Quantifying and modeling processes involved in the global carbon cycle is important to evaluate the temporal and spatial variability of these processes and understand the effect of this variability on future response to changing climate and land use patterns. Biomass accumulation and Net Primary Productivity (NPP) are large components of ecosystem carbon exchange with the atmosphere and thus are the focus of many modeling efforts. When scaling estimates of NPP temporally from days to years and spatially from square meters to landscapes and regions the spatial coherence of these processes through time must be taken into account. Spatial coherence is the degree to which pairs of sites across space are synchronous (i.e., correlated) through time with respect to a given process or variable. In this thesis I determined the spatial coherence of a major component of NPP, tree bole productivity (NPP[subscript B]), and examine how it influences scaling and our ability to predict NPP and forecast change of this flux. In Chapter 2 I developed and tested a method modeling radial tree increment growth from sub-sampled trees and estimating annual site-level biomass accumulation that allows quantification of the uncertainty in these estimates. Results demonstrated that a simple model using the mean and standard deviation of growth increments underestimated bole biomass increment in all three age classes examined by 1% at the largest sample sizes and up to 15% at the smallest sample sizes. The long term average NPP[subscript B] and inter-annual variability were also underestimated by as much as 10% and 22%, respectively. Stratification of trees by size in sampling and modeling methods increased accuracy and precision of estimates markedly. The precision of both models was sufficient to detect patterns of inter-annual variability. To estimate bole biomass accumulation with acceptable levels of accuracy and precision our results suggest sampling at least 64 trees per site, although one site required a sample size of more than 100 trees. In Chapter 3 I compared year to year variability of NPP for tree boles (NPP[subscript B]) for two adjacent small watersheds (second-growth and old-growth) in the western Cascades of Oregon using the methods developed in Chapter 2. Spatial coherence of NPP[subscript B] within and between watersheds was assessed using multivariate analysis techniques. NPP[subscript B] was found to be less coherent between watersheds than within watersheds, indicating decreased spatial coherence with differences in age class and increased spatial scale. However, a larger degree of spatial coherence existed within the old-growth watershed compared to the second-growth watershed, which may be a result of the smaller degree of variation in environmental characteristics in the former watershed. Within a watershed, potential annual direct incident radiation and heat load were more strongly associated with the variation of NPP[subscript B] than climate. Climatic factors correlated with the temporal variation of annual NPP[subscript B] varied between the two watersheds. Results suggest that inter-annual variability and spatial coherence of forest productivity is a result of both internal (e.g., environment and stand dynamics) and external (climate) factors. An unexpected conclusion was that spatial coherence was not consistent and changed through time. Therefore, the coherence of sites over time is not a simple relationship. Instead the patterns of spatial coherence exhibit complex behaviors that have implications for scaling estimates of productivity. This result also indicates that a correlation coefficient alone may not capture the complexity of change through time across space. In Chapter 4 I estimated year to year variation of NPP[subscript B] for eleven sites of varying age, elevation, moisture, and species composition in the Western Cascades of Oregon. Spatial coherence of tree growth within sites and NPP[subscript B] between sites was assessed using Pearson's product-moment correlation coefficient (r). Results suggest that spatial coherence is highly variable between sites (r=-O.18 to 0.92). The second-growth sites exhibited the greatest temporal variability of annual NPP[subscript B] due to the large accumulation of biomass during stand initiation, but old-growth sites exhibited the greatest variation of coherence of NPP[subscript B] between sites. In some years all sites behaved similarly, but for other years some sites were synchronous while others were not. As growth of individual trees and NPP[subscript B] at the site scale increased, inter-annual variability of those variables increased. Climate in part affected annual NPP[subscript B], but intrinsic factors and spatial proximity also affected the coherence between sites in this landscape. / Graduation date: 2006

