This thesis presents results from spatial-temporal and volumetric change analysis of blowouts on the Cape Cod National Seashore (CCNS) landscape in Massachusetts, USA. The purpose of this study is to use methods of analysing areal and volumetric changes in coastal dunes, specifically blowouts, and to detect patterns of change in order to contribute to the knowledge and literature on blowout evolution. In Chapter 2.0, the quantitative analysis of blowout change patterns in CCNS was examined at a landscape scale using Spatial-Temporal Analysis of Moving Polygons (STAMP). STAMP runs as an ArcGIS plugin and uses neighbouring year polygon layers of our digitized blowouts from sequential air photo and LiDAR data (1985, 1994, 2000, 2005, 2009, 2011, and 2012 for 30 erosional features, and 1998, 2000, 2007, and 2010 for 10 depositional features). The results from STAMP and the additional computations provided the following information on the evolution of blowouts: (1) both geometric and movement events occur on CCNS; (2) generation of blowouts in CCNS is greatest in 1985 and is potentially related to vegetation planting campaigns by the Park; (3) features are expanding towards dominant winds from the North West and the South West; (5) the erosional and depositional features are becoming more circular as they develop, (6) the evolution of CCNS blowouts follows a similar pattern to Gares and Nordstrom’s (1995) model with two additional stages: merging or dividing, and re-activation. In Chapter 3.0, the quantitative analysis of volumetric and areal change of 10 blowouts in CCNS at a landscape scale is examined using airborne LiDAR and air photos. The DEMs of neighbouring years (1998, 2000, 2007, and 2010) were differenced using Geomorphic Change Detection (GCD) software. Areal change was detected by differencing the area of polygons that were manually digitized in ArcGIS. The changes in wind data and vegetation cover were also examined. The results from the GCD and areal change analysis provide the following information on blowout evolution: (1) blowouts generate/initiate; (2) multiple blowouts can merge into an often larger blowout; (3) and blowouts can experience volumetric change with minimal aerial change and vice versa. From the analyzes of hourly Provincetown wind data (1998-2010), it was evident that blowouts developed within all three time intervals. The percentages of comparable winds (above 9.6 m s-1) were highest in 1998, 1999, 2007 and 2010. It is speculated that tropical storms and nor'easters are important drivers in the development of CCNS blowouts. In addition, supervised classifications were run on sequential air photos (1985 to 2009) to analyze vegetation cover. The results indicated an increase in vegetation cover and decrease of active sands over time. Two potential explanations that link increased vegetation to blowout development are: (1) sparse vegetation creates a more conducive environment for the initiation of blowouts by providing stability for the lateral walls, and (2) high wind events (e.g. hurricanes and nor'easters) could cause vegetation removal, allowing for areas of exposed sand for blowout initiation and development. / Graduate / 0799 / 0368 / kimia.abhar@gmail.com

Thesis (Ph. D.)--University of Chicago, 1911. / "Reprinted from the Botanical gazette, Vol. LIV, No. 4, 1912." "Literature cited": p. 304-305. Also available on the Internet.

As part of the Bagnold Dune campaign conducted by Mars Science Laboratory rover Curiosity, visible/near-infrared reflectance spectra of dune sands were acquired using Mast Camera (Mastcam) multispectral imaging (445-1013nm) and Chemistry and Camera (ChemCam) passive point spectroscopy (400-840nm). By comparing spectra from pristine and rover-disturbed ripple crests and troughs within the dune field, and through analysis of sieved grain size fractions, constraints on mineral segregation from grain sorting could be determined. In general, the dune areas exhibited low relative reflectance, a weak similar to 530nm absorption band, an absorption band near 620nm, and a spectral downturn after similar to 685nm consistent with olivine-bearing sands. The finest grain size fractions occurred within ripple troughs and in the subsurface and typically exhibited the strongest similar to 530nm bands, highest relative reflectances, and weakest red/near-infrared ratios, consistent with a combination of crystalline and amorphous ferric materials. Coarser-grained samples were the darkest and bluest and exhibited weaker similar to 530nm bands, lower relative reflectances, and stronger downturns in the near-infrared, consistent with greater proportions of mafic minerals such as olivine and pyroxene. These grains were typically segregated along ripple crests and among the upper surfaces of grain flows in disturbed sands. Sieved dune sands exhibited progressive decreases in reflectance with increasing grain size, as observed in laboratory spectra of olivine size separates. The continuum of spectral features observed between the coarse- and fine-grained dune sands suggests that mafic grains, ferric materials, and air fall dust mix in variable proportions depending on aeolian activity and grain sorting.

