While direct measurements of energetic demands are nearly impossible to collect on large cetaceans, comprehensive bioenergetic models can give insights on such parameters by combining physiological and ecological knowledge. This model was developed to estimate necessary food intake of gray whales, Eschrichtius robustus, of the Eastern North Pacific stock. Field Metabolic Rates (FMR) for gray whales were first estimated based on various assumptions (e.g. volumetric representation of gray whales, extent of their feeding season, and blubber depth distribution) using morphometric data, energetic costs, and food assimilation according to age and gender specific requirements. Food intake rates for gray whales of varying maturity and gender were then estimated based on FMR and caloric value of prey and compared to food intake rates of previous studies. Monte Carlo simulations and sensitivity analysis were performed to assess the model's predictions compared to observed field data from previous studies. Predicted average food intakes for adult male, pregnant/ lactating female, and immature whales were 475 ± 300, 525 ± 300 and 600 ± 300 kg d-1, respectively. Estimated blubber depths resulting from these food intakes were comparable to field data obtained from whaling data. Sensitivity analysis indicated food intake, from all parameters, as having the highest impact on the percent change in ending mass from a simulation. These food intake estimates are similar to those found in a previous study and fall within the range of food intake per body mass observed in other species of cetaceans. Though thermoregulation can be a factor in some cetaceans, it appears not to be an additional cost for gray whales as the present model's predicted lower critical temperatures for the whales (TLC) were below ambient temperatures. With temperatures increasing in the Bering Sea, the main prey of gray whales, ampeliscid amphipods, could be adversely affected, possibly resulting in increased food shortages leading to a surge in gray whale strandings.

Beluga whales are migratory over much of their range, congregating in small groups around shallow river estuaries in summer, and overwintering in large groups in areas with reliable open water. This complicates management issues because it is unclear if belugas from the common wintering ground represent one large group with exchange of individuals, or if each summer estuarine concentration should be managed as a separate stock. To examine the genetic structuring, we analyzed variation in mitochondrial DNA (mtDNA) restriction sites among 101 beluga whales from 10 regions across North America, including Greenland. Using 11 restriction enzymes, 9 haplotypes were identified among 71 whales. The remaining 30 whales were tested with only the six restriction enzymes found to identify polymorphisms. We found a marked segregation of divergent haplotypes for both sexes between eastern and western Hudson Bay. Haplotype 1 was found in 19 out of 21 animals on the east coast, while haplotype 5 was found in 18 out of 20 animals on the west coast. Sequence divergence among the 71 belugas was estimated to be 2.03%. Haplotypes fell into two major phylogenetic groups, labelled lineage I and II. Lineage I haplotypes occurred primarily in the St. Lawrence Estuary and the eastern Hudson Bay. Lineage II haplotypes occurred primarily along the western Hudson Bay, Southern Baffin Island, western Greenland, the Canadian high arctic, and the Beaufort Sea. These findings support the hypothesis that belugas exhibit maternally directed philopatry to summering grounds, and are consistent with the hypothesis that after deglaciation, the arctic was recolonized by at least two stocks of belugas divergent in their mtDNA, possibly representing Atlantic and Pacific stocks. / Thesis / Master of Science (MS)

No description available.

