can be viewed as separate parts that, taken together, reveal the comprehensive spectrum of iconicity. Starting from the language evolutionary debate, the individual chapters address iconicity on different linguistic levels. I present experimental evidence on sound symbolism, using the example of German Pokémon names, on iconic prosody, and on iconic words, the so-called ideophones. The results of the individual investigations point to the widespread use of iconicity in contemporary German. Moreover, this dissertation deciphers the communicative potential of iconicity as a force that not only enabled the emergence of language, but also persists after millennia, unfolding again and again and encountering us every day in speech, writing, and gestures.

La peur est ce qui permet de réagir à un stimulus aversif par une réponse de défense adaptée à la situation. Elle peut être générée par un ensemble de stimuli naturellement aversifs ou par des stimuli ayant acquis une valeur aversive par apprentissage associatif. Cette thèse a pour but d'étudier les caractéristiques et modifications de la réponse de peur à ces deux types de stimuli au cours de l'ontogenèse. Les études présentées ici utilisent un conditionnement de peur à l'odeur chez le rat qui associe une odeur à un stimulus aversif et permet d'induire très rapidement et durablement des mémoires de peur à l'odeur. La réponse de défense peut alors être étudiée à la fois envers l'odeur apprise et envers le stimulus naturellement aversif. Nous montrons en particulier que la réponse de peur à l'odeur apprise présente un décours temporel corrélé à la durée de l'intervalle de temps entre l'odeur et le stimulus aversif, permettant d'affirmer que les animaux mémorisent et estiment le temps, et ce dès les premiers âges étudiés, avant la maturation des structures cérébrales classiquement impliquées dans cette mémoire temporelle. Par ailleurs, nous nous sommes intéressés aux vocalisations ultrasonores émises en réponse au stimulus aversif et à leur modification au cours de l'ontogenèse. Nous avons mis en évidence deux types de vocalisations chez le raton, dont les caractéristiques et critères d'induction laissent présager un rôle différentiel qui reste à explorer. L'ensemble de ces travaux soulignent que, même si les réponses de défense du rat changent au cours du développement, la capacité à produire ces réponses de manière temporellement adaptée est observée dès le plus jeune âge / Fear allows individuals to react to an aversive stimulus by a defense response adapted to the situation. It can be triggered by naturally aversive stimuli or in response to stimuli that acquired an aversive valence through associative learning. This thesis investigated the characteristics and modifications of fear responses to these two types of stimuli throughout ontogeny. The studies presented here used olfactory fear conditioning in rat, in which an odor is paired with an aversive event and allows to rapidly induce long lasting odor fear memories. Defense responses can then be studied both to the learned odor and to the naturally aversive stimulus. We showed in particular that fear response to the learned odor presents a temporal pattern correlated with the duration of the time interval between the odor and the aversive event, showing that rats can learn about time and they do so at the youngest ages studied here, before the maturation of the brain structures classically involved in interval timing. We also studied the ultrasonic vocalizations emitted in response to the aversive stimulus and their changes throughout ontogeny. We described two types of vocalizations in pups that differ in their characteristics and emission context, suggesting they could have different functions, which needs further exploration. These thesis findings highlight that although the rat’s defense responses changes through ontogeny, the ability to produce temporally adapted responses occurs from the youngest age

