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The end-Triassic Mass Extinction (ETME) was one of the "Big 5" mass extinctions of the Phanerozoic and is thought to have been caused by a series of environmental changes triggered by the emplacement of the Central Atlantic Magmatic Province (CAMP). While the overall driver of the ETME is well-accepted, the specific roles of subsequent environmental changes in driving the extinction remain unresolved due to both spatial and temporal biases within existing geochemical records. Additionally, recent studies suggest that environmental deterioration may have preceded the emplacement of both CAMP and the ETME. Here, we present nitrogen isotope (δ15N) and iron speciation data from a sedimentary succession located in Grotto Creek, Alaska in order track changes in the marine nitrogen cycle and redox across the upper Norian Stage of the Triassic through lower Hettangain Stage of the Jurassic (~215 to 198 million years ago). The geologic succession at Grotto Creek represents deeper water marine deposition in eastern equatorial Panthalassa, likely capturing regional oceanic redox trends. δ15N records from Grotto Creek show a pronounced positive excursion of +3‰ that initiates in the upper Norian, peaks near the Norian-Rhaetian boundary (NRB; ~208.5), and declines through the ETME and into the Hettangian. Throughout this interval, iron speciation data show persistent anoxic conditions and occasional euxinia occurred in the local bottom waters. We propose that the positive δ15N and iron speciation data from Grotto Creek reflect a progressive deoxygenation of the upper water column with the expansion of the equatorial oxygen minimum zone (OMZ). Specifically, the δ15N excursion reflects a shift in the dominant processing of bioavailable nitrogen from uptake and nitrification under oxic conditions to incomplete denitrification followed by complete denitrification as deoxygenation progressed. Records from existing study sites in other regions of the Panthalassa reflect similar redox fluctuations and support that this deoxygenation was a regional phenomenon. Other recent studies of the Late Norian and Rhaetian also document declines in biodiversity that predate the ETME, as well as, carbon isotope excursions, likely reflecting perturbations to the carbon cycle. Together, this evidence suggests that increasingly low-oxygen conditions initiated at least ~8 Ma before the emplacement of CAMP and likely paved the way for the ETME. / Master of Science / The end-Triassic mass extinction (ETME) occurred ~ 201 million years ago and was associated with a large drop in global diversity. The extinction is thought to be due to a warming climate resulting from greenhouse gasses released during the eruption of the Central Atlantic Magmatic Province (CAMP). Although the ETME is one of the largest extinction events of the last ~500 million years, the environmental changes leading to the extinction, and before the CAMP eruption, are not well understood. We studied the Grotto Creek section in Alaska which is a sequence of sedimentary rocks that record the deep ocean environment from the ancient Panthalassa or Paleo-Pacific Ocean. This section also provides a relatively complete record of the time before, during, and after the ETME. Here, we present the stable isotopes of nitrogen (δ15N) from these rocks that show disturbances to the nitrogen cycling that occurred in the ocean more than ~ 8 million years before the ETME. We show that δ15N and iron data from Grotto Creek document falling ocean oxygen levels before the ETME. We also find that our δ15N agrees with other environmental proxies that also reflect declining oxygen levels in the oceans along with decreases in marine animal diversity well before the ETME. These geochemical records help to show a more complete picture of how environments changed in the Panthalassa before the ETME

Two experiments were carried out. The first study was designed to investigate the effects of percentage of reinforcement and the number of correct acquisition trials, when combined, on the number of responses to extinction and rate of lever-pulling of human subjects. The second study was designed to investigate the effects off percentage of reinforcement combined with number of reinforcements on the same dependent variables. Analysis of the data revealed: (1) resistance to extinction in both experiments, was inversely related to the percentage of reinforcement; (2) resistance to extinction was inversely related to the number of reinforcements but was not affected by the number of correct acquisition trials; (3) the mean rate of responding was inversely related to the number of reinforcements but was not affected by either percentage of reinforcement or number of correct acquisition trials; and (4) the number of responses per unit of time increased during acquisition and decreased during extinction in both studies. The results were discussed in relation to theories stemming from previous partial reinforcement studies. / Thesis / Master of Arts (MA)

The effects of intensity of the unconditioned stimulus (US) on the acquisition and extinction of the conditioned emotional response (CER) in rats were investigated. The US intensities studied were 0.28, 0.49, 0.85, 1.55, and 2.91 ma. Both acquisition and extinction of the CER were found to be monotonic functions of US intensity, with the higher US intensities producing more rapid acquisition and more resistance to extinction. The lowest shock intensity failed to produce suppression. The 0.49 ma. subjects typically showed a partial recovery of normal operant behavior after development of a fairly profound CER. The results were interpreted as consistent with the supposition that the CER is acquired in accordance with Pavlovian laws of classical conditioning. / Thesis / Master of Arts (MA)