Forest stands were studied to determine if old-growth forest structure could be mimicked in younger stands via overstory manipulation. Cover and species composition of understory plants were systematically sampled in sixteen thinned second-growth stands and sixteen adjacent unthinned second-growth Douglas fir (Pseudotsuga menziesii (Mirabel) Franco.) stands. The stands were thinned twenty-four to thirteen years ago. These were compared to seven nearby old-growth stands. Thinned and unthinned stands had matching elevations, aspect, and soils, yet differed primarily in management treatment. Leaf area indices were determined for these stands. Thinned stands differed from the old-growth and unthinned stands in having significantly higher cover values and species numbers, apparently resulting from increased light to the forest floor and a greater variety of microhabitats created by thinning. Young unthinned and old-growth stands were comparable in terms of cover and richness, but differed in species composition. Diversity indices showed no difference in species diversity between the three types of stands. Ordination of the species/sample data using Detrended Correspondence Analysis showed that understory species composition of the young unthinned and thinned stands was nearly identical. Species composition of old-growth stands differed from thinned and unthinned stands. The ordination indicated that age of the stands, structure of the canopy layers and climate were major determining factors in the species composition of the understory plant communities. Management manipulation of the second growth stands did not yield stands with understory vegetation communities that mimicked those of old-growth stands. The conclusions of this study were: 1) Shrub cover increased with thinning as compared to unthinned and old-growth stands. 2) Thinning increased the species richness of the stands, without increasing the number of exotics. 3) Diversity was not altered by thinning. Old-growth, thinned and unthinned stands did not differ in diversity values. 4) Patterns of community composition in thinned stands were more similar to unthinned equivalent stands than to nearby old-growth. / Graduation date: 1996

Graduation date: 1993

Eliot Glacier is a small (1.6 km2), relatively well-studied glacier on Mount Hood, Oregon. Since 1901, glacier area decreased from 2.03 ± 0.16 km2 to 1.64 ± 0.05 km2 by 2004, a loss of 19%, and the terminus retreated about 600 m. Mount Hood's glaciers as a whole have lost 34% of their area. During the first part of the 20th century the glacier thinned and retreated, then thickened and advanced between the 1940s and 1960s because of cooler temperatures and increased winter precipitation and has since accelerated its retreat, averaging about 1.0 m a-1 thinning and a 20 m a-1 retreat rate by 2004. Surface velocities at a transverse profile reflect ice thickness over time, reaching a low of 1.4 m a-1 in 1949 before increasing to 6.9 ± 1.7 m a-1 from the 1960s to the 1980s. Velocities have since slowed to about 2.3 m a-1 , about the 1940 speed.

Throughout many of the world’s mountain ranges snowpack accumulates during the winter and into the spring, providing a natural reservoir for water. As this reservoir melts, it fills streams and recharges groundwater for over 1 billion people globally. Despite its importance to water resources, our understanding of the storage capacity of mountain snowpack is incomplete. This partial knowledge limits our abilities to assess the impact that projected climate conditions will have on mountain snowpack and water resources. While understanding the effect of projected climate on mountain snowpack is a global question, it can be best understood at the basin scale. It is at this level that decision makers and water resource managers base their decisions and require a clarified understanding of basin's mountain snowpack. The McKenzie River Basin located in the central-western Cascades of Oregon exhibits characteristics typical of many mountain river systems globally and in the Pacific Northwestern United States. Here snowmelt provides critical water supply for hydropower, agriculture, ecosystems, recreation, and municipalities. While there is a surplus of water in winter, the summer months see flows reach a minimum and the same groups have to compete for a limited supply. Throughout the Pacific Northwestern United States, current analyses and those of projected future climate change impacts show rising temperatures, diminished snowpacks, and declining summertime streamflow. The impacts of climate change on water resources presents new challenges and requires fresh approaches to understanding problems that are only beginning to be recognized. Climate change also presents challenges to decision makers who need new kinds of climate and water information, and will need the scientific research community to help provide improved means of knowledge transfer. This dissertation quantified the basin-wide distribution of snowpack across multiple decades in present and in projected climate conditions, describing a 56% decrease in mountain snowpack with regional projected temperature increases. These results were used to develop a probabilistic understanding of snowpack in projected climates. This section described a significant shift in statistical relations of snowpack. One that would be statistically likely to accumulate every 3 out of 4 years would accumulate in 1 out of 20 years. Finally this research identifies methods to improved knowledge transfer from the research community to water resource professionals. Implementation of these recommendations would enable a more effective means of dissemination to stakeholders and policy makers. While this research focused only on the McKenzie River Basin, it has regional applications. Processes affecting snowpack in the McKenzie River Basin are similar to those in many other maritime, forested Pacific Northwest watersheds. The framework of this research could also be applied to regions outside of the Pacific Northwestern United States to gain a similar level of understanding of climate impacts on mountain snowpack. / Graduation date: 2012