Ecological evaluation techniques have been used to classify or evaluate the relative conservation worthiness of a portion of the Eastern Cape coastal zone from Cannon Rocks to Port Alfred. A system-based approach, using a basic GIS, was used to compare three aspects of the coastal zone. Two existing planning documents (EMATEK/s Coastal Vegetation Importance Rating plan and CPA's Cape Coastal Conservation plan) were statistically compared - based primarily on the size and number of the landscape fragments (environmental criteria polygons). The resultant plan, an Intermediate Sensitivity Class map, was further integrated with the biophysical aspect of the coastal zone - coastal dunes. The final map is the product of these overlays to be used as a-working overlay for coastal zone decision-making. This is an Environmental Sensitivity map, which has no particular direct reference to any particular biophysical criteria, which categorizes the coastal zone according to the environmental sensitivity or conservation worthiness of the site. The categories are listed according to a sliding-scale of sensitivity or worthiness. There is, however, it strong association between the final map and the vegetation status of the coastline, as a direct result of the vegetation rating map and indirectly from the perception based coastal conservation plan. The formation of most assessment or evaluation plans are based on the vegetation status of the area. This map can be overlayed over the particular stretch of the coastline which it covers and the particular conservation status noted. There are numerous techniques available for assessment and evaluation each having their own specific merits and advantages. This plan, however, is a first attempt at integrating other specific plans into a single working document for the coastal zone manager.

Arbuscular mycorrhizas (AM) are mutually beneficial symbiotic associations between the roots of plants and certain Zygomycetous fungi. The role of AM fungi in coastal sand dunes has been explored in many parts of the world, though little work has been conducted in South African dune systems. This study aimed to investigate the presence and extent of mycorrhizal colonisation of a coastal sand dune in South Africa. The roots of five plant species (Scaevola plumieri, Arctotheca populifolia, Ipomoea pes-caprae, Ehrharta villosa and Chrysanthemoides monilifera) were sampled along a foredune profile at Old Woman's River in the Eastern Cape. These roots were assessed for the percentage mycorrhizal colonisation they supported. Spores extracted from the rhizosphere sand of each plant species were counted and identified to genus level. Results were related to seasonality and the position of the plants along the profile. All plant species were found to be mycorrhizal. Percentage colonisation ranged from 0-92%, depending on plant species and season. Mycorrhizal colonisation was generally highest in the winter months, and especially so in I pes-caprae and E. villosa. The extent of various mycorrhizal structures in root tissue varied between plant species. Spore numbers ranged from 0-48 spores 100g-1 sand with highest numbers occurring in winter. S. plumieri and A. populifolia were associated with greatest spore abundance. Four fungal genera (Glomus, Acaulospora, Scutellospora and Gigaspora) were identified. Distribution of these genera showed seasonal variations between plant species. A bioassay, using Sorghum, was conducted to test the inoculum potentials of sand from the Scaevola hummock and the IpomoealEhrharta dune. Highest percentage colonisation occurred in plants grown in the Scaevola sand, which also had the lowest root and shoot measurements. The bioassay confirmed that AM propagules are present and viable, even in the mobile sand of the foredune. This study showed that mycorrhizal colonisation and spore numbers varied seasonally, but that the extent of this was dependent on plant species. The position of plants along the foredune profile tended to be less important than plant species. It is thought that the growth cycle and rooting system of each plant species determines seasonal cycles and abundance of AM fungi. Variation within fungal populations probably also impacts on this. Knowledge of the presence and distribution of AM fungi in this system paves the way for more detailed studies which need to examine the role of these endophytes in South African sand dunes.

Natural and anthropogenic environmental variables pressures exert fitness pressures on our flora and vegetation and necessitate an understanding of the mechanisms of plant responses to those pressures. Overgrazing and desertification are examples of the stressors that affect plant community composition, on both large and small scales. In order to investigate the plant community response to these stressors it has been shown that it is more practical and informative to group plant species according to their functional characteristics into functional types than attempting to predict plant response solely based on previous range distribution data. The present study involved a survey of the dunefield vegetation of several privately owned farms as well as a communal farm in the southern Kalahari. The farms were subjectively chosen to represent a gradient from underutilized to overutilized. Plant community surveys were conducted on the farms and the dominant plant species were sampled and researched for selected functional traits to explore possible relationships between traits and grazing pressure. Ordination techniques were used to identify plant functional grazing response types. The identification of plant functional types through easy measurable traits will be a great aid in modelling and longterm environmental monitoring approaches. Of the three dunefield habitats, the dune street habitat was identified as the most vulnerable to overgrazing. A reduction in cumulative cover and an increase in bare surface cover indicated increased grazing pressure in the dune crest habitat. In the dune streets shrub cover and annual grass cover increased when veld was overgrazed. Functional analysis of 112 Kalahari plant species for a set of 47 plant traits showed that species clustering were strongly related to vegetative characteristics. The emerging clusters were strongly associated with major life forms, which comprised phanerophytes of different size classes, geophytes and herbaceous perennials, as well as herbaceous annuals. These clusters were identifiable even with the inclusion of reproductive traits. Selected vegetative traits are good indicators of plant response to land-use intensity since these traits lend competitive advantages to plant persistence under utilisation pressure. Plant height, spinescence, specific leaf area, leaf dry matter content, leaf sclerophylly and stem density were found to be examples of such traits. Phanerophyte and chamaephyte dominated functional groups were mostly associated with lower range condition and hemicryptophyte dominated groups with higher range condition, whereas herbaceous therophyte dominated groups did not show any specific association. / Dissertation (MSc)--University of Pretoria, 2011. / Plant Science / unrestricted

