Human footprint - in which land is converted for human use - is a leading contributor to global habitat and biodiversity loss. The accelerated rate of human landscape change to meet our growing needs has led to the direct loss of critical habitat and shifts in species distributions, interactions, and behaviour. These altered conditions affect species’ ability to adapt to environmental stressors, while some species thrive and others decline. In North America, one ungulate has successfully invaded new habitat in conjunction with human land use – the white-tailed deer. Across the continent, the invasion of white-tailed deer has led to increased competition with other ungulate species including mule deer, moose, and woodland caribou. In regions with abundant apex predators, they have become a source of primary prey as their populations increase. The mechanisms by which deer occupy landscapes in the northern extents of their geographic range are not well studied outside of the winter months, or how deer respond behaviourally to various types of human disturbance in a predator-rich environment. To address these knowledge gaps, I examined population scale resource selection across seasons and individual movement behaviour in white-tailed deer in northeastern Alberta’s intensively developed oil and gas landscape. I used previously developed models of predator frequency to spatially extrapolate wolf and black bear occurrence across my study region as indicators of indirect predation risk. I used two approaches to habitat modeling to examine deer responses to various modes of human landscape change, including roads, seismic lines, and cut blocks in addition to predators and natural habitat. Deer were best described by cumulative effects – or the combination of all of these factors – across all seasons with proximity to linear features explaining the most variation among the parameters tested. Most prominently in winter, deer strongly selected for habitat features expected to contain abundant natural sources of forage, and linear features, despite a potential increased risk of predation by wolves – suggesting that deer make energetic trade-offs between forage availability and predation risk. At the individual level, deer significantly increased their rate of movement when occupying habitat associated with predation risk. I suggest that deer make greater energetic trade-offs during winter when mobility is limited to evade predators and energetic costs are higher. The continued use of anthropogenic features post-winter, increased rate of movement and spread of landscape occupancy by deer may allude to the importance of human disturbance in maintaining deer in northern climates. Linear corridors may be an important mechanism by which deer are able to successfully colonize new areas at the northern extents of their range. My results shed light on the drivers of deer distributions in human altered landscapes for managing populations where the invasion of deer is complicit in the decline of other ungulate species such as woodland caribou in Alberta’s boreal forest. / Graduate