Although nearshore macroinvertebrates are integral members of high mountain lentic systems, knowledge of ecological factors influencing their distributions is limited. Factors affecting distributions of nearshore macroinvertebrates were investigated, including microhabitat use and vertebrate predation, in the oligotrophic lakes of North Cascades National Park Service Complex, Washington, USA, and the conformity of distribution with a lake classification system was assessed (Lomnicky, unpublished manuscript; Liss et al. 1991). Forty-one lakes were assigned to six classification categories based on vegetation zone (forest, subalpine, alpine), elevation, and position relative to the west or east side of the crest of the Cascade Range. These classification variables represented fundamental characteristics of the terrestrial environment that indirectly reflected geology and climate. This geoclimatic perspective provided a broad, integrative framework for expressing the physical environment of lakes. Habitat conditions and macroinvertebrate distributions in study lakes were studied from 1989 through 1991. Distributions varied according to vegetation zone, elevation, and crest position, and reflected the concordance between habitat conditions and organism life history requirements. Habitat parameters affecting distributions included water temperature, the kinds of substrates in benthic microhabitats, water chemistry, and, to a limited extent, the presence of vertebrate predators. The number of taxa per lake was positively correlated with maximum temperature and negatively correlated with elevation. Forest zone lakes tended to have the highest number of taxa and alpine lakes the lowest. Substrates in nearshore microhabitats varied with vegetation zone. Organic substrates were more predominant than inorganic substrates in forest zone lakes. Organic substrates declined and inorganic substrates increased in the subalpine zone. There were virtually no organic substrates in alpine lakes. Taxa were placed into groups based on substrate preference. Ordinations indicated that the proportion of taxa in inorganic and organic-based substrate preference groups paralleled vegetation zone-substrate relationships. Lake water hardness and pH, as well as the presence of vertebrate predators affected the distribution of several taxa. Gastropods were limited to three forest lakes by their water hardness and pH requirements, and the dytiscid beetle, Potamonectes qriseostriatus appeared to be absent from forest lakes due, in part, to the pH requirements of this taxon. The distribution of three taxa (Taenionema, Ameletus, Desmona) appeared to be affected by the presence of vertebrate predators (salamanders and trout). Discriminant analysis was used to test the reliability of lake classification based on terrestrial characteristics. Discriminant analysis assigned lakes to categories based on similarities in kinds of substrates, substrate preference groups, and taxa. Strong concordance between both methods of lake classification supported the interconnection between terrestrial characteristics and processes and the abiotic and biotic conditions in lakes. / Graduation date: 1994

