At its inception, taxonomy simply provided guidelines for nomenclature. It has since developed into a science applied to problems relating to economics, conservation and even law. Taxonomy is arguably one of the most important pillars of the biological sciences, providing the framework from which all other studies are conducted. We expand this essential foundation by describing four new species of Trox in Chapter 2 of this dissertation - the first to be recorded in South Africa since Scholtz’s comprehensive revision of the family in 1980. All four new species are flightless and are restricted to densely vegetated areas. Based on their morphological characteristics, we conclude that the new species probably belong to the endemic South African “horridus”-group of Trox. At present, morphological studies suggest that Trogidae consists of only three genera – Polynoncus, Omorgus and Trox. The current consensus on the zoogeography of trogids, based on their current distribution patterns, is that they evolved in Central Pangaea, prior to the split that formed Laurasia and Gondwanaland. The Trox lineage is thought to have speciated in temperate Laurasia, invading Africa via a temperate faunal exchange route only after Gondwanaland had separated into the southern landmasses. However, in Chapter 3, our work, based on a molecular dataset, challenges these ideas. We investigated the phylogenetic relationships within the family by analysing the 16S ribosomal subunit gene on the mitochondrial genome, which has proved useful in investigating sub-familial relationships due to its fairly conserved nature. We performed both phenetic (Neighbor-Joining and Minimum Evolution) and likelihood (Parsimony, Maximum Likelihood and Bayesian Inference) analyses on the resulting molecular dataset. We found only partial support for the theories suggested by the morphological dataset. In all analyses, we find four major groups - Polynoncus, Omorgus, Holarctic Trox (Trox s. str.) and African Trox (Phoberus) - not three as suggested by previous morphological studies. We strongly support the promotion of Phoberus to full generic status as it is represented by a monophyletic group in all analyses. Given the evolutionary divergence suggested by our molecular phylogeny, we still propose a Pangaean origin for Trox sensu lato, but suggest that this lineage might have had its origin in what would become North Africa. It is likely to have inhabited the globe-spanning metamorphic geological features known as the Pan-African Belts, which would have offered the ancestral temperate biome preferred by this genus. We theorize that the formation of the Tethys Sea would have separated Trox s. str. from the Phoberus lineage. Copyright / Dissertation (MSc)--University of Pretoria, 2009. / Zoology and Entomology / unrestricted

New product development (NPD) is vulnerable to a wide variety of risks arising from within the firm or from the external environment. Existing categorizations of NPD project risks are partial or ill-defined and consequently there is no clear consensus among researchers and practitioners about what constitute NPD project risks. To address this gap, this thesis deploys a systematic literature methodology to inductively develop a comprehensive risk taxonomy from a review of 124 empirical studies. This taxonomy is then empirically validated through a survey capturing data from 263 NPD projects conducted by UK firms. The thesis further investigated the moderating effect of NPD project type (incremental or radical), firm size (SMEs and large firms) and industry sectors on the proposed risk taxonomy. Variation in the perceptions of NPD risk by different members of the team was explored as well. The findings revealed that the principal risk factors affecting NPD projects are technological rapidity risk, supply chain risk, lack of funding and resource risk. The risk profile of radical NPD projects differed to that of incremental projects. SMEs were more vulnerable to NPD project risks than large firms. Most risks influenced NPD projects equally across industrial sectors. Members of NPD project teams from different backgrounds or with different roles perceived risks differently. The proposed taxonomy and its subsequent empirical validation provides a comprehensive and robust taxonomy for identifying and managing risks associated with different types of NPD project conducted by firms of varying sizes from different industrial sectors.

Only thirty known species have populations at high latitudes in both hemispheres, this is, a bipolar distribution. Five of them belong to the genus Carex. Before attempting to elucidate the origins of such distributions, we need to resolve taxonomical problems that are typically encountered in such species. We focus on the Carex capitata complex, which includes Carex arctogena as a bipolar species, in worldwide scope sampling. A morphometric study and phylogenetic analyses based on maximum parsimony, Bayesian inference and Statistical Parsimony have revealed: a) C. capitata and C. arctogena are different species; b) C. anctarctogena is a synonym of C. arctogena and c) a great biodiversity previously undetected in western North America that could lead to the description of three new taxa (“Carex cayouetteana”, Carex sp. nov. 1 and 2) comprised in the so called “C. cayouetteana” lineage. More studies are needed in some C. capitata samples from Russia that appears in the molecular analyses in a strongly supported clade.

