An important portion of the large northern permafrost soil organic carbon (SOC) pool might be released into the atmosphere as greenhouse gases following permafrost thawing and subsequent SOC decomposition under future warming conditions, resulting in a warming amplification known as the permafrost carbon feedback. Improved knowledge about the amount, composition and distribution of the permafrost SOC pool is essential when assessing the potential magnitude and timing of the permafrost carbon feedback. This study investigates and compares the SOC storage, composition and distribution in two contrasting permafrost environments: a lowland tundra area in NE Siberia (Tiksi study site), and an alpine area in the Russian Altai Mountains (Aktru Valley study site). Soil pedons were sampled down to 1 m depth and analyzed for key soil properties, i.e., DBD, water content, coarse fraction content, %OC, %IC, C/N ratios and δ¹⁵N values. These soil properties are upscaled by vertical subdivisions based on land cover classes. The role of geomorphology in the accumulation and distribution of SOC in the alpine study site is tested by using a landform and a combined land cover-land form upscaling approach. The estimated mean SOC storage in the upper meter of soils in the alpine site is 3.5 ± 0.8 kg C m¯² compared to 21.4 ± 3.2 kg C m¯² in the lowland tundra site (95% confidence intervals). The inclusion of geomorphology in the upscaling in some cases allows identification of SOC hotspots and areas with very low SOC storage within former land cover classes, therefore improving the landscape SOC storage distribution in the area. The much lower SOC stocks in the alpine site of Aktru Valley can be largely explained by the presence of extensive unvegetated areas in high altitudes (60%), the occurrence active layers deeper than the active soil formation, the enhanced SOM decomposition due to coarse grained, well-drained non-frozen soils, and the negligible occurrence of peatlands and buried organics. Instead, the lowland tundra site in NE Siberia presents important amounts of relatively undecomposed SOM in the permafrost layer. Thus, under future climate warming, alpine permafrost environments such as Aktru Valley may become a net C sink due to an upward shift of vegetation zones and an increase in plant productivity and soil development. Contrarily, lowland tundra areas such as Tiksi may become important C sources since the small increase in C uptake by photosynthetic plants will be outweighed by the thawing and subsequent decomposition of the much larger permafrost SOC pool.

abstract: The plateau pika (Ochotona curzoniae), a small burrowing lagomorph that occupies the high alpine grassland ecosystems of the Qinghai-Tibetan Plateau in western China, remains a controversial subject among policymakers and researchers. One line of evidence points to pikas being a pest, which has led to massive attempts to eradicate pika populations. Another point of view is that pikas are a keystone species and an ecosystem engineer in the grassland ecosystem of the QTP. The pika eradication program raises a difficult ethical and religious dilemma for local pastoralists, and is criticized for not being supported by scientific evidence. Complex interactions between pikas, livestock, and habitat condition are poorly understood. My dissertation research examines underpinning justifications of the pika poisoning program leading to these controversies. I investigated responses of pikas to habitat conditions with field experimental manipulations, and mechanisms of pika population recovery following pika removal. I present policy recommendations based on an environmental ethics framework and findings from the field experiments. After five years of a livestock grazing exclusion experiment and four years of pika monitoring, I found that grazing exclusion resulted in a decline of pika habitat use, which suggests that habitat conditions determine pika population density. I also found that pikas recolonized vacant burrow systems following removal of residents, but that distances travelled by dispersing pikas were extremely short (~50 m). Thus, current pika eradication programs, if allowed to continue, could potentially compromise local populations as well as biodiversity conservation on the QTP. Lethal management of pikas is a narrowly anthropocentric-based form of ecosystem management that has excluded value-pluralism, such as consideration of the intrinsic value of species and the important ecological role played by pikas. These conflicting approaches have led to controversies and policy gridlock. In response, I suggest that the on-going large-scale pika eradication program needs reconsideration. Moderation of stocking rates is required in degraded pika habitats, and Integrated Pest Management may be required when high stocking rate and high pika density coexist. A moderate level of livestock and pika density can be consistent with maintaining the integrity and sustainability of the QTP alpine steppe ecosystem. / Dissertation/Thesis / Doctoral Dissertation Biology 2016