Made available in DSpace on 2016-07-27T14:21:35Z (GMT). No. of bitstreams: 1 Bertin Zarate Sanchez.pdf: 1661459 bytes, checksum: 6e55e7bdcfef51576a39b56d92c97883 (MD5) Previous issue date: 2006-06-05 / The aim of this work was: to evaluate the newborn s heart rate (HR) changes using live maternal and non-maternal vocalizations in adult-directed and motherese patterns. The work methodology was based in natural observation and continuous recording of the audio-cardiac behavior. The experiment was performed using a ABA design. From a population of seventy mother/newborn dyads that agreed to participate only fifteen were selected; they fulfilled the inclusion s criteria and completed the test. The participants were divided in two groups: drowsy (S) and wakefulness (V), in function of how alert they were during the test. The newborn s HR was observed in an ordered sequence of presentation and suppression periods of vocalizations. HR recordings were taken from a HR-counter in three seconds intervals. The data was analyzed using parametric tests (0.01). T-test were used to evaluate the significance of HR differences obtained in pre-stimulus post-stimulus and stimulation periods. Presentation and suppression of maternal and non-maternal adult-directed and motherese vocalizations differentiately modified the newborn s inner states according to gender and alertness state, suggesting the presence of perceptual stimulus processing abilities. These behaviors and abilities could be associated with a sensible period of attachment and bonding and deserve attention in future researches. In the Phonoaudiology field, the use of a single ABA drawing using live non-maternal motherese voice could turn make behavioral hearing screening of newborn infants, with (or without) risk of congenital deafness, easier. / Este trabalho teve por objetivo: avaliar as alterações do ritmo cardíaco observadas em crianças recém-nascidas durante a apresentação e supressão de estímulos vocais maternos e não-maternos em padrão adulto-natural e motherese. A metodologia utilizada foi a da observação natural e registro contínuo do comportamento audio-cárdico num delineamento de retirada de estímulo. De uma população de 70 díades mãe/recém-nascido contatadas e anuentes em participar da pesquisa foram selecionadas 15 por terem concluído satisfatoriamente a bateria de testes e atendido adequadamente os requisitos de inclusão na amostra. Em função do estado de alerta predominante durante os testes, os participantes foram divididos nos grupos em sono-leve e em vigília . A freqüência cardíaca (FC) dos recémnascidos foi observada numa seqüência ordenada de apresentações e supressões de vocalizações. Os registros foram feitos a intervalos de 3 segundos a partir da leitura visual do valor da FC em um freqüencímetro digital. Os dados foram analisados por meio de testes estatísticos paramétricos para os quais foi estabelecido o valor de 1% (0,01) como nível de significância. Por meio do teste t de Student foi avaliada a significância da diferença entre as médias da FC aferida em cada período de pré-estímulo, pós-estímulo e de estimulação. De acordo com o hipotetizado, pôde observar-se que diante da apresentação e supressão de vocalizações maternas e não-maternas em padrão adulto-natural e motherese modificaram-se os estados internos dos recém-nascidos diferenciadamente de acordo com o gênero e com o estado de alerta, sugerindo a presença de habilidades perceptuais de processamento auditivo dos estímulos. As implicações desses comportamentos e habilidades merecem atenção em estudos futuros, pois podem estar relacionadas ao período sensível de vinculação e apego da díade mãe-filho, aspecto de suma importância no âmbito da Psicologia e demais áreas correlatas. No campo da Fonoaudiologia a simplificação do método para um delineamento ABA, utilizando as vocalizações não-maternas motherese, poderá facilitar a triagem de crianças com (ou sem) risco de deficiência auditiva congênita.