The Upper Devonian of the central-southern Appalachians Valley and Ridge province of Virginia lacks stratigraphic resolution, revised formal nomenclature, and detailed biostratigraphic data. Eight of the most complete sections available in a three thousand square mile area were used to build a framework for revising the stratigraphy of the Upper Devonian strata in southwestern Virginia. Detailed lithologic descriptions of about four thousand feet (1.3 km) of rock were made at each outcrop. John Dennison's (1970 and 1976) nomenclature for the Upper Devonian along the Alleghany Front was successfully tested for it usefulness in Southwestern Virginia and are hereby applied to these rocks. The stratigraphic interval ranges in age from the Middle Devonian to the Lower Carboniferous. The stratigraphic units include the Middle Devonian Millboro Shale, the Upper Devonian Brallier, Scherr, Foreknobs (formally the "Chemung"), Hampshire, and the Lower Carboniferous Price Formation. The Brallier contains two members (Back Creek Siltstone and Minnehaha Springs), the Foreknobs is divided into five members (Mallow, Briery Gap, Blizzard, Pound, and Red Lick), and the lower Price is divided into three members (the Cloyd Conglomerate, Sunbury Shale, and the Ceres). 23046 fossils were collected and 160 taxa were identified. The biostratigraphic range of each taxon was compiled, analyzed, and then divided into biostratigraphic zones. 19 local biozones are described. The Frasnian/Famennian boundary is accurately placed based on occurrences of internationally known index fossils. The Frasnian/Famennian extinction event is recognized and is determined to have two pulses. The local biostratigraphic zonations doubled the resolution of previous studies. Lithostratigraphic and biostratigraphic data were combined to look at the timing and rates of events. The lithostratigrapic divisions were tested to see if their boundaries are or are not time transgressive. The Brallier/Foreknobs, Blizzard/Pound, and Pound/Redlick boundaries are crossed by biozones. Conversely the Frasnian/Famennian boundary crosses the lithologic boundary between the Pound and Red Lick Members. This documents the prograding nature of the clastic wedge. Composite biostratigraphic ranges correlate with ranges in New York and western Maryland. This detailed lithostraigraphic and biostratigraphic study documents a comprehensive and higher resolution understanding of the Upper Devonian in the Central-Southern Appalachian Basin. / Ph. D.

Three groups of rats were trained and extinguished with continuous (CRF) and partial (PR) reinforcement in the presence of two auditory stimuli differing in amount of similarity along a frequency dimension. Results revealed that in the groups where stimulus similarity was least, a within-subjects reversed partial reinforcement extinction effect (PREE) was found. The results also suggest that as similarity between the stimuli associated with CRF and PR decreased, the magnitude of the reversed PREE increased. This result is in contrast to Rashotte's (1971) predictions concerning the type of PREE produced as stimulus similarity varies along a single dimension. / Master of Science

The investigation sought to examine resistance to extinction (Rn) as a function of previous experience with downward shifts in reward magnitude. It was suggested that previous research conducted within the framework of the Spence-Amsel frustration hypothesis and the sequential hypothesis failed to administer sufficient postshift trials to adequately establish the relationships that may exist. Under one condition, four groups of rats received twenty extinction trials following forty postshift trials. Under another condition, four groups were extinguished following eighty postshift trials. An inverse magnitude of reward effect occurred in the preshift phases, however, which prevented an adequate analysis of either the shift or the Rn data, This unexpected effect was discussed within the framework of Black's incentive-motivation interpretation of reinforcement.