The structure and composition of mixed-conifer forest (MCF) in central Oregon has been altered by fire exclusion and logging. The resulting increased density, spatial contagion, and loss of fire resistant trees decrease the resiliency of this ecosystem to fire, drought, and insects. The historical and current composition and structure of MCF are characterized by steep environmental gradients and a complex mixed-severity fire regime. This inherent variation makes it difficult to determine the magnitude of anthropogenic effects and set objectives for restoration and management. As a result, there is a lack of consensus regarding how MCF should be managed and restored across the landscape. My primary research objectives were to: (1) Characterize the current structure and composition of MCF and how these vary with environmental setting; and (2) Characterize establishment and tree growth patterns in MCF in different environmental settings. To address these objectives, I collected field data on structure and composition and increment cores across a range of environmental conditions in MCF of the eastern Cascades and Ochoco Mountains. I used cluster analysis to identify four stand types based on structure and composition in the eastern Cascades study area and four analogous types in the Ochoco Mountains study area. Variation in understory composition and the presence of large diameter shade tolerant species distinguish each type. Stand types occupied distinct environmental settings along a climatic gradient of increasing precipitation and elevation. At relatively dry PIPO sites understories were dominated by ponderosa pine. At wetter PIPO/PSME and PIPO ABGC sites understories were dominated by shade tolerant species, but ponderosa pine was dominant in the overstory. At the coolest and wettest PIPO/PSME/ABGC sites understories were dominated by grand fir and shade tolerant species were common in the overstory. In the eastern Cascades current density of all live trees and snags was 432, 461, 570, 372 trees per hectare (TPH) for the four stand types identified. Stand types in the drier Ochoco Mountains were currently less dense at 279, 304, 212, and 307 TPH. Current MCF densities in both areas are 2-3 times higher than densities estimated for the late 19th and early 20th centuries from other studies in those two areas. Reconstruction of cuts in each stand type indicates that the density of large diameter ponderosa pine has been reduced by approximately 50% in all stand types in both study regions. Age histograms demonstrate that current density and composition of MCF stand types is a product of abrupt increases in tree establishment following fire exclusion in the late 19th century. The number of trees established increased after 1900 in all stand types, but the timing and composition of changes in establishment varied with climate. At dry PIPO sites increases in establishment were delayed until the 1920s and 1930s and were composed of ponderosa pine. At PIPO/PSME and PIPO/ABGC sites with intermediate precipitation, establishment was dominated by ponderosa pine prior to 1900, but after 1900 establishment was dominated by a large pulse of Douglas-fir and grand fir. At the wettest PIPO/PSME/ABGC there was less evidence of changes in structure and composition over time. My results indicate that compared to dry pine and dry-mixed conifer sites, relatively productive moist mixed-conifer sites were characterized by large changes in structure and composition. Such sites could be considered more ecologically altered by lack of fire than drier forest types that had high fire frequencies but slower rates of stand development and less plant community change. Radial growth patterns of cored ponderosa pines differed between the eastern Cascades and Ochoco Mountains. In the eastern Cascades mean growth rates and variance decreased during favorable climatic periods after 1900. This is likely related to increased competition, and provides evidence that current stand density lacks a temporal analog in the 18th and 19th centuries. Sensitivity of growth to climate and harvest suggest competition for water in the denser forest of the eastern Cascades, and indicates thinning will increase the diameter growth rate of large old pines. In the Ochoco Mountains, ponderosa pine tree growth was less responsive to climate prior to fire exclusion in the late 1800s, and growth did not respond to fire events. This suggests competition among trees was historically low in this region. After fire exclusion growth became more responsive to wet and dry climatic cycles, which may indicate that increased density and competition made trees more responsive to climate variability. Patterns of slow and fast growth appeared to differ between study regions and likely differ at the sub-regional scale. Further analysis of the relationship between growth and climate in different environmental settings is needed to distinguish where stand development has been modified by disruption of fire regimes. / Graduation date: 2013

Typescript. Includes vita and abstract. Includes bibliographical references (leaves 155-177). Also available online in Scholars' Bank; and in ProQuest, free to University of Oregon users.