Depuis quelques années, l’étude du code neuronal impliqué dans la perception des signaux de communication acoustique est devenue un domaine de recherche considérable. La littérature récente de ce domaine suggère que la discrimination entre ces signaux reposerait plutôt sur une organisation des décharges neuronales en motifs temporels que sur des variations globales de taux de décharge. Ma thèse a eu pour objectif de déterminer dans quelle mesure une régulation des inhibitions corticales peut (i) changer les motifs temporels déclenchés par des vocalisations conspécifiques et hétérospécifiques et (ii) modifier l’information portée par ces motifs sur l’identité des vocalisations. Nous avons enregistré l’activité neuronale dans le cortex auditif de cobayes anesthésiés en 16 sites corticaux lors de la présentation d’un jeu de vocalisations, et avons partiellement bloqué les inhibitions corticales par des applications de Gabazine (4minutes, 10µm). Dans ces conditions, les réponses évoquées sont plus fortes et les motifs temporels plus marqués. L’information mutuelle quantifiée au niveau de chaque site cortical est augmentée mais l’information populationnelle au niveau de l’ensemble des 16 sites enregistrés n’est pas modifiée, un effet qui peut s’expliquer par le fait que la redondance entre les sites corticaux est augmentée. Nous avons ensuite évalué dans quelle mesure une modulation noradrénergique était susceptible de mimer les effets d’un blocage partiel des inhibitions. Bien que les agonistes utilisés (α1, α2 et ) aient tous induit des modifications des réponses évoquées et de la reproductibilité des motifs temporels, aucun d’entre eux n’a induit de changements importants de l’information portée par les réponses neuronales aux vocalisations. En revanche, les effets induits par la phenylephrine, un agoniste α1, sont vraisemblablement sous-tendus par une action sur les inhibitions intra-corticales, ce qui rend plausible l’hypothèse d’une modulation noradrénergique des inhibitions corticales. Il est donc envisageable que l’action coordonnée de plusieurs systèmes neuromodulateurs puisse moduler les inhibitions corticales et ainsi changer la quantité d’information portée par les neurones corticaux sur l’identité des stimuli à discriminer. / Over the last 10 years, the neural code involved in the perception of acoustic communication signals has become a large area of researches. The recent literature suggests that the discrimination between these signals relies more on the temporal organization of neuronal discharges rather than on global changes of firing rate. My PhD thesis aimed at determining to what extent the regulation of cortical inhibition may (i) change the temporal patterns triggered by conspecific and heterospecific vocalizations and (ii) modify the information carried by these patterns on the vocalization identity. Neuronal activity was recorded in the auditory cortex of anesthetized guinea pigs in 16 cortical sites during presentation of a set of vocalizations, and a partial blockage of intra-cortical inhibition was performed by Gabazine application (4 minutes, 10μm). Under these conditions, evoked responses were stronger and the temporal patterns were reinforced. Mutual information quantified at each cortical site was increased but the information computed at the populationnal level did not change, an effect that could be explained by the fact that the redundancy between cortical sites was increased. We then assessed to which extent the noradrenergic modulation can mimic the effects of a partial blockage of inhibitions. Although all the tested drugs modulated both the evoked responses and the spike-timing reliability, none of the noradrenergic agonists used here (α1, α2 and ) induced significant changes in the information carried by neuronal responses. However, the effects induced by phenylephrine, an α1 agonist, seemed to involve an action on the intra-cortical inhibition, which suggests that a noradrenergic modulation of cortical inhibition can operate in the auditory cortex. It is therefore possible to envision that the coordinated action of several neuromodulatory systems modulates cortical inhibition and thus changes the information carried by cortical neurons on the stimuli identity.

Since the 1960s the focus of animal welfare has been mainly on the negative aspects of welfare and on minimizing and alleviating these negatives. Lately, the focus has been not only on the negative features of welfare that should be kept above a minimum standard, but also on the positive ones that should be enforced. Positive welfare goes a step beyond the common welfare approach and focuses additionally on the positive aspects that animals should have in their lives. This review analyses the behavioral indicators that have been proposed as positive welfare indicators of the domestic pig and have been studied theoretically or experimentally. Various behavioral indicators have been proposed but play is the positive indicator that has been studied the most, followed by exploratory and social affiliative behaviors. Vocalizations and ear and tail postures have also been proposed as promising positive welfare indicators but more research is needed to clearly understand the expression of these behaviors and the affective states that they indicate. According to the literature results, the Welfare Quality protocol for pigs is the only protocol that evaluates the positive welfare of the pig on the farm level. Play, exploratory and social affiliative behaviors are being measured. In addition, positive emotions are being measured via Qualitative Behavioral Assessment (QBA). / Efter 1960-talet, fokuserar djurvälfärden huvudsakligen på de negativa aspekterna av välfärd och på att minimera och lindra de negativa aspekterna. Under de senaste åren, har fokus legat inte bara på att minimera de negativa aspekterna och hålla dem över en minimistandard utan också på att förstärka de positiva. Positiv välfärd går ett steg framåt i det gemensamma välfärdssynsättet och fokuserar dessutom på de positiva aspekterna som djur bör ha i sina liv. Denna översyn analyserar de positiva beteendeindikatorer på djurvälfärd, de som har föreslagits som positiva välfärdsindikatorer för den domesticerade grisen och har studerats teoretiskt eller experimentellt. Bland alla indikatorer har lekbeteende studerats mest, följt av undersökingsbeteende och sociala affiliativa beteenden. Vokalisationer och öron- och svansställningar har också föreslagits som lovande positiva välfärdsindikatorer men fler studier bör göras för att tydligt förstå uttrycket av dessa beteenden och grisens känslor som de indikerar. Enligt litteraturen, är Welfare Quality protokoll för grisar det enda protokollet som utvärderar grisens positiva välfärd på gårdsnivå. Lekbeteende, undersökningsbeteende och sociala affiliativa beteenden mäts. Dessutom, positiva känslor mäts via QBA.

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