Passive acoustic monitoring has been used to investigate many aspects of marine mammal ecology, although methods to estimate absolute abundance and density using acoustic data have only been developed in recent years. The instrument configuration in an acoustic survey determines which abundance estimation methods can be used. Sparsely distributed arrays of instruments are useful because wide geographic areas can be covered. However, instrument spacing in sparse arrays is such that the same vocalisation will not be detected on multiple instruments, excluding the use of some abundance estimation methods. The aim of this thesis was to explore cetacean abundance and density estimation using novel sparse array datasets, applying existing methods where possible, or developing new approaches. The wealth of data collected by sparse arrays was demonstrated by analysing a 10-year dataset collected by the U.S. Navy's Sound Surveillance System in the north-east Atlantic. Spatial and temporal patterns of blue (Balaenoptera musculus) and fin whale (Balaenoptera physalus) vocal activity were investigated using generalised additive models. Distance sampling-based methods were applied to fin whale calls recorded by an array of Ocean Bottom Seismometers in the north-east Atlantic. Estimated call density was 993 calls/1000 km².hr⁻¹ (CV: 0.39). Animal density could not be estimated because the call rate was unknown. Further development of the call localisation method is required so the current density estimate may be biased. Furthermore, analysing a single day of data resulted in a high variance estimate. Finally, a new simulation-based method developed to estimate density from single hydrophones was applied to blue whale calls recorded in the northern Indian Ocean. Estimated call density was 3 calls/1000 km².hr⁻¹ (CV: 0.17). Again, density of whales could not be estimated as the vocalisation rate was unknown. Lack of biological knowledge poses the greatest limitation to abundance and density estimation using acoustic data.

Mysticetes, or baleen whales, make up one of two sub-orders of the Order Cetacea within the Class Mammalia (Drikamer et al, 2013). It contains three families, each characterized by baleen plates (Drikamer et al, 2013). These baleen plates consist of keratin that replaced the tooth structures for feeding in these mammals (Drikamer et al, 2013). Other biological characteristics of these animals include paired external nares (blowholes), a sternum consisting of a single bone, symmetrical skull and the most conspicuous of all, the body length of between ten to thirty metres (Drikamer et al, 2013). It is thought that the size characteristic was probably the main factor when categorising them as cetaceans, with the Latin word ‘Cetus’ referring to ‘sea monster’ (Carnaby, 2006). In this literature review, the focus will fall on two species of mysticetes from separate families, namely that of the southern right whale, Eubalaena australis of the Balaenidae family, and the humpback whale, Megaptera novaeangliae of the Balaenopteridae family. Emphasis in this review will fall on the aspect of site fidelity of the mother-calf pairs.

No description available.

Data collected during boat-based and aerial surveys were used to describe population structure, movements, temporal patterns of migration and skin condition of humpback whales in breeding sub-stock C1-S off southern Africa. Results confirmed that the migration route along the south coast of South Africa is linked to the winter ground off Mozambique. A lack of exchange between breeding sub-stocks C1-N and C1-S was found, suggesting that these are independent of each other. Molecular analysis revealed unexpected levels of population structure between the migration route and the winter ground of C1-S, as well as the possibility that this migration route is also utilised by some individuals from breeding sub-stock C3. A skin condition of unknown aetiology that primarily affects humpback whale mother-calf pairs was identified. The first assessment of its prevalence and severity was made, providing a baseline for future monitoring. Humpback whale abundance in an inshore region of Bazaruto Archipelago, Mozambique was estimated and attempts were also made to use the limited information off Plettenberg Bay/Knysna, South Africa. In addition to improving our understanding of humpback whales from Breeding Stock C, knowledge about another baleen whale species utilising the southwest Indian Ocean was extended. The first evidence of southern right whale presence off the coast of Mozambique since the cessation of whaling was documented. It remains unknown whether this is a remnant sub-stock or the recovering South African sub-stock reoccupying its historical range.

