Thesis (doctoral)--Swedish University of Agricultural Sciences, 2006. / Thesis documentation sheet inserted. Appendix includes reproductions of six papers and manuscripts co-authored with others. Includes bibliographical references. Also issued electronically via World Wide Web in PDF format; online version lacks appendix of papers; ill. in online version displays in color.

Död ved är en viktig del i ekosystem och har många olika funktioner. Många arter är beroende av att ha tillgång till död ved, exempelvis som födokälla, bo-/växtplats eller skydd. I kalhyggen, där den naturliga produktionen av död ved inte längre finns kvar, är det därför viktigt att det lämnas kvar en viss mängd, så att området bibehåller sina biologiska förutsättningar. I undersökningen jämfördes kalhyggen med produktionsskog för att se om det fanns någon skillnad i mängd död ved. Detta gjordes genom att inventera 5 lokaler av 70–100 årig gran- och tallskog där varje lokal innehöll ett kalhygge som sammanhängde med ett område produktionsskog. Resultatet visade att det inte fanns någon skillnad i volym död ved mellan kalhyggen och produktionsskog. I inventeringen kunde man dock se att mycket av den döda veden som uppmättes i kalhyggena hade en diameter runt 10 cm medan skogsområdena hade mer varierande omkretsar. En högre volym av död ved uppmättes i granskog än i tallskog. Således kan man dra slutsatsen att slutavverkning av en produktionsskog inte verkar ha någon större påverkan på hur stor volym död ved som finns tillgängligt. De uppmätta värdena ligger dock långt under vad som finns i naturskogar, vilket indikerar att man behöver göra insatser för att öka mängden död ved i många av våra skogar. / Woody debris is an important part in many ecosystems and has several different functions. Many species depend on the existence of woody debris, where it is used for example as a food source, nesting site, growing site or as a refuge. In clearcut areas, where the natural production of woody debris no longer exists, it is therefore important that a certain amount of woody debris is left behind to make sure that the area continues to have good biological function? In this investigation, I compared clearcut areas with production forest to see if there was a difference in the volume of woody debris. Five sites with 70–100-year-old spruce- and pine forests were inventoried. Every site consisted of one clearcut area that was adjacent to an area with production forest. No difference in volume of woody debris was found between the areas. The inventory showed that many of the measured trunk diameters that were found in the clearcut areas were around 10 centimeters, whereas in the areas with production forest the diameter varied more. Larger volumes of woody debris were found in spruce forests then in pine forests. Thus,the conclusion is that clearcutting an area with production forest does not seem to have a big impact on the volume of woody debris. Nevertheless, the measured amounts of woody debris are far less then what are found in natural forests, which indicates management needs to see what improvements can be done to increase the amount of woody debris in many of our forests.

Classical approaches to evaluate the stability of large woody debris (LWD) introduced in streams for habitat restoration or flood management purposes are usually based on inappropriate assumptions and hydraulic equations. Results suggest that the physics of small cylindrical elements located in large channels cannot be transferred to the case of a large roughness elements placed in small channels. The introduction of LWD in a small channel can generate a significant modification of the upstream hydraulic conditions. This modification has direct implications on the stability of the LWD. Experiments were performed in a controlled environment: a small stream section was represented by a low roughness rectangular flume and LWD were modeled with smooth PVC cylinders. Direct force measurements were performed with a load cell and results were used to identify an equation that evaluates the drag force acting on a large cylindrical element place in a rectangular channel. This equation does not depend on a drag coefficient. Water depths were also measured during the experiments and results were used to develop an approach that evaluates the upstream hydraulic impacts of a large cylinder introduced in a rectangular channel. The effect of the variation of the unit discharge (discharge per unit of width), cylinder size, cylinder elevation from the channel bed, and downstream hydraulic conditions, could be related to the upstream hydraulic conditions with relative success. Dimensionless parameters were developed to increase the versatility of the approach. The application of this approach to field cases is expected to require adjustments, mainly because of the roughness of natural environments differs from the smoothness of the controlled environment described in this work. / Applied Science, Faculty of / Civil Engineering, Department of / Graduate

Five hundred eighty red alder along a 2.16 km portion of the Little Creek riparian zone were assessed for mortality following the 2009 Lockheed Fire near Davenport, California. The study area was divided into burn severity zones and every red alder within the riparian zone was observed and assessed for mortality. Height, diameter at breast height (DBH), and age were collected on selected trees. An estimation of red alder large woody debris (LWD) input to Little Creek since a 2010 LWD survey (Smith, 2010) was calculated using average red alder tree dimensions applied to Smalian’s formula (MLNRO, 2011). The mean proportion of dead red alder in the non-burn zone and burn-zone was .11 and .45 respectively. Volume of red alder LWD in Little Creek is estimated to have increased approximately 245% since the fire. Given other factors may have played a role in red alder mortality in Little Creek, the analysis reported here supports fire also contributed to accelerated mortality of red alder established following the catastrophic debris torrent in 1955 in Little Creek. This study provides useful information regarding fire and red alder and establishes baseline conditions of the Little Creek riparian zone following the 2009 Lockheed Fire.