The southern fox squirrel (Sciurus niger niger) has the northernmost part of its range in Virginia. For the past 100 years, southern fox squirrels have been declining due to habitat fragmentation, cover type conversion, and fire suppression. Decrease in growing season burns, hardwood encroachment and forest mesophication have transformed pine hardwood woodlands and pine (Pinus spp.) savanna habitats that southern fox squirrels prefer to hardwood dominant habitats that eastern gray squirrels (Sciurus carolinenisis) prefer. These habitat changes have the potential to increase competition among the two species. The main objectives of my study were to investigate the general resource needs, occupancy, and home range of southern fox squirrels as well as the impact of resource partitioning and possible competition with eastern gray squirrels in the Piedmont and Coastal Plain of Virginia. I captured, radio collared and tracked four individuals at Big Woods Wildlife Management area and Piney Grove Complex using 95% and 50% kernel density estimate. I found an average male home range 173.49 ha (SE = 25.73, N = 2) and 40.62 ha (SE = 5.87, N = 2) and an average female home range of 28.51 ha (SE = 0.49, N = 2) and 4.71 ha (SE = 0.34, N = 2). I then identified the second and third order habitat selection in which my top models identified selection for pine savanna cover types (β = 2.095, SE = 0.158), increasing number of burns since 2019 (β =1.24, SE = 0.098), and decreased time between burns (β = -0.233, SE = 0.097). I used two-species occupancy modeling which reflected that gray squirrel occupancy increased with increasing time since last prescribed burn. However, southern fox squirrel occupancy, in the absence of gray squirrels, decreased with increasing time since last burn. My informed single-season occupancy model confirmed that southern fox squirrel occupancy decreased with time since the last burn. Presence in the absence of gray squirrels suggests that southern fox squirrels are selecting habitats on BWPGC with respect to both resource needs and competition with gray squirrel. Additionally, my level-of-effort (LOE) analysis indicated that 7 consecutive days of camera trapping without a southern fox squirrel detection would provide 90% confidence of the species' absence in areas burned 2 or more years prior to sampling in southeastern Virginia. Further management for southern fox squirrels in the future should focus on high rotational (short fire return interval) burns in areas of savanna as well as pine-hardwood mixed areas and hardwood-pine savanna ecotones. / Master of Science / The southern fox squirrel (Sciurus niger niger) is a subspecies of fox squirrel that ranges from southeastern Virginia down to northern Florida. All throughout its range in the Southeast, southern fox squirrel habitat has been fragmented from natural mixed pine-hardwood woodland forests to agriculture and high rotation pine plantations. Additionally, habitat has been further transformed by the lack of prescribed fire as a management tool on the landscape. This has in turn created sparse and fragmented local populations of southern fox squirrels as well as possible competition with gray squirrels. Further, the southern fox squirrel has not been studied in Virginia in over 20 years and management recommendations are lacking. I studied the resource needs, occupancy, home range and competition of southern fox squirrels in two physiographic regions of Virginia: the Coastal Plain and Piedmont regions. The Coastal Plain field site was Big Woods Wildlife Management Area and The Nature Conservancy's Piney Grove- both adjacent to each other. The Piedmont field site was Military Training Center Fort Barfoot. Here I utilized camera trapping, nest box monitoring, live trapping, and radio tracking to assess the resources they are utilizing in each area through home range analyses. I found that southern fox squirrels are selecting areas that have low fire return intervals and are located in pine savanna habitats. Therefore, fire should be prioritized as a management tool for southern fox squirrel habitat in pine savanna areas. I also used camera trapping data to identify the possible competition among gray and fox squirrels and fox squirrel detection through occupancy modeling. My findings reflected that there is apparent competition between southern fox squirrels and eastern gray squirrels and that southern fox squirrels are selecting heavily burned areas not only for their resource needs, but also because gray squirrels are absent. I concluded through my studies that the southern fox squirrel currently occupies southeastern Virginia, particularly in the Coastal Plain, however at low numbers. This could be due to suitable habitat on Big Woods/Piney Grove, but the surrounding habitat is of marginal quality. Further, in documenting southern fox squirrels, multi day camera surveys in mixed pine-hardwood woodland and pine savannas should be prioritized. Also, in aims to increase the presence of southern fox squirrels on the landscape, short rotation prescribed burning should be prioritized as well as additionally considering meditation among gray squirrel and fox squirrel competition.

Sagebrush (Artemisia spp.) is an important source of food and cover for many animals, especially during winter months. Understanding how wildlife species respond to sagebrush management actions can help improve conservation planning. Dixie harrow is a method of improving spring/summer habitat for many herbivores by reducing sagebrush cover to stimulate the growth of grasses and forbs. These treatments, however, may influence the quantity and quality of sagebrush available to greater sage-grouse (Centrocercus urophasianus; hereafter, sage-grouse) during winter. We evaluated the effects of Dixie harrow on sage-grouse resource selection during winter (Chapter 1) and on the nutritional value of sagebrush (Chapter 2). We were unsure what effect Dixie harrow would have on the nutritional value of sagebrush, but hypothesized that sage-grouse would select for untreated areas because they contained a higher quantity of food and cover. We captured 81 sage-grouse and fit them with GPS transmitters. Using 6,728 winter locations, we modeled third-order resource selection. Further, we collected samples of sagebrush plants that sage-grouse had eaten from (n = 54), samples of sagebrush plants passed by but not eaten from (n = 54), as well as samples from random locations inside (n = 60) and outside Dixie harrow treatments (n = 60). Contrary to our hypothesis, sage-grouse selected for Dixie harrow treatments during winter. We found that sage-grouse selectively browsed sagebrush plants with increased nutritional value, and that sage-grouse browsed plants inside treatments more frequently than outside the treatments, but Dixie harrow treatments had no measurable effect on the nutritional value of sagebrush. Based on our results, Dixie harrow treatments performed at the southern extent of the sage-grouse range will create habitat that sage-grouse prefer during winter, but we were unable to ascertain why sage-grouse select for Dixie harrow treatments during winter.

No description available.

