Pentadesma butyracea Sabine est l’une des quatre espèces du genre Pentadesma endémique de l’Afrique. Elle est distribuée de la Sierra Léone au Gabon dans deux grands types d’habitats :les forêts denses humides discontinues du domaine guinéo-congolais (Haute- et Basse-Guinée) et le domaine soudanien du couloir sec du Dahomey (assimilé à une barrière à l’échange de gènes et d’espèces entre les deux blocs guinéo-congolais). Dans ce dernier, l’espèce se retrouve dans des galeries forestières et occupe une place capitale dans le développement socio-économique des communautés locales en raison des multiples biens et services que procurent ses produits (alimentation, médecine et pharmacopée traditionnelle, etc.). Cependant, des pressions d’origines multiples, telles que le ramassage des graines pour fabriquer du beurre, la fragmentation de l’habitat et sa destruction en faveur du maraîchage, les pratiques culturales inadaptées, les incendies, font peser de lourdes menaces sur l’espèce.<p>Le but de ce travail est d’acquérir les connaissances requises pour la conservation et la gestion durable des ressources génétiques de l’espèce. Trois objectifs ont été définis :(i) étudier la phylogéographie de l’espèce, (ii) étudier sa variabilité morphologique et génétique au Bénin et (iii) caractériser sa biologie de reproduction. En amont de ces travaux, nous avons développé onze marqueurs microsatellites nucléaires chez P. butyracea (chapitre 2). Ils ont été utilisés pour l’étude de la phylogéographie et la diversité génétique de P. butyracea (chapitres 3 et 5), ainsi que pour étudier la dépression de consanguinité et les paramètres de son système de reproduction (chapitre 7).<p>La caractérisation de la répartition spatiale des lignées génétiques de régions intergéniques de l’ADN chloroplastique et de l’ADN ribosomal (ITS) a détecté deux lignées génétiques allopatriques entre le Haut et le Bas-Guinéen, indiquant une forte différenciation génétique et un signal phylogéographique. L’analyse des microsatellites détecte trois pools géniques correspondant aux trois régions étudiées (Haute Guinée, Dahomey Gap et Basse Guinée). La diversité génétique est faible dans le Dahomey Gap, modérée dans le Haut-Guinéen et élevée dans le Bas-Guinéen. Ces résultats indiquent une séparation très ancienne des populations d’Afrique centrale et d’Afrique de l’ouest, alors que celles du Dahomey Gap pourraient résulter des forêts denses humides de l’Afrique de l’ouest lors de la période Holocène humide africaine. Dans ce couloir sec, les populations ont subi une forte dérive génétique, potentiellement due à des évènements de fondation. Au Bénin, deux groupes éco-morphologiques ont été détectés suivant un gradient nord-sud, contrastant avec deux pools géniques présentant une distribution est-ouest. <p>P. butyracea est une espèce auto-compatible majoritairement allogame. La corrélation de paternité est plus élevée aux niveaux intra-fruit vs. inter-fruits, et au sein d’une population de petite taille vs. de grande taille. Les principaux pollinisateurs au Bénin sont deux oiseaux (Cyanomitra verticalis, Cinnyris coccinigastrus) et trois abeilles (Apis mellifera, Meliponula togoensis, Hypotrigona sp.). La productivité totale en fruits augmente en fonction de l’âge de l’arbre et varie en fonction de l’année, atteignant un pic pour les arbres ayant un diamètre de 60-80 cm. Les graines sont récalcitrantes et ont une teneur en eau de 42.5 ± 2.9 %. <p>L’analyse des paramètres de reproduction et de diversité génétique, associés aux facteurs écogéographiques, nous a permis de proposer un échantillon de neuf populations représentatives de la diversité à l’échelle du Bénin, dans la perspective d’une conservation in situ. Le succès de celle-ci dépendra des efforts conjugués des communautés locales, de la recherche forestière et de la définition d’un cadre législatif par le politique pour la protection des habitats. La conservation ex situ est envisagée sous forme d’un verger rassemblant diverses origines, présentant l’intérêt supplémentaire de permettre d’étudier les contributions de la diversité génétique et de la plasticité phénotypique à la variation phénotypique. / Pentadesma butyracea Sabine is one of the four species of the endemic genus Pentadesma in