Those who frequent the coastline may be accustomed to seeing driftwood washed onshore, some of it having seemingly found a home there for many years, others having been freshly deposited during the last set of storms; However, if a passerby were to take a closer look at the driftwood on the coastline, they may notice that some of these logs – also known as Large Woody Debris (LWD) – are anchored in place, a practice which is generally used for the purpose of stabilizing the shoreline or reducing wave-induced flooding. Records of existing anchored LWD project sites date back to 1997 and anecdotal evidence suggests that the technique has been used since the mid-1900’s in coastal British Columbia (BC), Canada, and Washington State, USA. Now, with an increased demand for natural and nature-based solutions, the technique is again gaining popularity. Despite this, the design of anchored LWD has largely been based on anecdotal observations and experience, as well as a continuity of design practices from the river engineering field. To date, there is no known peer-reviewed literature on the design or efficacy of LWD protection systems in a coastal environment. In 2019, the “Efficacy and Design of Coastal Protection using Large Woody Debris” research project was initiated to determine if LWD are effective at stabilizing the shoreline under wave action, if they are effective at reducing wave run-up, and if they are durable enough to meet engineering requirements for shore protection. In addition, the project aimed to determine the optimum configuration of LWD for design purposes. To meet these objectives, this study included the following work: (1) field studies of existing LWD installations, (2) experimental modeling of beach morphology with and without LWD structures, (3) experimental modeling of wave run-up with and without LWD structures, and (4) development of preliminary design guidance. The first phase of the project included field investigations at 15 existing anchored LWD sites in coastal BC and Washington State. Site characteristics, design techniques, and durability indicators were examined and correlated to a new design life parameter: ‘Effective Life’. Six primary installation techniques were observed: Single, Multiple, Benched, Stacked, Matrix, and Groyne. Observed durability and/or performance issues included: missing LWD, erosion, arson, wood decay, and anchor corrosion/damage. The Effective Life of anchored LWD was found to be strongly correlated to the tidal range and the upper beach slope for all installation types, and the LWD placement elevation relative to the beach crest elevation for single, shore-parallel structures. The many noted durability issues and ineffectiveness as mitigating erosion indicates that existing design methods for anchored LWD have not generally been effective at providing coastal protection and meeting engineering design life requirements. A comprehensive set of over 60 experimental tests were completed as part of the overall research program. Thirty-two (32) tests were analyzed as part of this study relating to the morphological response of a gravel beach with and without various LWD configurations. The tests were conducted within a wave flume at the National Research Council’s Ocean, Coastal and River Engineering Research Centre (NRC-OCRE), at a large scale (5:1) based on site characteristics and LWD design characteristics made during the previous field investigations. Tests were also conducted to assess experiment repeatability, sensitivity to test duration, sensitivity to wave height, wave period, and relative water level, influence of regular waves, and influence of log roughness. The position of the most seaward LWD (whether considering distance or elevation) was found to be strongly linked to morphological response. A theoretical relationship was developed between LWD elevation and sediment volume change. Configurations which included LWD placement below the still water level, such as the Benched configuration, were found to be most effective at stabilizing the beach profile. As part of the experimental modeling program, 24 tests were also conducted for the purpose of estimating the effect of LWD design configuration on wave run-up. In total, six different beach and LWD configurations were tested under a base set of four regular wave conditions. The study findings indicated that anchored LWD may increase wave run-up relative to a gravel beach with no structures. In particular, configurations with more logs tended to result in higher wave run-up. However, additional research is needed on the effect of LWD on wave run-up to confirm and expand these findings. There are a number of potential engineering, ecological, social, and economic benefits associated with anchored LWD installations if designed, installed, and monitored appropriately for the site conditions and user needs. To realize these potential benefits, significant additional research is needed on the topic. One of the most significant barriers to usage is a lack of information on how to effectively anchor LWD structures. However, this research project provides a baseline for future comprehensive studies on the effect and design of coastal protection using LWD. The project provides preliminary design considerations for the usage of LWD as coastal protection and contributes to the growing body of literature on nature-based solutions.