Silicic volcanism in the central Oregon Cascade range has decreased in both the size and frequency of eruptions from its initiation at ~40 Ma to present. The reasons for this reduction in silicic volcanism are poorly constrained. Studies of the petrogenesis of these magmas have the potential for addressing this question by providing insight into the processes responsible for producing and erupting silicic magmas. This study focuses on two extensive and well-preserved ash-flow tuffs from within the ~4-8 Ma Deschutes Formation of central Oregon, which formed after the transition from Western Cascade volcanism to the modern High Cascade. Documentation of outcrop extent, outcrop thickness, clast properties, and samples provide the means to estimate a source location, minimum erupted volumes, and to constrain eruptive processes. Major and trace element chemistry of glass and minerals constrain the petrogenesis and chemical evolution of the system. The tuffs selected for this study, the Lower Bridge and McKenzie Canyon, are the first known silicic units originating from the Cascade Arc following the reorganization from Western Cascade to High Cascade Volcanism at ~8 Ma. These eruptions were significant in producing a minimum of ~5 km�� DRE each within a relatively short timeframe. These tuffs are sourced from some vent or edifices related to the Three Sisters Volcanic Complex, and capture an early phase of the volcanic history of that region. The chemical composition of the tuffs indicates that the Lower Bridge erupted predominately rhyolitic magma with dacitic magma occurring only in small quantities in the latest stage of the eruption while McKenzie Canyon Tuff erupted first as a rhyolite and transitioned to a basaltic andesite with co-mingling and incomplete mixing of the two magma types. Major and trace element concentrations in minerals and glass indicate that the basaltic andesite and rhyolite of the McKenzie Canyon Tuff were well convected and stored in separate chambers. Geothermometry of the magmas indicate that the rhyolites are considerably warmer (~850��) than typical arc rhyolites. Trace element compositions indicate that both the Lower Bridge and McKenzie Canyon Tuff experienced mixing between a mantle derived basaltic melt and a rhyolitic partial melt derived from gabbroic crust. Rhyolites of the Lower Bridge Tuff incorporate 30-50% partial melt following 0->60% fractionation of mantle derived melts. The McKenzie Canyon Tuff incorporates 50-100% of a partial melt of a mafic crust with up to 15% post mixing fractionation. The results of this study suggest that production of voluminous silicic magmas within the Cascade Arc crust requires both fractionation of incoming melts from the mantle together with mixing with partial melts of the crust. This provides a potential explanation for the decrease in silicic melt production rates from the Western Cascades to the High Cascades related to declining subduction rate. As convergence along the Cascade margin became more oblique during the Neogene, the consequent slowing rate of mantle melt production will result in a net cooling of the crust, inhibiting the production of rhyolitic partial melts. Without these partial melts to provide the rhyolitic end member to the system, the system will evolve to the mafic melt and fractionation dominated regime that has existed along Cascadia throughout the Quaternary. / Graduation date: 2013

The fisher (Martes pennanti) is a medium sized member of the mustelid family that once roamed the forests of Washington and whose historic range in the western United States once spread throughout the northern Rocky Mountains, the Cascade and Coast Ranges, and the Sierra Nevada (Carroll, Zielinski, and Noss 1999; Powell 1993, Spencer et al. 2011). Due to pressures from trapping and habitat fragmentation, the abundance of the species in the western United States has decreased dramatically and is thought to be limited to several small, isolated populations. In 2008, fishers were reintroduced to the Olympic Peninsula; however, bobcat (Lynx rufus) predation in the first years is thought to have killed off a significant portion of the released fisher hindering their ability to establish a self-sustaining population (Lewis et al. 2011). Other studies in the western United States have shown that bobcats can be a dramatic force on small or isolated fisher populations. The coniferous forest of the southern Washington Cascades is the possible site of a release of currently extirpated fishers. My research examines the distribution of bobcats in the region and explores the implication this and the habitat variables of the area have for a future reintroduction of fisher. The workflow of the research was a stepwise process of: 1) surveying forested areas in the southern Washington Cascades for the presence and absence of bobcat and acquiring previously completed survey data 2) using a classification tree to model the correlation of bobcat presence or absence with forest variables and 3) applying these relationships to spatial analysis the creation of maps showing areas of high ranking fisher habitat. The classification tree modeled the correlation between the forest variables and the results of the surveys, which included 145 bobcat absence observations and 39 presence observations. The model highlighted a 95% probability of absence above 1,303 m in elevation, 73% probability of absence in areas under 1,303 m in elevation and with a tree diameter value under 43.45 cm, 57% probability of absence in areas between 1,070 m and 1,303 m in elevation and with a tree diameter value above 43.45 cm, and an 89% probability of bobcat presence in areas under 1,070 m in elevation with a tree diameter value above 43.45 cm. I applied an upper elevation limit of 1,676 meters as a threshold for suitable habitat and only considered habitat suitable in cells with a tree diameter above 29 cm. The three locations highlighted as the most suitable areas for reintroduction due to a large amount of the highest ranking habitat and the largest aggregations of suitable habitat cells were around the William O. Douglas Wilderness that straddles the border of the Gifford Pinchot National Forest (GPNF) and the Wenatchee National Forest, another location in the Norse Peak Wilderness northeast of Mount Rainier, and a third location in Indian Heaven Wilderness in the southern portion of the GPNF.