Despite the prevalent distribution of bobcats in western Virginia and the broader region of Appalachia, there is a paucity of information on their spatial ecology in this region. Due to the unique ecological conditions of Appalachia, and increasing public interest surrounding the role of bobcats as predators in the region, there is a need for local information on bobcat ecology. I utilized data from 20 GPS collared bobcats (14M, 6F) to investigate bobcat spatial ecology in the mountains of Western Virginia. Average resident male home range size was 33.9 ± 2.6 km^2, nearly 3 times larger than average resident female home range size (12.1 ± 2.4 km^2). Seasonal areas of use did not differ in size among seasons, but exhibited minor shifts in location and shape. Average male movement rates (232.3 ± 12.0 meters/hour) were 1.5 times greater than average female movement rates (154.4 ± 8.9 meters/hour). Male movement rates increased during the dispersal season and female movement rates increased during the denning/kitten-rearing season. Second order resource selection analysis indicates that bobcats of both sexes select home ranges at higher elevations than expected at random, and that selection varies between sexes and among seasons. Third order resource selection analysis indicates that bobcats select for locations near open canopy structure, and avoid forest interior. These findings build upon previous findings regarding bobcat diet and population dynamics to provide a comprehensive understanding of basic bobcat ecology in western Virginia, and will inform further research investigating predator/prey interactions. / Master of Science / Despite the prevalent distribution of bobcats in western Virginia and the broader region of Appalachia, there is a paucity of information on their spatial ecology in this region. Due to the unique ecological conditions of Appalachia, and increasing public interest surrounding the role of bobcats as predators in the region, there is a need for local information on bobcat ecology. I utilized data from 20 GPS collared bobcats (14M, 6F) to investigate bobcat spatial ecology in the mountains of Western Virginia. I found that male bobcats use more space than females, with home ranges nearly 3 times larger and movement rates 1.5 times higher. I found that home ranges do not expand or contract throughout the year, but do shift in shape slightly. I found that bobcats select home ranges at higher elevations, indicating that home ranges are predominantly located on ridges. I found that selection of home ranges differed between sexes and seasons, indicating that shifts in home range shape throughout the year reflect varying habitat selection. Within their home ranges, I found that bobcats select for areas of open canopy resulting from fields, fire, and timber harvest; and avoid areas of forest interior. These findings build upon previous findings regarding bobcat diet and population dynamics to provide a comprehensive understanding of basic bobcat ecology in western Virginia, and will inform further research investigating predator/prey interactions.

[pt] Nos últimos anos, o uso de dispositivos móveis em grades computacionais tem sido alvo de crescente investigação. Entretanto, um problema mais desafiador, referente ao estabelecimento dinâmico de grades móveis, baseadas exclusivamente em redes sem fio ad hoc, ainda tem sido pouco investigado. Uma contribuição desta tese é a proposta de uma arquitetura de software específica para grades móveis, que pode ser igualmente aplicável a redes sem fio infraestruturadas e ad hoc. Em grades fixas, a responsabilidade de prover um serviço computacional é compartilhada entre dispositivos com relativa abundância de recursos, se comparadas a grades móveis. Nestas últimas, é interessante que a descoberta e a seleção de recursos para execução de tarefas sejam tratadas conjuntamente, de modo a promover a seleção automática dos dispositivos com maior disponibilidade de recursos, considerando-se os requisitos da aplicação. Entretanto, tais elementos têm sido tradicionalmente tratados em separado na literatura relacionada a grades móveis e, em grande parte das abordagens existentes, assume-se que a seleção de recursos seja executada de forma manual pelos usuários da grade móvel. Esta tese propõe, como uma outra contribuição, um protocolo que integra as fases de descoberta e seleção automática de recursos em grades móveis, permitindo que a provisão de serviços computacionais seja escalonada entre os dispositivos com maior disponibilidade dos recursos requeridos pelo serviço. Devido à característica dinâmica dos recursos que correspondem às requisições dos usuários em uma grade móvel (por exemplo, tempo de CPU livre e memória disponível), o protocolo baseia-se unicamente no envio, sob demanda, de requisições via broadcast. No entanto, principalmente em redes sem fio ad hoc de saltos múltiplos, essa estratégia pode acarretar uma sobrecarga nos dispositivos envolvidos, tanto na difusão de requisições quanto no encaminhamento de respostas. Uma terceira contribuição desta tese é o desenvolvimento de um mecanismo que permite reduzir a sobrecarga, devido à difusão de mensagens de resposta, por meio da supressão de respostas excedentes ao longo da rede. O mecanismo, embora implementado no contexto do protocolo proposto nesta tese, pode ser aplicado também a outros protocolos de descoberta baseados no envio de requisições via broadcast. Os resultados experimentais, obtidos em redes de testes e em plataformas de simulação, mostram que o protocolo proposto provê um balanceamento de carga eficiente entre os dispositivos, mediante o aumento do número de requisições. Além disso, pode-se observar que o mecanismo de supressão de respostas é escalável no que diz respeito ao crescimento do número de dispositivos, em comparação com protocolos de descoberta baseados puramente no envio de requisições por broadcast em redes sem fio ad hoc. / [en] In the last few years, the use of mobile devices in computational grids has seen a growing interest. Nevertheless, a more challenging issue, the dynamic establishment of mobile grids on wireless ad hoc networks, has been so far only partially addressed. The first contribution of this thesis is the proposal of a software architecture for mobile grids that can be used for both infrastructured and ad hoc wireless networks. In the execution of conventional applications in grids, the responsibility to provide the service is shared among the most resourceful mobile devices. In mobile grids, it is fundamental that resource discovery and selection of resources are jointly handled. This calls for a mechanism that promotes the automatic selection of the best resource providers amongst the discovered nodes, taking into account the requirements of the application. Discovery and selection, however, have been traditionally handled separately and in most approaches the selection of resources and services requires explicit intervention by the user of the mobile grid. As a second contribution of this thesis, we propose a protocol that integrates the phases of resource discovery and automatic selection in mobile grids, allowing that computational resource provisioning is scheduled among the most resourceful nodes. Due to the dynamics of the resources needed in a mobile grid (for example, free CPU time and available memory), the protocol is based solely on demand- driven broadcasts. However, mainly in multihop ad hoc wireless networks, this strategy can incur in overhead at the involved devices, due to the diffusion of requests and replies. A third contribution of this thesis is the development of a mechanism that allows to reduce this overhead by means of the suppression of redundant replies in the network. The mechanism has been implemented in the context of the proposal protocol, but can be applied as well to other query-based discovery protocols based on broadcasts. The experimental results obtained from executions in a testbed and through simulations show that the proposed protocol provides efficient load balancing between devices with an increasing number of requests. Moreover, it can be observed that the mechanism for suppression of replies scales well with respect to an increasing number of devices when compared to other discovery protocols in wireless ad hoc networks that are purely based on requests via broadcast.