Behavioural responses can reveal important fitness trade-offs and ecological traps in evolutionarily novel contexts created by anthropogenic stimuli, and are of increasing conservation concern due to possible links to population-level impacts. This thesis illustrates the use of proxies for energy acquisition and expenditure within multivariate and state-based modelling approaches to quantify the relative time and energetic costs of behavioural disturbance for a deep-diving marine mammal (Physeter macrocephalus) in foraging grounds in Kaikoura Canyon (New Zealand) and near Lofoten Islands (Norway). A conceptual framework is first developed to identify and explore links between individual motivation, condition and external constraints to behavioural disturbance [Chapter 1]. The following chapters then use data from behavioural response studies (BRS) to: 1) derive biologically relevant metrics of behaviour [all chapters], 2) investigate effects of boat-based focal follows and tagging procedures [Chapters 2-3], and 3) relate responses to specific disturbance stimuli (distance, approach, noise) from whale-watching [Chapter 2], naval sonar and playback of presumed natural predator (killer whale Orcinus orca) sounds [Chapter 4]. A novel hidden state model was developed to estimate behavioural budgets of tagged sperm whales from multiple streams of biologging (DTAG) data [Chapter 3]. Sperm whales traded off time spent at foraging depths in a non-foraging and non-resting state in response to both tag boat presence, 1-2 kHz naval sonar (SPL 131-165 rms re 1μPa) and mammal-eating killer whale sound playbacks, indicating that parallel non-lethal costs were incurred in both anthropogenic disturbance and presumed antipredatory contexts. While behavioural responses were highly variable by individual, biologically informed state-based models appeared effective to control for variability in energy proxies across different functional contexts. These results and Chapter 5 “linking buzzes to prey” demonstrate that behavioural context is a signal that can aid understanding of how individual non-lethal disturbance responses can impact fitness.

In killer whales, fish- versus mammal-eating ecological differences are regarded as key ecological drivers of sociality, but the potential influence of specific target prey characteristics remains unclear. This thesis aimed to study the social patterns and dynamics of Icelandic killer whales feeding upon herring, a schooling prey that undergoes frequent changes in distribution and school size. I used a multi-disciplinary approach combining photo-identification and genetic data to understand the sociality, role of kinship and genetic differentiation within the population. Individuals sighted in summer-spawning and overwintering herring grounds during at least five separate days (N = 198) were considered associated if photographed within 20 seconds of each other. Photo-identified individuals were genotyped (N = 61) for 22 microsatellites and mitochondrial DNA control region (611 bp). The population had weak but non-random associations, fission-fusion dynamics at the individual level and seasonal patterns of preferred associations. The society was significantly structured but not hierarchically. Social clusters were highly diverse and, whilst kinship was correlated with association, it was not a prerequisite for social membership. Indeed, some cluster members had different mitochondrial haplotypes, representing separate maternal lineages. Individuals with different observed movement patterns were genetically distinct, but associated with each other. No sex-biased dispersal or inbreeding was detected. This study revealed that the Icelandic population has a multilevel society without clear hierarchical tiers or nested coherent social units, different from the well-studied salmon- (‘residents') and seal-eating populations in the Northeast Pacific. In the Icelandic population kinship drives social structure less strongly than in residents. These findings suggest effective foraging on schooling herring in seasonal grounds promotes the formation of flexible social groupings which can include non-kin. Killer whale sociality may be strongly influenced by local ecological context, such as the characteristics of the specific target prey (e.g., predictability, biomass, and density) and subsequent foraging strategies of the population.