Africa. The species is distributed from Sierra Leone to Gabon in two major types of habitats: the discontinuous and dense Guineo-Congolian rainforests (Upper and Lower Guinea) and the Sudanian domain of the dry corridor of Dahomey (considered as a barrier to the exchange of genes and species between Upper and Lower Guinea). In the latter, the species is found in gallery forests and plays a vital role in the socio-economic livelihood of local communities due to the various resources and services that provide its products (food, medicine and traditional, etc.). However, pressure from many sources including the collection of seeds to make butter, habitat fragmentation and its destruction for market gardening, inadequate agricultural practices, fires, are serious threats to the species.<p>The aim of this work was to acquire appropriate knowledge for the conservation and sustainable management of genetic resources of the species. Three objectives were defined (i) study the phylogeography of the species; (ii) evaluate its morphological and genetic variability in Benin; and (iii) characterize its reproductive biology. In a preliminary work, eleven nuclear microsatellite markers of P. butyracea were developed (Chapter 2). They were used for the study of phylogeography and genetic diversity of P. butyracea (chapters 3 and 5), and to study the inbreeding depression and parameters of its breeding system (Chapter 7).<p>The characterization of the genetic lineages and their spatial distribution using intergenic regions from chloroplast DNA and ribosomal DNA (ITS) region detected two allopatric genetic lineages between Upper and Lower Guinea, indicating a high genetic differentiation and a phylogeographic signal. Microsatellite markers allowed us to detect three genepools matching with the three studied regions (Upper Guinea, Dahomey-Gap and Lower Guinea). Genetic diversity was low in the Dahomey Gap, moderate in Upper Guinea and high in Lower Guinea. These results indicate an ancient separation of populations from Central and West Africa, while those from Dahomey Gap could originate West African rainforests (Upper Guinea) during the African humid Holocene period. In this dry corridor, populations experienced high genetic drift, possibly due to founding events. In Benin, two eco-morphological groups were detected following a north-south gradient, contrasting with two gene pools presenting an east-west distribution.<p>Pentadesma butyracea is a self-compatible, mainly allogamous species. The correlation of paternity was higher within-fruit vs. among-fruits, and in population of small size vs. large size. The main pollinators in Benin are two birds (Cyanomitra verticalis, Cinnyris coccinigastrus) and three bees (Apis mellifera, Meliponula togoensis, Hypotrigona sp.). Total productivity in fruit increases with tree age and varies yearly, reaching a peak for trees of 60-80 cm of diameter class. Seeds are recalcitrant (i.e. they cannot be conserved at low temperature), having a water content of 42.5 ± 2.9% at maturity.<p>The analysis of reproduction and genetics parameters, associated with eco-geographic factors, enabled us to select nine populations representative of the diversity in Benin, from the perspective of in situ conservation. The success of the latter will depend on combined efforts of local communities, forest research and an adequate legislative framework for the protection of habitats. Ex situ conservation is envisaged as an orchard assembling various origins, and would have the additional advantage of allowing to study the contribution of genetic diversity and phenotypic plasticity to phenotypic variation. / Doctorat en Sciences / info:eu-repo/semantics/nonPublished