This paper examines and summarizes the data standard XBRL. The paper explains taxonomies and describes the most important institutions involved in the development and adoption of XBRL as a format for reporting financial and non-financial information in the world. Most attention is devoted to the implementation of XBRL in various parts of the world and in the Czech Republic. The practical part is focused on the possibilities of using XBRL in the Czech Republic and the evaluation of readiness of the Czech environment for the adoption of XBRL in comparison with other countries in the world.

This dissertation represents a study on Ceratocystis spp. with hat-shaped ascospores. Previously only six species of Ceratocystis with this spore form were known. These include C. fimbriata, C. pirilliformis, C. albofundus, C. moniliformis, C. moniliformopsis and C. acericola. In this study, we have discovered and tentatively described three new species with hat-shaped ascospores. One of these group with the larger C. fimbriata clade, while the other two reside within the larger C. coerulescens clade. Chapter one provides a concise summary of the literature pertaining to the genus Ceratocystis. The intention of the chapter is to introduce readers to this important genus of plant pathogens and to provide a background regarding their taxonomy, ecology, biochemistry and variation in species. Emphasis is placed on the distinction of Ceratocystis from Ophiostoma, as ell as on those Ceratocystis spp. with hat-shaped ascospores that negatively impact upon plantation forestry species. This chapter shows how Ceratocystis spp. associated with hardwood species in commercial forestry plantations have increased in number and it provides the background for research presented in the following five chapters. In the second chapter of this dissertation, a new species of Ceratocystis was discovered amongst isolates from the Himalayan mountain range of Bhutan. This fungus, in association with the bark beetle Ips schmutzenhoferi, is responsible for large-scale deaths of Himalayan spruce trees in Bhutan. The fungus is morphologically very similar to C. moniliformis and C. moniliformopsis, but differences in culture morphology, survival at different incubation temperatures and DNA sequence data based on three different gene regions supported the fact that this is a unique species. The fungus has thus been tentatively described as Ceratocystis bhutanensis prov. nom. In chapter 3, I consider isolates of a Ceratocystis sp. recently discovered associated with dying clove trees in Northern Sulawesi, Indonesia. The fungus was found at a very high level of incidence, but was at first identified as C. fimbriata, based on morphological characteristics. Differences were observed in cultures of this fungus when they were compared with C. fimbriata especially in terms of colony colour and growth at different temperatures. Morphological differences were also observed when the clove fungus was compared with C. fimbriata isolates. When three different DNA gene regions were sequenced and compared, it was clear that this fungus represents a new species. The fungus is, therefore, tentatively described as Ceratocystis polychrome prov. nom. in this dissertation. Ceratocystis polychrome prov. nom. Isolates obtained from cloves in Sulawesi displayed three distinctly different culture morphologies. In Chapter 4 of this dissertation we used DNA sequence data and microsatellite markers to consider whether these differences could be observed at the molecular level. Comparisons of sequence data for the ITS region gave no distinction between any of the morphological groups. A total of 50 isolates were studies using microsatellites markers developed for C. fimbriata. No distinction could be obtained between isolates representing the three different culture morphological groups. The 50 isolates were subsequently treated as one population in further analyses. With the aid of the microsatellite markers, it was shown that this population probably originated from Sulawesi and that it benefits form sexual outcrossing. In chapter five of this dissertation, a study was undertaken to consider the taxonomic status of C. moniliformis. Consideratble variation has been noted in different descriptions of this species. It also has a very wide host and geographic distribution raising speculation that C. moniliformis represents a species complex rather than a single taxon. Based on morphological and DNA sequence data from three gene regions, isolated from Sumatra were described as a new species, which we have tentatively named C. tribiliformis prov. nom. The other C. moniliformis isolates were all the same, despite the fact that they originated from a wide range of hosts and areas. The fungus correctly bearing the name C. moniliformis, C. moniliformis sensu stricto, therefore does not seem to tepresent a species complex. Species such as the closely related C. tribiliformis prov. nom., C bhutanensis prov. nom., C. omanensis prov. nom. and C. moniliformopsis all belong to the larger C. moniliformis sensu lato group, and all have hat-shaped ascospores, conical spines on the ascomatal bases, disc-shaped bases to the ascomatal necks and are phylogenetically closely related to C. moniliformis. Studies presented in this dissertation provide considerable new knowledge regarding various Ceratocystis spp. with hat-shaped ascospores. Three new species are described and I have also been able to show that C. moniliformis sensu stricto is monophyletic. Two of the species (C. tribiliformis prov. nom. and C. bhutanensis nom. prov.) group within the larger C. coerulescens clade while C. polychrome prov. nom. groups within the larger C. fimbriata clade. Studies in this dissertation have also improved our knowledge of the identity of several species previously incorrectly identified as either C. moniliformis or C. fimbriata. What has clearly emerged from this dissertation is the need for a monograph of Ceratocystis to include all new species and to thoroughly consider the population biology and ecology of all species. / Dissertation (MSc)--University of Pretoria, 2008. / Microbiology and Plant Pathology / MSc / Unrestricted