La mise en place des premiers chemins de fer en Savoie est un sujet qui est longtemps resté dans l'ombre des études sur la percée du tunnel du Mont-Cenis. Cependant, l'histoire de ce mode et les intentions sous-jacentes à la construction de ce réseau sont bien antérieures à la mise en exploitation de cet ouvrage (1871) et même des débuts de son percement (1857). Ce tunnel est évidemment un élément essentiel, voire central, en ce qu'il accompli la quête de continuité modale ferroviaire à l'échelle de la liaison Europe du nord-péninsule italienne. Mais, il est aussi l'aboutissement d'un processus technique, économique et politique qui débute dans les années 1830. L'intérêt du chemin de fer en Savoie ne réside ainsi pas uniquement dans la prouesse technique qu'il représente. Il convient, à la lumière des évolutions de l'historiographie récente, d'interroger cet objet singulier à plus d'un titre. Cette singularité tient à deux éléments majeurs autour desquels il est possible de problématiser le traitement proposé : l'antériorité à la plupart des chemins de fer établis dans les Alpes, et le contexte politique particulièrement instable. La question de l'unité italienne, l'annexion de la Savoie à la France, l'évolution des relations franco-sardes puis franco-italiennes contribuent à dessiner ce chemin de fer qui en est une émanation plus ou moins directe. D'un outil de liaison entre un centre et sa périphérie « d'outre-monts » (époque sarde), ce chemin de fer devient un outil de projection internationale d'échelle européenne avec le tunnel, avant de devenir un enjeu d'intégration d'une périphérie dans le territoire français. La dimension locale et sociale ainsi que les questions touchant aux représentations ne sont toutefois pas à négliger, comme nous y invitent les récentes évolutions qui peuvent se caractériser par le passage d'une histoire des transports à une histoire de la mobilité. La réflexion proposée tend, en considérant ces éléments, à interroger le chemin de fer en Savoie des années 1830 aux années 1880 comme le miroir d'une convergence d'ambitions sur un même territoire politiquement intermédiaire. Le chemin de fer en Savoie, tout en demeurant un instrument politique et une aventure financière qui doit s'adapter au contexte géopolitique, ne serait-il pas plus profondément un nœud de confrontation entre recherches d'opportunités locales et d'intérêts internationaux ? La recherche de rentabilité des acteurs financiers (Compagnie Savoyarde, Compagnie Victor-Emmanuel, Compagnie du PLM) qui prennent part à la construction et à l'exploitation du réseau qui se met progressivement en place à travers et sur le territoire savoyard, se heurtent aux visées de l'Etat sarde, puis français. Ces derniers entendent de leur côté faire jouer principalement un rôle politique au chemin de fer. Ces intérêts politiques se heurtent à leur tour aux ambitions commerciales des puissances européennes, notamment de la Grande-Bretagne, qui voient dans le chemin de fer à travers la Savoie – et surtout dans le tunnel du Mont-Cenis – un axe de transit à portée internationale vers la Méditerranée et donc vers les Indes. Les élites locales, de leur côté, voient dans le chemin de fer un puissant moteur de développement économique et un moyen pour leur territoire d'exister dans la politique nationale. Quant au reste de la population savoyarde, il demeure en dehors des débats qui entourent les intentionnalités et la matérialisation du chemin de fer, tout en subissant les bouleversements qui découlent de son implantation. Fascinations et peurs se mêlent dans leurs représentations de cette incarnation du progrès. Le chemin de fer se pose ainsi en point de rencontre et d'articulation des diverses lignes de force qui font la trajectoire du territoire savoyard. / The implementation of the first railway in Savoy is a subject that has long been in the shadow of the breakthrough studies of the Mont -Cenis . However, the story of this mode and the intentions behind the construction of this network are prior to the operation of this book ( 1871 ) and even the beginnings of