La diadromie représente un cycle de vie qui implique obligatoirement des migrations entre la mer et l’eau douce. La diadromie s’est maintenue dans différentes lignées évolutives de poissons, mettant ainsi en évidence l’existence d’avantages adaptatifs comme la capacité à coloniser des milieux vierges ou de mieux exploiter la variabilité spatio-temporelle de l’environnement. Une hypothèse pour expliquer l’origine et le maintien de la diadromie postule que les migrations entre mer et rivière ont pour but de placer les individus dans le milieu qui favorisera la croissance avant la première reproduction. Une croissance en mer et une reproduction en rivière (anadromie) serait favorisée si la productivité primaire en eau douce est inférieure à celle des milieux marins environnants tandis que la situation inverse (catadromie ; reproduction en mer, croissance en rivière) est attendue quand le différentiel de productivité est en faveur des eaux douces. Les résultats de notre étude réalisée à l’échelle du globe confirment cette hypothèse et montrent également que les poissons diadromes se trouvent majoritairement dans des rivières à faible richesse spécifique et/ou ouvertes à la colonisation après le retrait des glaciers au Quaternaire. Leur capacité à coloniser de nouveaux milieux permet aux diadromes d’être très présents sur les îles océaniques. Si la biodiversité sur les îles océaniques a été largement étudiée, très peu d’études ont porté sur les poissons d’eau douce qui s’y trouvent. Notre étude des peuplements de poissons des îles polynésiennes confirme la généralité d’un modèle proposé pour les faunes terrestres prédisant une relation en dôme entre richesse spécifique et âge de l’île. Nos résultats montrent également que la richesse diminue avec la distance au centre de biodiversité le plus proche et augmente avec l’altitude maximale de l’île.Malgré leur succès évolutif, les poissons diadromes se révèlent très vulnérables face aux activités humaines (surexploitation, pollution de l’eau, barrages…). De ce fait, de nombreuses espèces sont classées comme vulnérables par l’IUCN. Paradoxalement, notre étude sur les caractéristiques biologiques et écologiques des espèces qui se sont éteintes récemment à l’échelle du globe n’identifie pas la diadromie comme un facteur aggravant. Cette étude montre que le critère biologique principal menant à l’extinction est l’endémisme et une aire de distribution géographique restreinte. Bien que souvent en déclin, beaucoup d’espèces diadromes ont jusqu’ici échappé à l’extinction du fait de leur aire de distribution initiale de grande taille, résultat probable de leur grande capacité de dispersion. Un très bon exemple est l’esturgeon européen, Acipenser sturio, initialement largement distribué en Europe mais qui après un fort déclin à partir de 1850 ne se reproduit plus que dans le seul bassin de la Garonne. Pour mieux suivre le déclin de cette espèce au cours du temps, nous avons étudié sa distribution avant 1850 à l’aide de données archéozoologiques. L’analyse de ces données suggère une diminution des populations très ancienne, initiée il y a 2500 ans. A l’aide de températures passées reconstituées, nous avons montré que des températures élevées expliquent en partie l’occurrence d’esturgeons dans les sites archéologiques mais sans contribuer à expliquer son déclin. Ces résultats suggèrent un impact de l’homme précoce sur les populations d’esturgeons. Cette thèse a permis de synthétiser les connaissances sur les poissons diadromes concernant leur distribution géographique à diverses échelles de temps et d’espace, et de contribuer à une meilleure compréhension de leur diversité et de leur vulnérabilité. / Diadromous species exhibit a life-cycle implying migrations between freshwater and ocean. Diadromy is observed in many fish lineages suggesting that adaptive traits are associated with this strategy such as better dispersal ability and more efficient tracking of the spatio-temporal variability of the environment. One of the hypotheses that have been put forward to explain the evolution and persistence of diadromy states that the purpose of migrations is to select environmental conditions that will optimize pre-reproductive growth. Growing at sea and reproducing in river (anadromy) is supposed to be favored when freshwater primary productivity is lower in freshwater than in the nearby sea. The reverse (growing in freshwater, reproducing in ocean; catadromy) is expected when productivity is higher in sea than in freshwater. According to our study, conducted at a global scale, this hypothesis holds true. In addition it is shown that diadromous species are overrepresented in species poor rivers and/or in those that have been open to colonization after the retreat of glaciers after the last glacial maximum. Because of their dispersal ability, diadromous species are also over-represented on oceanic islands. Biodiversity on islands is classical topic of biogeography but yet few studies have dealt with freshwater fishes. In our study of fish communities from the Polynesian islands, we demonstrate the generality of a model initially built to explain terrestrial biodiversity on oceanic islands. As predicted by this model, species richness peaks at peaks at intermediate island age. Also revealed by our results are a positive relationship between island elevation and species richness and a decrease in species richness as the distance from the nearest biodiversity hotspot increases. Diadromy has been a successful strategy over geologic times but in face to human activities it seems to contribute to extinction proneness, as exemplified by the numerous diadromous species red listed by IUCN. Paradoxically, our study about the biological and ecological traits shared by the freshwater fishes that have been recently globally extinct does not suggest that diadromy has been a major factor. This study points to endemism and restricted geographic range size as the major determinants of extinction. It seems that because of their dispersal abilities, diadromous species tend to have widespread historical distributions and, even if many species are declining, this prevented most of them from being entirely extirpated. This is well exemplified by the European sturgeon (Asipenser sturio), formerly widely distributed over Europe and now restricted to one reproducing population in the Garonne River, because of dramatic population collapses after 1850. To have a better idea of the fate of this species before 1850, we analyzed archaeozoological records. According to our analyses, sturgeons started to decline a long time ago, about 2500 years ago. Using reconstructed past temperatures, our analyses show that a high temperature increases the probability of finding this species in archeological remains but cannot explain entirely the observed decline. This suggests that human activities impacted sturgeon populations well before the industrial revolution. This thesis by synthesizing species occurrence data over different spatial and temporal scales contributed to a better knowledge about the diversity and vulnerability of diadromous fishes.