This study uses a spatial model to visualize LWD mobility areas in an approximate 1km reach of Cummins Creek, a fourth-order stream flowing through an old-growth Sitka spruce-western hemlock forest in the Oregon Coast Range. The model solves a LWD incipient motion equation for nine wood size combinations (0.1m, 0.4m, 1.7m diameters by 1.0m, 6.87m, 47.2m lengths) during the 2-year, 10-year, and 100-year discharge events. Model input variables were derived from a combination of field survey, remotely sensed, and modeled data collected or derived between June 2010 and July 2011. LWD mobility map results indicate the 2-year discharge mobilizes all modeled diameters, but mobile piece lengths are shorter than the bankfull channel boundary. Mobility areas for each wood size combination increases with discharge; 10-year and 100-year discharge events mobilize wood longer than average bankfull width within a confined section of the main stem channel, and mobilize LWD shorter than bankfull width within the main stem channel, side channels, and floodplain. No discharge event mobilizes the largest LWD size combination (1.7m / 47.2). Recruitment process was recorded for all LWD during June 2010, revealing that all mobile wood in the study reach was shorter than bankfull width. Based on these conflicting results, I hypothesize the distribution of wood in Cummins Creek can be described in terms of discharge frequency and magnitude, instead of as a binary mobile/stable classification. Mobility maps could be a useful tool for land managers using LWD as part of a stream restoration or conservation plan, but will require additional calibration.

京都大学 / 0048 / 新制・課程博士 / 博士(理学) / 甲第18107号 / 理博第3985号 / 新制||理||1575(附属図書館) / 30965 / 京都大学大学院理学研究科生物科学専攻 / (主査)教授 朝倉 彰, 講師 宮崎 勝己, 教授 疋田 努 / 学位規則第4条第1項該当 / Doctor of Science / Kyoto University / DGAM

A frequent source of pine tree mortality in recent years can be attributed to pine bark beetles and their vectored Ophiostomatoid fungi, an organism that has been observed to attract subterranean termites that preferentially recruit to this downed woody debris. This interaction may significantly modify biogeochemical fluxes in bark beetle mass mortality events, but studies are often limited to singular regions or single pine-dominated ecosystems. Two studies were designed to test the interactive effects of these associations on terrestrial biogeochemical cycles and decomposition processes, the first using replicated field trials in Mississippi, Arizona, and Honduras across two years and the second across five years in Mississippi. Both studies utilized Ophiostomatoid inoculation and invertebrate exclusion techniques to determine the individual and interactive contributions from biotic factors on biogeochemical cycling. Local drivers were found to have greater influence on biogeochemical cycling and decomposition before climate drivers overwhelmed their influence in later years.

Recent and anticipated global change has focused concern on the role of forests in ecosystem functioning and carbon sequestration. Of key importance is identifying relevant factors that drive carbon and nutrient dynamics and the consequences of changes in these processes. Systems undergoing invasion by invasive woody species are particularly prone to changes. This study examined the branch and crown biomass and decay dynamics for the invasive Chinese tallow tree in Mississippi, USA as well as the influences of stand and site conditions on biomass modeling and arthropod contributions to fine woody debris decay. Coupling biomass and decay models presents a method for modeling carbon sequestration and nutrient turnover rates at the stand level. These predictions will aid our understanding of the consequences of ecosystem change, especially those driven by invasive species.

Intercropping switchgrass (Panicum virgatum) between tree rows within young pine (Pinus spp.) plantations is a novel method to generate lignocellulosic biofuel feedstocks within intensively managed forests. Intensively managed pine supports diverse avian assemblages potentially affected by establishment and maintenance of a biomass feedstock. I sought to understand how establishing switchgrass on an operational scale affects bird communities within intercropped plantations as compared to typical intensively managed loblolly pine (Pinus taeda) plantations. I conducted breeding bird point counts, nest searching and monitoring, and coarse woody debris (CWD) surveys following establishment of intercropped switchgrass stands (6 replicates), traditionally-managed pine plantations, and switchgrass-only plots (0.1 km2 minimum) in Kemper Co., MS from 2011 to 2013. I found establishment of intercropping did not affect downed CWD, but reduced standing snags and green trees. I detected 59 breeding bird species from 11,195 detections and modeled nest survivorship for 17 species. Neotropical migrants and forest-edge associated species were less abundant in intercropped plots than controls for two years after establishment, and more abundant in year three. Short distance migrants and residents were scarce in intercropped and control plots initially, and did not differ between these treatments in any year. Species associated with pine-grass habitat structure were less abundant initially in intercropped plots, but converged with pine controls in subsequent years. Switchgrass monocultures provided minimal resources for birds. There was no evidence supporting an effect of intercropping on songbird nest survivorship. I found evidence for dominance of one species, yellow-breasted chat (Icteria virens), over another, indigo bunting (Passerina cyanea) in competition for nest sites, which illustrates how songbirds competing for nest sites can coexist in sympatry without the dominant species driving subordinate competitors to local extirpation. This dissertation, and related publications, are among the earliest research on wildlife response to intercropping. Forest managers implementing intercropping within pine plantations where vertebrate conservation is a management priority should be aware of potential changes to snag-utilizing species from reductions in green trees and snags. Songbird populations may lag behind traditional management for up to two years following establishment of switchgrass. Intercropping neither positively nor negatively affected songbird nest survival.