its opening ( 1857 ) . This tunnel is obviously an essential or central element, it made ​​the quest for continuity modal rail across Europe connecting the North Italian peninsula. But it is also the result of a technical, economic and political process that began in the 1830s. The interest of railway in Savoy and lies not only in the technical achievement it represents. It should , in the light of developments in recent historiography , examine this singular object in more than one way . This singularity is due to two major elements around which it is possible to problematize the proposed treatment: prior to most railways established in the Alps, and particularly unstable political context. The question of the unification of Italy , the annexation of Savoy to France , the evolution of Franco-Sardinian Franco-Italian relations and help shape the railway which is a more or less direct emanation . On the linker between the center and the periphery of " beyond the mountains " ( Sardinian time) , the railway becomes a tool for screening international European level with the tunnel before becoming an issue of integration a periphery in the french territory. Local and social issues as well as the performances are, however, not be neglected , as we invite recent developments that may be characterized by the passage of a story of transport at a story of mobility . The proposed reflection tends considering these elements , to question the railway Savoie 1830s to the 1880s as the mirror of a convergence ambitions in the same territory through politically. The railway in Savoy, while remaining a political instrument and a financial adventure that must adapt to the geopolitical context , would it not deeper node confrontation between local research opportunities and international interests ?

Datainsamling har genomförts i Tarfaladalen under 5 dagar vid månadsskiftet mellan mars och april 2017. Insamlingen resulterade i 36 BTS-mätningar (Bottom Temperature of Snow cover) som därefter har använts tillsammans med data från tidigare insamlingar, till att skapa en sammanställd modell över förekomsten av permafrost omkring Tarfala. En statistisk undersökning syftade till att identifiera meningsfulla parametrar som permafrost beror av, genom att testa de oberoende variablerna mot BTS i en stegvis regression. De oberoende faktorerna höjd över havet, aspekt, solinstrålning, vinkel och gradient hos sluttningar producerades för varje undersökt BTS-punkt i ett geografiskt informationssystem.                 Den stegvisa regressionen valde enbart höjden som signifikant variabel, höjden användes i en logistisk regression för att modellera permafrostens utbredning. Den slutliga modellen visade att permafrostens sannolikhet ökar med höjden. För att skilja mellan kontinuerlig, diskontinuerlig och sporadisk permafrost delades modellen in i tre zoner med olika sannolikhetsspann. Den kontinuerliga permafrosten är högst belägen och därav den zon där sannolikheten för permafrost är störst, denna zon gränsar till den diskontinuerliga permafrosten vid en höjd på 1523 m. Den diskontinuerliga permafrosten har en sannolikhet mellan 50–80 % och dess undre gräns på 1108 m.ö.h. separerar den diskontinuerliga zonen från den sporadiska permafrosten / A field data collection has been carried out in Tarfala valley at the turn of March to April 2017. The collection resulted in 36 BTS-measurements (Bottom Temperature of Snow cover) that has been used in combination with data from earlier surveys, to create a model of the occurrence of permafrost around Tarfala. To identify meaningful parameters that permafrost relies on, independent variables were tested against BTS in a stepwise regression. The independent variables elevation, aspect, solar radiation, slope angle and curvature were produced for each investigated BTS-point in a geographic information system.                 The stepwise regression selected elevation as the only significant variable, elevation was applied to a logistic regression to model the permafrost occurrence. The final model showed that the probability of permafrost increases with height. To distinguish between continuous, discontinuous and sporadic permafrost, the model was divided into three zones with intervals of probability. The continuous permafrost is the highest located zone and therefore has the highest likelihood, this zone delimits the discontinuous permafrost at 1523 m a.s.l. The discontinuous permafrost has probabilities between 50-80 % and its lower limit at 1108 m a.s.l. separates the discontinuous zone from the sporadic permafrost.