Please read the abstract in the section front of this document. / Thesis (PhD)--University of Pretoria, 2008. / Microbiology and Plant Pathology / Unrestricted

Magister Scientiae (Biodiversity and Conservation Biology) - MSc (Biodiv and Cons Biol) / This thesis explores the diversity, spatial patterns and community structure for the sponges (Porifera) along the west and south coasts of South Africa. Species collected were identified to the lowest level of lowest taxonomic unit possible (either species or genus). The study site was divided into areas and in each of these we documented the spatial diversity and in so doing were able to assess the variation of sponge communities between the west and south coasts. The total number of species recorded for this deep-water region was eighty-three of which nineteen are described. The south coast was more diverse than the west coast and eleven species were found to be common to both coasts. The analysis based on location and depth showed that both coasts are significantly different to each other. We determined that these areas are biogeographically separated. Species contributing toward the dissimilarity between both coasts include Suberites carnosus, Myxilla (Burtonanchora) sp 1, Rossella antarctica, Tetilla capillosa and Haliclona sp. Patterns of species richness showed an increase in diversity from the west to south. It was found that species richness increases with depth for both coasts but only up to 350 m for the west coast and 200 m for the south coast. However, the sampling effort was determined to possibly have not been enough to gain a full understanding of species richness for the entire study area as the number of species was correlated with sampling effort. Estimated richness found that higher richness of sponges could still be found within most of the best bins and for each coast. An estimate of samples needed both each depth bin per coast showed that more samples would be needed on the south coast and this is possibly due to the greater variety and variability of the species found on the coast. The sponge community on the south coast was found to have no significant difference in pattern with some of the depth bins, whereas depth plays a role in sponge community on the west coast. Species of Suberites were dominant at depths lower than 200 m while Hamacantha (Vomerula) esperioides was dominant between 200 and 350 m with Tetilla capillosa dominated depths lower than 350 m. The thesis is concluded with an overview of what is now known and what still needs to be discovered and determined to further enhance biodiversity knowledge in the country.