Coastal dune vegetation plays a key role in dune formation and stabilization through sediment trapping and erosion control. To restore degraded dunes, revegetation of dune building species is critical. Planting density has been found to effect growth of marsh species, with closer plantings alleviating stress through facilitation. As coastal dunes are high stress environments, it is expected that dune species may also exhibit facilitative interactions based on the Stress Gradient Hypothesis. Therefore, planting grasses in clumped configurations may lead to more successful dune revegetation. The objective of this research was to determine how planting density affects the growth of two dominant dune grasses along the US Atlantic coast, Ammophila breviligulata and Uniola paniculata, through field surveys of natural distribution, density, and a manipulation study of planting densities. Natural distribution differed between the two species with A. breviligulata occurring at lower dunes and U. paniculata occurring at higher dunes. Ammophila breviligulata occurred more densely than U. paniculata. Planting density of U. paniculata had an effect on growth parameters (shoot length, stem number, and ramet number) but not survival with dispersed plantings (50 cm apart) having higher growth than clumped plantings. The effect of density planting on growth parameters may impact dune building processes. Sparsely planted U. paniculata may have greater sediment capture compared to densely planted U. paniculata due to greater stem number and biomass resulting in taller, steeper dunes.

Ecological restoration has the potential to stem the tide of habitat loss, fragmentation and transformation that are the main threats to global biological diversity and ecosystem services. Through this thesis, I aimed to evaluate the ecological consequences of a 33 year old rehabilitation programme for coastal dune forest conservation. The mining company Richards Bay Minerals (RBM) initiated what is now the longest running rehabilitation programme in South Africa in 1977. Management of the rehabilitation process is founded upon the principles of ecological succession after ameliorating the mine tailings to accelerate initial colonisation. Many factors may detract from the predictability of the ecological succession. For example, if historical contingency is a reality, then the goal of restoring a particular habitat to its former state may be unattainable as a number of alternative stable states can result from the order by which species establish. Succession appears to be a suitable conceptual basis (at this stage in regeneration at least) for the restoration of coastal dune forest. Patterns of community characteristics observed in rehabilitating coastal dune forest sites were similar to those predicted by ecological succession, with few exceptions. Changes in the species pool such as the establishment of strong dominants may lead to divergence of regenerating trajectories away from the desired endpoints. The species composition of herbaceous plants in regenerating coastal dune forest sites became increasingly uniform as the time since disturbance increased. Despite initially becoming more similar they II deviated away from an undisturbed reference site. Contrary to our expectations, non-native species did not contribute the most to dissimilarity. The deviation from the reference forest is attributable to the higher abundance of a native forest specialist in the reference site and the higher abundances of native woodland adapted species in the rehabilitating sites. Changes in the disturbance regime under which species have evolved may lead to arrested succession. The rehabilitation of coastal dune forest relies on the Acacia karroo successional pathway which, has been criticised because Acacia dominated woodlands may stagnate succession. The patterns of species composition within regenerating coastal dune forest are a response to the canopy characteristics and represent an early stage in forest succession. Succession did not appear to be stagnant. Ecological succession does not pay much heed to the role that the surrounding landscape composition can play in the assembly of communities. The theory of Island biogeography provides predictions about how landscape composition influences community assembly. Landscape spatial parameters, measuring edge, isolation, and area explained the patch occupancy of the several bird and tree species, however, responses to patch characteristics were varied and idiosyncratic. For restoration to succeed, managers need to consider the spatial configuration of the landscape to facilitate colonization of rehabilitating patches. From this thesis and previous work, it appears that processes are in place that will lead to the reassembly of dune forest communities. As the rehabilitating sites are at an early stage of regeneration this may take some time to give rise to these coastal dune forest communities, and the management of rehabilitating coastal dune forest must allow for this. In addition, it is III important to remember that time may be interacting with the landscapes spatial attributes, which may limit the presence of certain species. / Thesis (PhD)--University of Pretoria, 2011. / Zoology and Entomology / unrestricted