La connaissance de la distribution spatiale des espèces et leurs déterminants constituent les principaux thèmes de la biogéographie et de l’écologie. Cette thèse a été réalisée sur les Fabaceae de l’Afrique Centrale :Burundi, République Démocratique du Congo et Rwanda. Composées de trois sous-familles (Caesalpinioideae, Faboideae et Mimosoideae), les Fabaceae sont présentes dans tous les biomes terrestres du monde. La présente étude a pour objectif d’évaluer et localiser la diversité spécifique, déterminer les zones de conservation des Fabaceae, identifier leur modèle de distribution spatiale et examiner l’impact potentiel de la déforestation sur les espèces indicatrices de cette famille. Les facteurs environnementaux déterminant la distribution des espèces endémiques de Fabaceae, les zones favorables à leur présence et l’évaluation de leur état de conservation ont également été analysés. L’étude a consisté à vérifier et à digitaliser tous les échantillons d’herbiers des Fabaceae conservés au Jardin Botanique National de Belgique et à l’Université Libre de Bruxelles. Les Systèmes d’Information Géographique ont été largement utilisés. Les analyses spatiales ont montré une distribution non uniforme de la diversité spécifique des Fabaceae. Les zones où la diversité spécifique coïncide avec celle des espèces endémiques ont été suggérées comme prioritaires pour la conservation. L’analyse de groupement appliquée sur le jeu de données des Mimosoideae a mis en évidence trois régions floristiques majeures. Ces trois régions correspondent aux régions phytogéographiques définies par White (1979, 1983). L’étude des Caesalpinioideae a confirmé l’existence d’espèces indicatrices de territoires phytogéographiques de Ndjele (1988). L’étude de cas réalisée au Katanga (R.D. Congo) a montré que la déforestation constitue une menace potentielle pour ces espèces, les plus vulnérables étant inféodées aux habitats forestiers. La distribution potentielle des espèces endémiques de Fabaceae (Caesalpinioideae) a montré une convergence entre les cartes de la distribution actuelle et celles issues de la distribution potentielle. Des sites où les espèces n’ont jamais été signalées ont été identifiés. La distribution potentielle a démontré le rôle des déterminants environnementaux dans la distribution des espèces endémiques. Cette étude a également prouvé que d’importantes proportions des zones de diversité des espèces endémiques ne sont pas couvertes par la conservation actuelle de l’Afrique Centrale. Cette thèse confirme que l’Afrique Centrale est une zone importante de conservation de la biodiversité. L’étude devrait être poursuivie en étendant la distribution potentielle aux Fabaceae non endémiques afin de définir leurs aires de distributions, critère indispensable pour évaluer le degré de vulnérabilité d’une espèce. La création de nouvelles aires protégées renforcerait le système actuel de conservation dans cette région./Knowledge of the spatial distribution of species and its determinants constitutes a principal theme in biogeography and ecology. This dissertation focused on the Fabaceae family of Central Africa: Burundi, the Democratic Republic of the Congo and Rwanda. Composed of three subfamilies (Caesalpinioideae, Faboideae and Mimosoideae), Fabaceae species are present in all terrestrial biomes of the world. The current study aimed to assess and locate Fabaceae species diversity, to determine potential conservation zones, to identify the spatial distribution pattern and to examine the potential impact of deforestation on some indicator species of this family. The environmental factors driving the distribution of endemic Fabaceae have been identified, as well as the potential areas of occurrence; their conservation status in the region has also been studied. The current study has verified and digitized all the herbarium samples kept at the National Botanical Garden of Belgium and the Université Libre de Bruxelles regarding the Fabaceae family. Geographic Information Systems have been used frequently. Spatial analysis showed an uneven distribution of Fabaceae species diversity. The zones where species richness depends on endemic species diversity are suggested to have priority for conservation. A cluster analysis applied to the dataset of Mimosoideae has highlighted three major floristic regions. These three regions correspond to existing phytogeographical regions defined by White (1979, 1983). A study on Caesalpinioideae species of Central Africa has confirmed the existence of indicator species of phytogeographic territories of Ndjele (1988). A case study conducted in Katanga (D.R Congo) showed that deforestation constitutes a potential threat for these species, the most vulnerable being those strictly confined to forest habitats. The potential distribution of endemic