Le charbon de bois (biochar), est un amendement qui améliore les propriétés physico-chimiques du sol, augmente le stockage du carbone et les productions agricoles. Les anciens sites de production de charbon permettent d'évaluer son impact directement sur le terrain et sur le long terme. Dans les Alpes italiennes on a échantillonné une série de charbonnières daté de 1858 ainsi que les sols de prairies adjacents, ne contenant pas de charbon. 80±21% du carbone provenant de la production du charbon est toujours présent dans le sol et a un temps de résidence moyen de 650±139 ans. Le contenu des nutriments et leur biodisponibilité sont plus élevés dans les charbonnières par rapport à la prairie alentour et, sont plus élevés aujourd'hui qu'en 1858. L'ajout de charbon apporte des nutriments au sol, mais à court terme les ions Ca2+, K+, SO42- et Mg2+ sont lixiviés sous forme de cendres. Le charbon s'avère capable de retenir les dépositions atmosphériques de PO43-, NH4+, NO3- et, sur le long terme, de K+. L'augmentation du contenu en nutriments, la diminution de l'hydrophobie et de la densité apparente du sol augmentent la productivité et la valeur nutritionnelle des espèces alpines fourragères (Festuca nugrescens Lam. et Trifolium pratense L.). La croissance des plantes est limitée par l'azote sur les charbonnière et par le phosphore dans les prairies non-amendés ou amendés récemment. Nous concluons que le charbon/biochar peut être une stratégie pour stocker le carbone dans les sols, augmenter la production de biomasse et la qualité du fourrage des prairies Alpines à long terme. Toutefois les opérations d'enfouissement peuvent être complexes à cause de la géomorphologie des Alpes. / Charcoal or biochar is proposed as a soil amendment to improve physio-chemical soil properties, increase soil carbon (C) stocks and agricultural yields. Ancient charcoal hearths provide an opportunity to investigate its impact under field conditions and in the long term. A series of charcoal hearths and adjacent charcoal-free soils under grassland in the Italian Alps abandoned in 1858 was sampled.80±21% of the C originating from ancient charcoal is still present in the soil today and has a Mean Residence Time of 650±139 years. The content of total and available nutrients is higher in the hearths soils compared to the surrounding grasslands and it is higher today compared to 1858. The input of charcoal directly adds nutrients to soils but Ca2+, K+, SO42- and Mg2+ are leached in the short term after application, as they are lost in the form of ashes. Charcoal is able to retain atmospheric depositions of PO43-, NH4+, NO3- and in the long term K+.The increase in soil nutrient content and decreases in hydrophobicity and bulk density translated into higher plant growth and nutritional values of two alpine fodder species (Festuca nigrescens Lam. and Trifolium pratense L.). Plant growth was N-limited in the charcoal hearths soils and P-limited in the surrounding grasslands not amended or recently amended with charcoal/biochar.We can conclude that charcoal/biochar is a long term strategy to store carbon in soils, improve biomass productivity and fodder quality in alpine grasslands. However charcoal/biochar incorporation into soil can be complex because of the geomorphology of the Alps.