I Dune: Messiah (1969) sker en förändring av klimatet, en terraformning som människan initierar. Vatten introduceras i rikliga mängder på planeten Arrakis och detta påverkar planetens ekologi såväl som människans kultur. Uppsatsen granskar Frank Herberts roman genom att teoretiskt utgå ifrån ett ekokritiskt perspektiv och tematiskt ifrån vatten, dess varande och relation till människan samt planeten. Människans medvetenhet av sig själv som en art på planeten och sin kulturella koppling till både planeten och vatten är centralt i analysen. Klimatförändringen som sker när människan terraformar planeten och hur detta påverkar människan behandlas också. Även postkolonialism väger in i analysen.  I Dune: Messiah är vattnet, människan och planeten starkt kopplade till varandra. Människan utgår från sin antropocentriska världssyn och betraktar planeten som en plats vilken kan förbättras till människans fördel. Eftersom fremenfolkets kultur är djupt grundad i naturen skapar terraformningen av planeten en identitetskris. Kulturen blir kopplingen som får människan att vilja bevara planeten som den är. Vattnets värde kulturellt men även som valuta gör den till en maktskapande vara. För människan är terraformningen en strävan mot förbättring, vilket speglar de teman av stora strukturer som finns genomgående i Dune-serien. Den här långsiktiga visionen som Frank Herbert målar upp, avbildar människan med en större medvetenhet, en möjlighet att tänka stort. Jag menar (likt andra inom ekokritiken) att detta större tänkande är en väg framåt som ger mänskligheten ett större perspektiv – därmed förståelse av – ekologiska förhållanden. / In Dune: Messiah (1969) the climate changes due to terraforming caused by humans. Water is introduced on the planet Arrakis, and this affects the planet’s ecology as well as the people’s culture. This paper is an exploration of Frank Herbert’s novel, theoretically anchored in ecocriticism and thematically in water, its nature of existence, relationship to humankind and the planet. Humanity’s awareness of themselves as a species on the planet, and their cultural connection to both the planet and water is central to the analysis. The climate-change which occurs when humanity terraforms the planet and how this affects humankind is also touched upon. Post-colonialism also plays a role in the analysis. Water, people, and the planet are strongly connected to each other in Dune: Messiah. Humans see the planet as a place which can be improved to their own benefit due to their anthropocentric worldview. Because the Fremen culture is deeply rooted in nature, an identity-crisis occurs when the planet is terraformed. It is culture that enables humans to want to preserve the planet as it is. Water is a resource that creates power due to its cultural and economical value. The terraforming is humanity aiming for improvement, which mirrors the themes  of larger structures that can be found continuously in the Dune-series. This long-term vision which Herbert paints, pictures humans with a broader consciousness, a capability to think in a larger magnitude. This I (as others within ecocriticism), consider to be a path forwards which offers a broader perspective – therefore an understanding of – ecological conditions.

Stability of coastal systems are threatened by oceanic and atmospheric drivers of climate change. Sea-level rise compounded with increased frequency and intensity of storms emphasizes need for protection of inner island systems by dune formations. Dune building processes are affected by interactions between growth of ecosystem engineering dune grasses and environmental factors associated with disturbance such as sand burial and salt spray. Climate change may also cause latitudinal expansion of some species, resulting in emergence of competitive interactions that were previously absent. Topographic structure of coastlines, traditionally influenced by sand burial, could change as a result of competition emergence. My goal was to determine if species functional trait responses to common abiotic factors are altered by novel and current biotic interactions. I performed a multi-factorial greenhouse experiment by planting three common dune grasses (Ammophila breviligulata, Uniola paniculata, and Spartina patens) in different biotic combinations, using sand burial and salt spray as abiotic stressors. I hypothesized that biotic interactions will cause these dune grasses to shift functional trait responses to abiotic factors that are associated with dune building. I found that plants consistently decreased in biomass when buried. I also found that competition between A. breviligulata and U. paniculata negatively affected dune building function traits of A. breviligulata. This indicates that competition with U. paniculata could alter dune structure. In comparison A. breviligulata had a positive interaction with S. patens, which increased functional trait responses to abiotic stress. Last, we found that competitive intransitivity could occur between these species. My results can be used to make predictions on cross-scale consequences of novel competitive events. This experiment also provides evidence that consideration of local biotic interactions is important in understanding connections between plant level dynamics and large-scale landscape patterns in high stress environment.