Fabaceae (Caesalpinioideae) showed a clear convergence between the current distribution maps and the potential distibutions. Sites where those species had not been reported before have been identified. The potential distribution enabled to identify the importance of the different environmental factors for each endemic species’ distribution. This study also showed that large regions characterized by important endemic species diversity are not covered by current zones of conservation in Central Africa. This dissertation confirms Central Africa as an important zone for biodiversity conservation. The current research should be completed by the potential distribution of non endemic Fabaceae species of Central Africa, an important criterion to evaluate their degree of vulnerability. Creation of new protected areas would reinforce the current status of conservation in the region. / Doctorat en Sciences / info:eu-repo/semantics/nonPublished

La compensation écologique, envisagée dans le respect de la séquence éviter-réduire-compenser les impacts, cherche à apporter des réponses à la crise actuelle d’érosion de la biodiversité. L’objectif de la thèse est de comprendre quels sont les enjeux organisationnels et institutionnels de la mise en œuvre du principe de compensation écologique pour les impacts sur les écosystèmes aquatiques marins et continentaux en Europe et aux Etats-Unis. La thèse mobilise le cadre théorique de l’économie néo-institutionnelle. Les organisations de type hiérarchique (compensation au cas par cas) ont pour principales limites d’avoir une efficacité écologique discutable et de ne pas permettre le suivi et le contrôle des mesures compensatoires ce qui conduit souvent à ce que la compensation ne soit pas ou peu mise en œuvre. Aux Etats-Unis, les banques de compensation consistent en une réponse anticipée et mutualisée aux dommages à l’environnement. Il s’agit d’une forme organisationnelle alternative hybride fortement régulée à mi-chemin entre la hiérarchie et le marché. Cependant, les acteurs ont des stratégies collectives et individuelles qui engendrent des négociations ayant des conséquences sur les enjeux de développement économique et de maintien des objectifs de conservation de la biodiversité. Le système des banques de compensation crée un double phénomène de redistribution des coûts de transaction et de diminution de ceux-ci mais il semble que l’évolution de l’ensemble du cadre d’application de la compensation écologique américain permet aujourd’hui une complémentarité institutionnelle entre ces deux formes organisationnelles plutôt qu’une disparition des formes hiérarchiques. / Biodiversity offset, as the last step of the mitigation hierarchy (avoid/reduce/offset), aims to address the current biodiversity loss crisis. The main goal of this PhD, which relies on the new institutional economics approach as a theoretical framework, is to understand the organizational and institutional issues of implementing biodiversity offsets policies for impacts on continental and offshore wetlands in Europe and the United-States. Hierarchical organizations (i.e. permittee responsible mitigation) often lead to little to no implementation of biodiversity offsets because the ecological effectiveness of case-by-case compensatory measures is unproven and the monitoring and control of numerous measures are difficult. In the United-States, the mitigation banking system aims to compensate for multiple small impacts on wetlands by carrying out restoration projects on fewer but larger wetland areas in order to reach the goal of no net loss of biodiversity. This hybrid organizational form is highly regulated and therefore stands halfway between hierarchical and market organizations. However, actors have individual and collective strategies that generate negotiations with consequences on economic development and biodiversity conservation stakes. The mitigation banking system leads to a decrease of transaction costs and a change of the actors bearing them. It seems that the evolution of the whole framework of biodiversity offsets policy implementation in the United-States leans toward an institutional complementarity between these two organizational forms rather than a disappearance of hierarchical organizations.