Le contrat est un outil destiné à servir les intérêts des parties par la mise en oeuvre d'un projet dont le résultat est désiré. Exécuté conformément aux stipulations convenues, le contrat est censé procurer au créancier la satisfaction qu'il escompte, tout en libérant le débiteur de son obligation.Toutefois, dans certaines hypothèses, et sans que le contrat fasse l'objet d'une exécution reflétant ce qui a été convenu à l'origine, le créancier est satisfait tandis que le débiteur est libéré du poids de sa dette. En effet, en dépit de sa nature prévisionnelle, le contrat ne s'achève pas nécessairement par un paiement, ce dernier ne constituant pas l'unique forme de dénouement satisfactoire et libératoire. La satisfaction substitutive fait appel à différents mécanismes issus du droit des contrats et du régime général des obligations. L'étude de cette notion offre une grille de lecture apportant une lumière nouvelle sur des concepts classiques mais qu'il semble a priori impossible de relier, tant leurs natures juridiques et fonctions respectives semblent inconciliables. Par un examen de leur structure et de la logique qui les inspire, il apparaît toutefois possible de mettre en évidence des problématiques convergentes et des conséquences pratiques susceptibles d'impacter la technique contractuelle. Face à l'imprévu, la meilleure stratégie consiste à recentrer le contrat sur son véritable objectif, à savoir la réalisation d'un dénouement optimal, quitte à ce que celui-ci s'écarte du projet initialement convenu. La satisfaction substitutive illustre la recherche permanente de la solution la plus économiquement avantageuse, que ce soit par les parties, le juge ou la loi. / The contract is a commitment aimed to serve the interests of the contracting parties through the execution of a project carrying a desired outcome. Executed accordingly to what has been agreed originally, the contract is supposed to provide to the creditor the satisfaction he is expecting, while the debtor is being released from the obligation. However, in some hypotheses, and while the contract is not performed the way it should beaccording to what was agreed, the creditor benefits from a satisfaction where as the debtor isnot anymore under the burden of his debt. Indeed, despite its forward-looking nature, the contract does not always end by a payment, as this form of termination does not constitute the single way to provide satisfaction to the creditor and discharge to the debtor. The substitute satisfaction, as an atypical form of termination, relies on different mecanismscoming from contract law and general rules of obligations. Studying this notion gives keys forunderstanding that bring a new light on some classical concepts which seem difficult to linktogether, since their structure and logics appear irreconcilable. By examining their structureand their logic, it appears possible to highlight convergent problematics and practicalconsequences that can have impact on contractual technique. To face the unforeseen, the best strategy is to refocus the contract on its true objective, which is to bring an optimal termination, even if there is a discrepancy between what was agreed andwhat is done. Substitute satisfaction illustrates the permanent search for the most worthwhile solution on an economic point of view by the contracting parties, the judge or the law.

The present thesis examined extinction of conditioned fear in the developing rat. In the adult rat, the hippocampus is thought to be important for the context-specificity of extinction. Because the hippocampus is a late-maturing structure, it was hypothesised that context-modulation of extinction may be different across development. The first series of experiments investigated reinstatement of extinguished fear in the developing rat (Chapter 2). The results showed that P24 rats exhibited context-specific reinstatement. On the other hand, P17 rats did not exhibit reinstatement of extinguished fear following a US reminder treatment. The failure to see reinstatement in P17 rats was not due to the reminder treatment being ineffective in these rats because the same treatment alleviated spontaneous forgetting in rat this age. The second series of experiments then examined the renewal effect and GABAergic involvement in extinction in P24 and P17 rats (Chapter 3). It was observed that P24 rats displayed renewal whereas P17 rats did not. Also, pre-test injection of FG7142 recovered extinguished fear in P24 rats but not in P17 rats, even across a range of doses. This failure to see any FG7142 effect on extinction in P17 rats was not due to the lack of responsiveness to this drug in these rats because FG7142 was found to be effective in alleviating spontaneous forgetting in rats this age. The third series of experiments then examined the effect of temporary inactivation of the amygdala on extinction and re-extinction in the developing rat (Chapter 4). It was observed that extinction retention is impaired in both P24 and P17 rats if the amygdala is inactivated during extinction training. Interestingly, when a CS that had been previously extinguished and then re-trained was re-extinguished, re-extinction was amygdala-independent if initial extinction occurred at 24 days of age but amygdala-dependent if initial extinction occurred at 17 days of age. That is, amygdala involvement in re-extinction was dissociated across development. Taken together, these experiments provide strong evidence for fundamental differences in mechanisms underlying fear extinction across development. The implications of the findings were discussed in light of the theoretical and neural models of extinction.