<p>The purpose of this project, as a minor field study, is to study the fishermen and their beliefs and worship in Việt Nam. Since the worship of the fishermen exists all along the coast from the Gulf of Thailand in the south to Ha Long Bay and beyond in the north, the study is concentrated to the coastal area of central Việt Nam – that is in the Quảng Nam surroundings, mainly Hội An but also Đà Nẵng. The aspects of the representation of the worship in society as well as how it is looked upon by society, both religiously (according to tôn giáo) and politically, will also be dealt with. This study is based upon observation and non-structured interviews. Although partially being based upon Grounded theory, the study is theoretically inspired by Graham Harvey’s book Animism. Respecting the Living World. The outcome shows that the Cá Ông worship is based upon historical events and experiences. The worship, which is carried out among all fishermen, is similar throughout the coast though the dates of worship may vary. Politically the religion is not necessarily threatened, although the 1992 Constitution is somewhat ambiguous. In society, the religion is respected, but peasants do not intermingle in festival activities concerning the fishermen’s traditions. There are links to be found dealing with the arguments of Harvey when it comes to animist beliefs and features of Cá Ông.</p> / <p>Le but de cette étude de champ c’est d’étudier les pêcheurs de Viét Nam et leur confianceen et leur affection à Cá Ông. Puisque la confiance des pêcheurs dit-on existe le long du chemin du sud , de la baie de Thailand, à Ha Long Bay au nord, l’étude a été concentrée à la côte du Viét Nam central surtout la province de Quang Nam, concentrée à Hôi An et aussi Dà Nang.</p><p>Comment la croyance des pêcheurs est representée dans la société et comment la société regarde la pratique vont être traité d’un perspectif politique et religieux. L’étude s’est basée sur l’observation participante sur des interviews semistructurelles. Partiellement la théorie est liée à l’Animisme respectant le monde vivant de Graham Harvey basé sur Grounded Theory. Le résumé montre que la croyance en Cà Ông est basée sur des événementset des experiences historiques. La prière pratiquée par tous les pêcheurs est faite à la même facon même si les dates pour les festivaux et les ceremonies diffèrent. Politiquement la croyance est respectée et en partie reconnue, mais les paysans n’assistent pas aux festivals concernant des traditions des pêcheurs.</p><p>Il y a des parallèles entre l’opinion d’Harvey sur l’Animisme et les parties animistes trouvées dans la croyance en Cá Ông.</p> / <p>Nội dung của bài học tự do này là học và nguyên cứu về những người ngư dân ở việt nam về sự tín ngưỡng và tôn thờ của họ đối với Cá Ông (Whale). Theo những tin đồn của những người ngư dân đã cho thấy Cá Ông tồn tại dài theo biển của ThaiLan và xuống dưới phía nam của vịnh Hạ Long và sau đó tiếp tục lên phía bắc và ở đây bài luận này chỉ chú trọng nhất đến những ngư dân của vùng biển trung tâm Việt Nam tức là tỉnh Quảng nam, trong đó có Đà nẵng và chú trọng nhất là ở Hội An. Những gì tin, mê tín và tín ngưỡng của ngư dân là điển quan trọng được diễn biến thực tế trong xã hội đời sống ngay nay, và những gì xã hội thấy được sự thử thách sẽ diễn ra cả hai bên, như tôn giáo và cũng như trong lĩnh vực chính trị. Trong bài luận này cơ bản là gặp gở sinh viên đã trực tiếp quan sát và học hỏi những tư liệu cũng như những cuộc thẩm vấn thực tế. Lý thuyết của bài học này là một phần có liên quan đến Grahamn Harvey’s Animism. (Thuyết duy linh đối với duy vật) Respecting the Living World (Tôn trọng thế giới của sinh vật sống). Phần kết luận cho chúng ta thấy rằng sự tồn tại của Cá Ông là căn bản của những câu chuyện đã xảy ra va kinh nghiệm của ngư dân. Lễ cúng cá ông, được ngư dân thường tổ chức ở vùng biển và diễn ra hàng năm như thường lệ, nhưng ngày va mùa cúng thì có vùng lại khác nhau. Trong lĩnh vực chính trị cho thấy, sư tín ngưỡng của của ngư dân đã được tôn trọng và một phần được ông chứng trong xã hội, nhưng không phải ai cũng thờ cúng cá ông ví dụ, những người làm nghề nông thì khác chẳng hạn, họ không cúng Cá Ông như ngư dân làm biển. Ngược lại Lễ cúng cá ông đã trở thành một phong tục truyền thống của riêng ngư dân. Quan điểm của Harvey’s về thuyết duy linh đối với duy vật nó song song với nhưng linh thiêng vể, thần linh mà con người đối với Cá Ông.</p> / Uppsatsen har senare publicerats i modifierad form som boken: "Lantz, Sandra (2009). Whale Worship in Vietnam. Religionsvetenskapliga studier från Gävle 4. Uppsala: Swedish Science Press."