The most contemporary approach to biodiversity conservation within South Africa is that of community-based initiatives, which seek to combine biodiversity conservation with socioeconomic development. As a challenge to the Western, science laden approaches to conservation there is an increasing need for community initiatives to reflect the values of local communities. Values of local communities and the management body, CapeNature, with regards to Driftsands Nature Reserve, Cape Town, were captured and analytically coded through the qualitative methods of interviewing and participant observation in order to develop a grounded theory and model. A discussion of the expressed values suggests that community-based conservation initiatives are doing little to include community values even though there is a large degree of agreement between these and corporate values. As such, it is questioned whether community-based conservation can be practised within an organisation which, due to procedures and protocols, is top-down in its approach. / Geography / M. Sc. (Geography)

This study aimed to investigate the spatial extent of Land-Use Land-Cover (LULC) change and the implications for ecosystem services in order to develop a sustainable land-use management framework for traditional authorities in South Africa. Effectively, this study undertook an insightful examination of the impacts that arise from policy decisions and practices, which unfortunately were found to be ineffective. The methodologies and approaches used in this study included both quantitative and qualitative techniques. The critical quantitative method employed in this research was the use of survey questionnaires to collect primary data. Qualitative approaches, such as one-on-one and key informant interviews, were used to triangulate the findings. Remote sensing and geographic information system (GIS) methods were used to investigate changes in LULC from 1990 to 2018 through the use of data obtained from the South African National Land-Cover project. Stochastic models were used to predict future LULC changes from 2018 to 2050. The Co$ting Nature Policy Support System was used to identify and undertake economic valuation of services provided by ecosystems. Statistical analysis using the Statistical Package for the Social Sciences was used to identify correlations and the reliability of the data, while graphs and tables were generated to identify patterns and lessons from the research. Between 1990 and 2018, significant changes in land cover were noticed for thickets and dense bush, woodlands, waterbodies, subsistence agriculture, and built-up areas. Woodlands changed by over 1 000 hectares (ha) per year, while thickets decreased by over 900 ha per year. Drivers of these changes include deforestation, among others. Future predictions for LULC revealed that between 2018 and 2050, almost 500 ha of woodlands would be lost to built-up areas. The aggregate value of the services flowing from ecosystems was found to be R9 509 044 608.00. A significant issue was that 90% of the traditional leaders interviewed could not positively respond to whether they knew the extent of the land they presided over, which raised questions regarding the effectiveness of their management systems. Recommendations were made in this study to address the limitations identified in the land-use management practices by adapting elements of the main theoretical frameworks, namely the Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services framework; the Drivers, Pressures, States, Impacts and Responses framework; the sustainability theory; and the hierarchy of plans, into a new framework designed specifically for traditional leaders, titled the “Traditional leaders land-use decision support framework”. / Environmental Sciences / D. Phil. (Environmental Management)

Environmental positive externalities from public provision, such as the benefits yielded from the public measure of nature conservation, are often not internalized. Potential sub-optimal public service provision can be expected from such a condition, leading to inefficiency, if the benefits at a greater territorial scale are not acknowledged. This holds particularly true for intergovernmental fiscal relations in a decentralizing multi-tiered governmental system. Moreover, in developing countries the fiscal capacities to perform measures of ecological public functions are limited with their fiscal needs for these functions often appearing to outweigh their fiscal capacities. Research at the interface of the economic theory of fiscal federalism, the sustainability concept, and policies related to conservation and the environment is relatively new. Furthermore, in the literature on environmental federalism the emphasis tends to be comparatively less on the benefits of positive environmental externalities. The essential contribution of this study is an extension of this research field that is still in its infancy by applying the specific case of Indonesia as the context, on account of this tropical country‟s ecological significance as well as its recent