Abstract Deciduous shrubs form the tallest type of vegetation in arctic-alpine areas and are important for ecosystem function. In the southern part of the Eurasian tundra zone, willows (Salix spp.) are the most common species in the shrub layer. In the alpine areas of Northern Fennoscandia, willow shrubs are characteristic to areas between tree line and treeless tundra heaths. Vertical structure and composition of willow-characterized tundra vegetation is affected by a variety of ecological factors including climate and herbivory. In turn, the abundance of the willow canopy affects understory species in several ways that still remain inadequately understood. In this PhD work I describe compositional differentiation of willow-characterized vegetation by using a large data set spanning from north-western Fennoscandia to the Yamal Peninsula in north-western Siberia. I studied environmental factors affecting willow-characterized vegetation and willow growth by using correlative analyses. The factors under investigation were latitude, distance from the sea, depth of thaw, position in the slope, industrial disturbance and reindeer grazing. In addition, I examined the relationships between the shrub biomass estimate and composition and species richness of understory vegetation. The effects of reindeer grazing on vegetation in an alpine forest-tundra ecotone were studied experimentally using reindeer-proof exclosures. I found that willow-characterized vegetation is floristically variable and comprises at least eight vegetation types. The most abundant willow thickets typically have a forb-rich understory. The growth of willow increased along with increasing summer temperatures. However the height of willow was more determined by distance from the sea, thaw depth and slope position. Reindeer grazing decreased the abundance of willow and changed the composition of understory vegetation. In addition, industrial activities were detected to have destructed shrub vegetation and turned it into graminoid-dominated vegetation. Shrub canopies facilitated forbs but decreased the cover of all the other groups including dwarf shrubs, bryophytes and lichens. The species richness of vegetation decreased along with increasing shrub abundance. My study shows that arctic-alpine willow vegetation is more diverse than previously thought. There is a predictable relationship between summer temperatures and willow growth. However, the results also show that there are many factors, both physical and anthropogenic, that are likely to complicate this pattern. Most important of these counteracting effects are industrial activities and reindeer grazing. In the areas where shrubs grow in abundance, the species richness of understory vegetation is likely to decrease and forbs are likely to replace other tundra species.

A detailed understanding of cloud microphysical processes is crucial for a large range of scientific disciplines that require knowledge of cloud particles for accurate climate and weather prediction. This thesis focuses on 3 measurement campaigns, encompassing both airborne and ground based measurements of the microphysical structures observed in cold, warm and occluded frontal systems around the United Kingdom, stratocumulus clouds in the Arctic and many different clouds observed over a 6 week period at a high-alpine site in the Swiss Alps. Particular attention was paid to the origin and distribution of both primary and secondary ice and the dominant features associated with ice phase processes. During investigation of cold, warm and occluded frontal systems associated with mid-latitude cyclones around the U.K., secondary ice was often found to dominate the number and mass concentrations of ice particles in all systems. The presence of large liquid droplets was sometimes observed in close proximity to regions of secondary ice production. The existence of these provides a possible mechanism by which rime-splintering is greatly enhanced through the creation of instant rimers as the large drops freeze. In-situ measurements during the cold frontal case were used to calculate rates of diabatic heating during a comparison between bin-resolved and bulk microphysics schemes. Observations in arctic stratocumulus clouds during spring and summer seasons revealed higher ice concentrations in the summer cases when compared to the spring season. This is attributed to secondary ice production actively enhancing ice concentrations in the summer, due to the higher temperature range the clouds spanned. At Jungfraujoch in the Swiss Alps, ground based measurements allowed us to obtain high spatial scale resolution measurements of cloud microphysics and we found transitions between high and low ice mass fractions that took place on differing temporal scales spanning seconds to hours. During the campaign measurements of aerosol properties at an out of cloud site, Schilthorn, were made. When analysing a Saharan Dust Event that took place a possible link between the number of U.V. fluorescent particles and the number of ice particles was found in the temperature range around -10 ºC.

The Carex nardina complex is a group of one to three species (C. nardina, C. hepburnii, C. stantonensis) and six taxa of unispicate sedges (Cyperaceae), the taxonomy of which has been controversial since the 1800s. As initial DNA phylogenies suggested that the complex was nested within Carex section Filifoliae and sister to C. elynoides, a species often confused with C. nardina and sympatric with it in the western North American Cordillera, analyses were conducted to determine whether C. hepburnii, C. stantonensis and other infraspecific taxa could be the result of hybridization. Morphometric and molecular analyses found no substantial evidence for hybridization and supported the recognition of no taxon beyond C. nardina. Consequently, this study concludes that the complex comprises a single variable species, Carex nardina, distributed throughout arctic North America south through the western Cordillera to New Mexico with a minor portion of its range in northeastern Russia, northwestern Scandinavia and Iceland.