The western toad (Anaxyrus boreas, formerly Bufo boreas) is one of many amphibian species considered to be at risk of extinction (COSEWIC status is Special Concern). I examined habitat use patterns of the western toad using several methods to gain a better understanding of its habitat requirements. I examined the relationship between relative abundance of the western toad and two sympatric amphibian species (wood frog, Lithobates sylvaticus; and boreal chorus frog, Pseudacris maculata) and habitat features at eight scales of spatial extent at 24 wetlands in the Lake Utikuma region of Alberta, Canada. I radio-tracked adult western toads in three study areas in the Aspen Parkland and Boreal regions of north-central Alberta to examine 1) whether patterns of habitat selection change with different scales of spatial extent, spatial resolution, habitat composition, temporal period, and between males and females during the active period, 2) habitat used for hibernation, and 3) factors influencing the timing and nature of movements to hibernation sites. I found that the abundance of the three amphibian species was best described at different spatial extents and was related to the biology of each species. Resource Selection Function (RSF) models, created using radio-telemetry data, indicated that habitat selection was scale-dependent for western toads; differences in selection were observed among study designs, study areas, time periods, and sexes. Predictive ability did not differ significantly among study designs. However, models that were created using a fine-grained map and home-range spatial extent generally produced models with greater predictive ability than models using a coarse-grained map or population-range extent. During the active season toads selected open habitat types such as wet shrub, disturbed grass, and crop/hay fields. Western toads hibernated terrestrially in pre-existing tunnels and the majority of toads hibernated in forest stands dominated by spruce. Toads used hibernation sites 1461936 m from breeding ponds and 68% of hibernacula were communal. Arrival at and entry into hibernation sites was influenced by temperature and/or day length; larger toads moved to hibernation sites later in the year. My research results can be used to identify and protect habitat for western toads in Canada. / Environmental Biology and Ecology