developments during the fiscal decentralization process. The overall aim of this study is to assess the possibilities of ecological fiscal transfers as a set of instruments in the public sector to internalize environmental externalities. To this end, the study traces the development as well as the current state of intergovernmental fiscal transfers in Indonesia in terms of ecological purposes. On the basis of this knowledge, the study offers new policy perspectives by proposing a number of policy options for ecological fiscal transfers in the context of the functioning fiscal transfer system and institutions between the national and the subnational (province and local) governments as well as among jurisdictions at the same governmental level. The incorporation of an explicit ecological indicator into general-purpose transfers is the first option. The second option is derived from a revenue-sharing arrangement. In this arrangement, two sub-options are proposed: first, shared revenues from taxes are distributed on the basis of the ecological indicator and, second, shared revenues from natural resources are earmarked for environmental purposes. Finally, the third option suggests an extension of existing specific-purpose transfers for the environment. The potential and limitations of the respective options are addressed. Additionally, a short treatment is given to the discourse on the possible mobilization of fiscal resources in the context of tropical deforestation and global climate change. The research concentrates mainly on the first option, namely the incorporation of an ecological indicator into the structure of general-purpose transfer allocation. In order to substantiate an explicit ecological dimension in the transfer, it extends the present area-based approach by introducing a protected area indicator while maintaining the remaining socio-economic indicators in the fiscal need calculation. The parameter values of area-related indicators are adjusted and subject to the properties of the existing formula. The simulation at the provincial level yields the following results. First, more provinces lose rather than gain from the introduced ecological fiscal transfer when compared to the fiscal transfer that they received in the reference fiscal year. Second, on average the winning provinces obtain a higher level of transfer from the introduction of an ecological indicator in the fiscal need calculation. The extent of the average decreases for the losing provinces, however, it is lower compared to the extent of the average gain by their winning counterparts. In terms of spatial configurations of the general-purpose transfer with an ecological indicator, provinces in Papua would benefit most from the new fiscal regime whereas provinces in Java and Sulawesi, with a few exceptions, would suffer a transfer reduction. Kalimantan and Sumatera show a mixed pattern of winning and losing provinces. The analysis on the equalization effects of the general-purpose transfers makes the following important contributions. It suggests that, first, the transfers are equalizing and, second, the introduction of the protected area indicator into the structure of these transfers plays a significant role in the equalizing effect, particularly in the presence of provinces with a very high fiscal capacity and when the area variable is also controlled. All of these new insights are imperative in the design of fiscal policy which intends to integrate explicit ecological aspects into the instruments of intergovernmental fiscal transfers. Since a formula-based fiscal transfer distribution is intrinsically zero-sum, the aforementioned configuration of winning and losing jurisdictions is conceivable. Among other future perspectives, it is the task of further research to explore ecological fiscal transfer instruments and associated measures that on the one hand seek to induce the losing provinces to join their winning counterparts and, on the other hand, are still subject to the requirements of the rational fiscal transfer mechanism.

Wir befinden uns inmitten einer globalen Biodiversitätskrise mit steigender Aussterberate, die die natürliche Hintergrundrate bereits jetzt um das Tausendfache überschreitet. In terrestri-schen Lebensräumen geht vom Landnutzungswandel die größte Gefahr für die Biodiversität aus. Besonders die ursprünglich hohe Vielfalt im mitteleuropäischen Grünland ist seit der In-dustrialisierung durch Nutzungsintensivierung und -aufgabe stark zurück gegangen. Traditionell genutztes, beweidetes Grünland ist in Mitteleuropa selten geworden. Dennoch deuten einige aktuelle Studien auf seine große Bedeutung für den Erhalt der Biodiversität in der Agrarland-schaft hin. Allerdings fehlen umfassende, systematische Untersuchungen für viele traditionelle Grünlandsysteme. Um diese Wissenslücke zu verringern, untersuchte ich in meiner Dissertation die Relevanz voralpiner, traditionell bewirtschafteter Allmendweiden als Refugium für bedrohte Zielarten (Baumpieper [Anthus trivialis], Blaukernauge [Minois dryas]) sowie für arten- und individuenreiche Zoozönosen (Heuschrecken [Orthoptera]) der mitteleuropäischen Agrarlandschaft. Gepaarte Vergleiche von Allmendweiden und angrenzenden Flächen mit identischen Standortbedingun-gen (Referenzflächen) ermöglichten den Vergleich der Arten bzw. -gruppe anhand von Umwelt-parametern in den Vegetationstypen Grünland auf Mineralboden und offenen Mooren. Der Un-tersuchungsraum in Südbayern stellt den Verbreitungsschwerpunkt für Allmendweiden und intakte bis naturnahe Moorökosysteme in Deutschland dar. Meine Analysen zeigen, dass sich der Landnutzungswandel drastisch auf die Referenzflä-chen, die früher überwiegend Teil der Allmendweiden waren, auswirkte: Im Vergleich zu den Weiden weisen sie (i) homogene Strukturen auf Landschafts- und Habitatebene, (ii) geringe Anteile nährstoffarmer Habitate und (iii) eine geringe Besiedlung von Vögeln und Insekten auf. Somit sind sie weitgehend ungeeignet für den Erhalt der Biodiversität. Die einzige Ausnahme innerhalb der Referenzflächen bilden traditionell genutzte Heu- und Streuwiesen. In den Allmendweiden sorgt die traditionelle Beweidung (0.5–2.0 Großvieheinheiten/ha von Mai–Oktober) für eine hohe strukturelle Vielfalt mit nährstoffarmen Habitaten und fließende Übergänge zwischen Wald und Offenland. So können saisonal bzw. innerhalb des Lebenszyklus wechselnde Ansprüche vieler typischer Agrarlandschaftsarten im eng verzahnten Mosaik wert-voller Habitate erfüllt werden. Der Erhalt der Allmendweiden ist für das Überleben von Agrar-landschaftsarten im Untersuchungsgebiet essenziell. Eine Ausdehnung der Allmendweiden-Nutzung auf angrenzende Flächen ist unbedingt zu empfehlen.

Habitat destruction and modification are major causes cited for the decline of amphibians worldwide (Wake and Morowitz 1991). Depressional wetlands on Pinckney Island NWR in Beaufort County, South Carolina were ditched and drained during the 1950?s for agricultural development. Wetlands were restored by filling ditches with existing spoil. I surveyed herpetofaunal and vegetation communities to determine responses to wetland restoration on Pinckney Island from 2004-2005. I selected ten wetlands each in pine and maritime habitats, sampling prior to and after restoration. I recorded 14 reptile and 9 amphibian species using time-constrained searches and funnel trap arrays. I documented (first record on the Island) Fowler?s toad (Bufo woodhousei) and many-lined salamander (Stereochilus marginatus). There were differences between amphibian species richness recorded using time constrained surveys and funnel trap arrays, amphibian species abundance between pine and maritime forest, and between number of Eastern spadefoot toads (Scaphiopus holbrooki) before and after restoration.

Agri-environment schemes (AESs), ecological focus areas (EFAs), and organic farming are the main tools of the common agricultural policy (CAP) to counteract the dramatic decline of farmland biodiversity in Europe. However, their effectiveness is repeatedly doubted because it seems to vary when measured at the field-versus-landscape level and to depend on the regional environmental and land-use context. Understanding the heterogeneity of their effectiveness is thus crucial to developing management recommendations that maximize their efficacy. Using ensemble species distribution models and spatially explicit field-level information on crops grown, farming practice (organic/conventional), and applied AES/EFA from the Integrated Administration and Control System, we investigated the contributions of five groups of measures (buffer areas, cover crops, extensive grassland management, fallow land, and organic farming) to habitat suitability for 15 farmland bird species in the Mulde River Basin, Germany. We used a multiscale approach to identify the scale of effect of the selected measures. Using simulated land-use scenarios, we further examined how breeding habitat suitability would change if the measures were completely removed and if their adoption by farmers increased to meet conservation-informed targets. Buffer areas, fallow land, and extensive grassland were beneficial measures for most species, but cover crops and organic farming had contrasting effects across species. While different measures acted at different spatial scales, our results highlight the importance of land-use management at the landscape level—at which most measures had the strongest effect. We found that the current level of adoption of the measures delivers only modest gains in breeding habitat suitability. However, habitat suitability improved for the majority of species when the implementation of the measures was increased, suggesting that they could be effective conservation tools if higher adoption levels were reached. The heterogeneity of responses across species and spatial scales indicated that a mix of different measures, applied widely across the agricultural landscape, would likely maximize the benefits for biodiversity. This can only be achieved if the measures in the future CAP will be cooperatively designed in a regionally targeted way to improve their attractiveness for farmers and widen their uptake.