Ce travail de thèse avait pour objectif (i) de quantifier les contraintes métaboliques en slalom (SL), en slalom géant (GS) pour des durées d’efforts courtes et longues, représentatives des temps extrêmes de compétitions contemporaines ; (ii) de comparer les demandes métaboliques entre le ski alpin in situ (SL et GS) et différentes motricités (bonds latéraux sur box, slide, course à pied, cyclisme, bonds latéraux sur trampoline). Quelle que soit la durée de GS en compétition, le métabolisme aérobie est prédominant dans la fourniture d’énergie. En revanche, les contributions respectives des métabolismes aérobie et glycolytique à la fourniture d’énergie ne sont pas significativement différentes en SL. Par ailleurs, la course à pied et le cyclisme sont les motricités investiguées les moins spécifiques de SL et GS. En revanche, la motricité des bonds latéraux sur box et celle des bonds latéraux sur trampoline représentent des similarités physiologiques intéressantes par rapport au SL et GS. A fortiori, la motricité du slide est la plus spécifique de SL et GS au regard de la spécificité de ses paramètres physiologiques (part d’implication des métabolismes aérobie et glycolytique, débits énergétiques total, glycolytique et aérobie, valeurs de [La]pic et de Δ[La]). Subséquemment, l’objectif de l’étude n°3 était de déterminer la reproductibilité d’un test physique de 80s utilisant la motricité du slide. Le 80s-slide-test est un test physique spécifique du ski alpin et reproductible. En effet, le nombre total d’appuis, V̇ O2pic, V̇ O2moy et FCmoy sont reproductibles. Ainsi, le 80s-slide-test constitue une alternative pertinente aux tests physiques non spécifiques du ski alpin menés en laboratoire. Enfin, l’objectif de l’étude de cas n°4 était de décrire la stratégie d’allure du skieur alpin au cours d’une manche longue de GS. Le skieur alpin adopterait une stratégie d’allure au cours d’une manche longue de GS en diminuant l’intensité de ski de manière anticipée dès la section 1 (premières 52s) puis durant la section 2 (dernière 29s) lors d’un GS long (81s). / The objective of this thesis was (i) to assess metabolic constraints in slalom (SL) and giant slalom (GS) during short and long effort according to extrem contemporary races duration ; (ii) to clarify motor skills (box bouncing, sliding, running, cycling, trampoline lateral bouncing) energetically specificity to alpine ski racing. Regardless of GS length the aerobic system was the primary energy system involved. Aerobic and glycolytic energy system contribution were not different in SL. Furthermore running and cycling were most unspecific motor skills investigated regarding SL and GS. On the other hand box bouncing and trampoline lateral bouncing presented interesting physiological specificities. Moreover no significant difference in aerobic and glycolytic energy system contribution, total energy output, glycolytic energy output, aerobic energy output, phosphagen energy output, [La]peak, Δ[La] were observed between sliding and both GS and SL. Subsequently the aim of the third study was to determine the test-retest reliability of a 80s test involving sliding motror skills. The 80s-slide-test is a specific and reliable test. Indeed the 80s-slide-test showed large reliability for total push-offs number, V̇ O2peak, V̇ O2moy, FCpeak and FCmoy. Hence 80s-slide-test is a relevant alternative to lab unspecific test for alpine ski racers. Finally the aim of the fourth study was to analyse pacing strategy in long GS. Alpine ski racer may run pacing strategy skiing decreasing intensity during section 1 (first 52s) and during section 2 (last 29s) in long GS (81s).

The aim of the diploma thesis is sustainable tourism in the mountainous environment. Alpine habitats include a valuable natural environment where tourism can leave lasting effects. Research focuses on the operation of mountain chalets in the High Tatras and the influence of alpine tourism on the environment in terms of three pillars of sustainable development using qualitative methods, namely the triangulation method. Key words: sustainable tourism, mountain chalets, alpine tourism, High Tatras