Call-response surveys were conducted to better delineate and estimate the population of the nationally threatened least bittern and their habitat requirements in southern Manitoba, Canada. Other marsh bird species whose populations are believed to be declining due to wetland loss throughout, or in parts of, their range were also surveyed including the American bittern, pied-billed grebe, sora, Virginia rail and yellow rail. Surveys were conducted during the 2003 and 2004 breeding seasons within 46 different wetlands. Least bitterns were encountered on 26 occasions at 15 sites within 5 wetlands. The sora was the most abundant and widely distributed target species and was encountered on 330 occasions in 39 of the 46 surveyed wetlands. Yellow rails were not detected during either survey year due to survey methodology. Use of the call-response survey protocol led to an increase in the numbers of all target species detected. This increase was more significant for the least bittern, sora and Virginia rail. Habitat was assessed as percent vegetation cover within a 50-m radius around the calling sites, and forest resource inventory data were used in a Geographic Information System to determine the landscape composition within a 500-m radius around the sites and within a 5-km radius around the wetlands surveyed. Logistic regression analyses were used to evaluate the relationship between the presence of the target species and the site and landscape characteristics. The target species responded differently to different site and landscape characteristics. Least bittern and pied-billed grebe selected areas with higher proportions of Typha spp. and tall shrubs; American bittern also selected areas with higher proportions of tall shrubs. At the 5-km scale, the American bittern responded positively to the amount of wetland and some positive trends were also detected for the pied-billed grebe. Sora and Virginia rail were not associated with any of the measured landscape characteristics. One of the most important steps towards the conservation of marsh bird species in Manitoba and elsewhere is the development, adoption, and implementation of a standardized survey protocol. Based on the results of the present study, I recommend that future surveys include both a passive and call-broadcast period for marsh bird species. Future surveys should be conducted in both the morning and evening and sites should be visited 3 times each during the breeding season. In southern Manitoba, call-response surveys should begin as early as the beginning of May to ensure the survey incorporates the period of peak vocalization. I recommend that future yellow rail surveys be conducted after dark. In this study many of the target species selected sites that had a greater area of wetland habitat surrounding them. Future wetland conservation efforts should focus on the protection and/or restoration of wetland complexes to ensure that remaining wetlands do not become smaller and increasingly isolated from one another. In addition, the Rat River Swamp was found to be the most productive marsh complex for least bittern in southern Manitoba. Measures should be taken to protect this area from future development and alteration. / May 2006

No description available.

Call-response surveys were conducted to better delineate and estimate the population of the nationally threatened least bittern and their habitat requirements in southern Manitoba, Canada. Other marsh bird species whose populations are believed to be declining due to wetland loss throughout, or in parts of, their range were also surveyed including the American bittern, pied-billed grebe, sora, Virginia rail and yellow rail. Surveys were conducted during the 2003 and 2004 breeding seasons within 46 different wetlands. Least bitterns were encountered on 26 occasions at 15 sites within 5 wetlands. The sora was the most abundant and widely distributed target species and was encountered on 330 occasions in 39 of the 46 surveyed wetlands. Yellow rails were not detected during either survey year due to survey methodology. Use of the call-response survey protocol led to an increase in the numbers of all target species detected. This increase was more significant for the least bittern, sora and Virginia rail. Habitat was assessed as percent vegetation cover within a 50-m radius around the calling sites, and forest resource inventory data were used in a Geographic Information System to determine the landscape composition within a 500-m radius around the sites and within a 5-km radius around the wetlands surveyed. Logistic regression analyses were used to evaluate the relationship between the presence of the target species and the site and landscape characteristics. The target species responded differently to different site and landscape characteristics. Least bittern and pied-billed grebe selected areas with higher proportions of Typha spp. and tall shrubs; American bittern also selected areas with higher proportions of tall shrubs. At the 5-km scale, the American bittern responded positively to the amount of wetland and some positive trends were also detected for the pied-billed grebe. Sora and Virginia rail were not associated with any of the measured landscape characteristics. One of the most important steps towards the conservation of marsh bird species in Manitoba and elsewhere is the development, adoption, and implementation of a standardized survey protocol. Based on the results of the present study, I recommend that future surveys include both a passive and call-broadcast period for marsh bird species. Future surveys should be conducted in both the morning and evening and sites should be visited 3 times each during the breeding season. In southern Manitoba, call-response surveys should begin as early as the beginning of May to ensure the survey incorporates the period of peak vocalization. I recommend that future yellow rail surveys be conducted after dark. In this study many of the target species selected sites that had a greater area of wetland habitat surrounding them. Future wetland conservation efforts should focus on the protection and/or restoration of wetland complexes to ensure that remaining wetlands do not become smaller and increasingly isolated from one another. In addition, the Rat River Swamp was found to be the most productive marsh complex for least bittern in southern Manitoba. Measures should be taken to protect this area from future development and alteration.