Landwirt:innen prägen Agrarlandschaften und beeinflussen die Biodiversität maßgeblich. Ein tiefgehendes Verständnis ihrer Entscheidungen zu biodiversitätsfreundlichen Maßnahmen (BFFM) ist essenziell für wirksame Biodiversitätsstrategien. Diese sollen nicht nur Anreize schaffen, sondern auch Maßnahmen auf Landschaftsebene koordinieren. Ziel dieser Dissertation ist es, die Faktoren des Biodiversitätsmanagements europäischer Landwirt:innen zu untersuchen, insbesondere ihre Verhaltensweisen, Wahrnehmungen und räumlichen Entscheidungsprozesse. Die Arbeit umfasst drei aufeinander aufbauende Studien: (i) eine systematische Literaturanalyse zu Motiven und Hindernissen für BFFM, (ii) qualitative Interviews zur Wahrnehmung von Biodiversität und deren Schutz sowie (iii) ein räumliches Entscheidungsexperiment zu Präferenzen für die Platzierung linearer Biodiversitätsmaßnahmen. Die Ergebnisse zeigen, dass betriebliche Entscheidungen durch ein komplexes Zusammenspiel verschiedener Faktoren beeinflusst werden. Über wirtschaftliche und praktische Gesichtspunkte hinaus sind diese Entscheidungen tief in Wertvorstellungen und der Wahrnehmung von Biodiversität verwurzelt. Wenngleich die meisten Befragten ihre landwirtschaftlichen Tätigkeiten möglichst wenig beeinträchtigen wollten, erkannten sie die ökologische Relevanz ihrer Arbeit und waren offen für Anpassungen, sofern sich Möglichkeiten zur Vernetzung bestehender Habitate ergaben. Die erheblichen regionalen Unterschiede verdeutlichen die Notwendigkeit, Biodiversitätsstrategien an spezifische sozioökonomische, kulturelle und ökologische Kontexte anzupassen. Die Berücksichtigung intrinsischer Werte und der kulturellen Verbundenheit mit dem Land kann die Akzeptanz und das Engagement fördern. Zudem kann eine gezielte Abschätzung der Bereitschaft zur Habitatvernetzung die Planung ökologischer Netzwerke optimieren und Transaktionskosten senken, indem Wildtierkorridore mit Präferenzen der Landwirt:innen abgestimmt werden. / Farmers are key actors in agricultural landscapes and have a profound impact on biodiversity. Understanding their decisions to adopt biodiversity-friendly farming measures (BFFM) is crucial for developing effective conservation strategies that not only motivate farmers but also coordinate their efforts to achieve greater biodiversity outcomes at the landscape scale. This dissertation aims to explore the factors that influence farmers’ biodiversity management across Europe, integrating behavioural, perceptual, and spatial dimensions of their decision-making processes. Three interrelated studies were conducted: (i) a systematic literature review synthesising evidence on motivations for and barriers to BFFM adoption, (ii) qualitative interviews exploring farmers’ perceptions of biodiversity and its management, and (iii) a spatial choice experiment investigating preferences for the placement of linear biodiversity measures. The studies show that farmers’ biodiversity management decisions are shaped by a complex interplay of factors. Beyond economic and practical concerns, these decisions are deeply rooted in farmers’ values and perceptions of biodiversity. Although most farmers sought to minimise disturbance to their farming activities, they recognised the ecological significance of their practices at the landscape level and were willing to adapt them where opportunities arose to connect habitats. The considerable regional variability indicates that biodiversity conservation strategies need to be tailored to the specific socio-economic, cultural, and environmental contexts. Conservation measures that resonate with farmers’ intrinsic values and cultural connections to the land may increase adoption and commitment. In view of the habitat fragmentation, predicting farmers’ willingness to participate in connectivity measures can inform efficient ecological network planning and reduce transaction costs by aligning potential corridors with their preferences.