Leguminosae - Mimosoideae possui cerca de 3300 espécies subordinadas a cerca de 80 gêneros, distribuídas principalmente nas regiões tropicais e subtropicais, com alguns gêneros se estendendo até áreas de clima temperado. O presente trabalho refere-se ao estudo taxonômico das espécies do gênero Stryphnodendron, que apresenta distribuição neotropical com limite norte de ocorrência a Nicarágua e como limite sul o Estado do Paraná no Brasil. A realização desse estudo foi baseada em levantamento bibliográfico, consultas aos acervos dos principais herbários brasileiros e do exterior, e em expedições de coleta em áreas de ocorrência natural dos táxons do gênero. Stryphnodendron reúne 36 táxons, dos quais cerca de 89% ocorrem no Brasil e aproximadamente 50% são exclusivos do território brasileiro, ocorrendo em diversos tipos de vegetação, mas principalmente em Cerrados e na Floresta Amazônica. Apenas quatro espécies não têm registro de coleta no Brasil: S. excelsum Harms (Nicarágua, Costa Rica e Panamá), S. levelii R.S.Cowan (Venezuela), S. moricolor Barneby & Grimes (Guiana Francesa) e S. porcatum D.A.Neill & Occhioni f. (Equador). Com base na morfologia externa, Stryphnodendron pode ser caracterizado por ser inerme, apresentar ápice dos ramos dotados de indumento ferrugíneo, folhas bipinadas (dotadas de nectários extraflorais), flores pentâmeras, sépalas e pétalas unidas, corola de 2-6mm de comprimento, dez estames livres ou unidos muito próximo à base, ovário curto-estipitado, flores reunidas em espigas, além de frutos tipo legume nucóide ou folículo, septados e sementes com endosperma e pleurograma do tipo apical-basal. Como resultado da revisão, foram definidos quatro padrões de distribuição das espécies, dezoito lectotipificações são aqui designadas, seis novas sinonimizações, duas novas combinações, um restabelecimento de espécie, uma mudança de status nomenclatural, três espécies excluídas e dois nomes mantiveram-se como nomes duvidosos. Foram ainda detectadas nove espécies novas: Stryphnodendron conicum, Stryphnodendron dryaticum, Stryphnodendronfasciatum, Stryphnodendron holosericeum, Stryphnodendron orinocense, Stryphnodendron procerum, Stryphnodendron riparium, Stryphnodendron velutinum e Stryphnodendron venosum. / Leguminosae - Mimosoideae comprises 80 genera and about 3,300 species, mainly distributed in tropical and subtropical regions, although some genera can reach temperate areas. We reviewed species from Stryphnodendron, which presents Neotropical distribution with northern limit in Nicaragua and southern limit in Paraná state, Brazil. This study was based on bibliographical review, consults to the main Brazilian, North American and European herbaria, and field expeditions to the genus\' natural occurrence areas. Stryphnodendron includes 36 taxa; about 89% of which occur in Brazil, and ca. 50% of these are exclusive of this country. It can be found at several kinds of vegetations, mainly at Cerrado and Amazon Forest. Only four species have not been reported to Brazil: S. excelsum Harms (Nicaragua, Costa Rica and Panama), S. levelii R.S.Cowan (Venezuela), S. moricolor Barneby & Grimes (French Guiana) and S. porcatum D.A.Neill & Occhioni f. (Ecuador). Based on its external morphology, Stryphnodendron can be characterized by its spineless habit, ferruginous terminal portion of shoots, bipinnate leaves (with extrafloral nectaries), flowers with 5¬fused sepals, 5-fused petals, corolla 2 to 6 mm long, 10 stamens, the filaments free or sometimes united just near the base, short-stipitate ovary, spike-arranged flowers, nucoid legume or follicle fruits (sensu Barroso et al. 1999), septate, and endospermous seeds with apical-basal pleurograms. We defined four distribution patterns within Stryphnodendron, twelve lectotipifications, six new synonyms, two new combinations, one species reestablishment, one nomenclatural status change, three excluded species and two were kept as doubtful names. Nine new species were also detected: Stryphnodendron conicum, Stryphnodendron dryaticum, Stryphnodendronfasciatum, Stryphnodendron holosericeum, Stryphnodendron orinocense, Stryphnodendron procerum, Stryphnodendron riparium, Stryphnodendron velutinum e Stryphnodendron venosum.

京都大学 / 0048 / 新制・課程博士 / 博士(理学) / 甲第22631号 / 理博第4620号 / 新制||理||1664(附属図書館) / 京都大学大学院理学研究科生物科学専攻 / (主査)教授 田村 実, 教授 工藤 洋, 教授 永益 英敏 / 学位規則第4条第1項該当 / Doctor of Science / Kyoto University / DGAM

Aquatic plants are a heterogeneous assemblage of species that, although surviving in similar habitats, have evolved from very different genetic and ecological backgrounds. However, many aquatics share a number of anatomical, morphological, metabolic and reproductive adaptations, which have arisen independently in remarkable similarities (through convergence and parallelisms) in many unrelated groups. Despite their evolutionary uniqueness, aquatic plants are markedly underrepresented in contemporary biosystematic studies. Moreover, the taxonomic evaluation of numerous aquatic plant groups is intricate due to the strong morphological reduction and a high degree of phenotypic plasticity. This thesis focuses on two notoriously challenging aquatic plant groups, Callitriche and Ranunculus sect. Batrachium. The combination of several approaches (genome size estimation, chromosome counting, sequencing of nrDNA ITS and plastid trnT-trnL regions, examination of herbarium collections) was applied in order to improve our knowledge on principal evolutionary processes such as hybridization, polyploidization and cryptic variation and demonstrate their role on the shaping of overall aquatic plant diversity. The distribution of particular species in the Czech Republic was mapped for the first time. For both groups,...