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O presente trabalho trata de efeitos de borda, especialmente sobre solo e serapilheira. Foi dividido em dois capítulos independentes, mas complementares: uma revisão bibliográfica e um estudo de caso em cerrado. Revisão bibliográfica. Os estudos sobre borda tiveram, inicialmente, um enfoque exclusivamente voltado aos interesses humanos, ganhando o enfoque ecológico atual há menos de três décadas. Assim, o principal objetivo desta revisão bibliográfica foi gerar um panorama sobre como o assunto \"borda\" vem sendo estudado pelo mundo, de 1990 a 2007, por meio de buscas em periódicos disponíveis online. De forma geral, o número de publicações sobre o tema \"borda\" vem aumentando visivelmente com o passar dos anos. No entanto, a região inter-tropical (latitudes entre 23°26\'22\'\'N e 23°26\'22\'\'S) é a que demanda maior preocupação na atualidade por apresentar menor número de publicações. Estudo de caso. A crescente substituição de áreas de cerrado por cultivos de cana-de-açúcar e eucaliptos tem gerado intensa fragmentação de habitats nativos no estado de São Paulo, deixando-os sujeitos aos efeitos de borda. Neste capítulo, o principal objetivo foi verificar os efeitos de borda causados por monoculturas de cana-de-açúcar e eucaliptos sobre os nutrientes do solo e da serrapilheira em três fragmentos de cerradão, na região nordeste do estado de São Paulo (municípios de Santa Rita do Passa Quatro e Luiz Antônio), visando a conservação das áreas estudadas. Análises físicas e químicas de amostras de solo superficial (0-15 cm) e de serrapilheira, coletadas a distâncias de 0, 10, 20, 40, 80 e 160 m da borda entre os fragmentos nativos e a matriz (monocultura), bem como no interior do fragmento e nos dois tipos de matriz demonstraram a existência de um gradiente na quantidade dos nutrientes, evidenciando efeitos de borda causados por cultivos agrícolas. / This thesis tells about border effects, especially on soil and litter. It contains two independent, but complementary, chapters: a literature review and a case study in cerrado. Literature review. The first studies on edge were exclusively focused on the man\'s point of view, and the focus turned to ecological concern less than three decades ago. Thus, the main objective of this literature review was to get a broad view on how the subject \"border\" is being studied around the world, from 1990 to 2007, by searching in scientific journals available online. In general, the number of publications on the theme is increasing outstandingly with time. But the inter-tropical region (latitudes between 23°26\'22\'\'N and 23°26\'22\'\'S) is the one with fewer publications and requires awareness. Case study. The increasing substitution of cerrado environments by sugar cane and eucalyptus plantations in São Paulo State has caused severe fragmentation of the native habitats, which become exposed to edge effects. The main objective of this chapter was to verify edge effects caused by sugar cane and eucalyptus monocultures on the nutrients of cerrado soil and litter. The research was carried out in three cerradão fragments in the Northeast of São Paulo State (Santa Rita do Passa Quatro and Luiz Antônio municipalities), aiming at the conservation of the studied areas. Chemical and physical analyses of surface soil (0-15 cm) and litter, collected at 0, 10, 20, 40, 80 and 160 m of the edge between the native fragment and the matrix (monoculture), as well as inside the fragment and the monocultures, showed a gradient in the amount of nutrients, supporting the existence of border effects caused by the agriculture.

In the wetter climates associated with the coastal forests of northwestern North America, coarse woody debris (CWD) accumulations in the form of snags, downed boles, and large branches can be large in natural forest ecosystems. Seedlings often regenerate on stumps and downed logs in the understory of old-growth coastal forests. The question remains though, whether CWD is a necessary component for seedling survival and growth in forests managed for commodity production. This study addresses one concern of forest managers: is there an immediate nutritional or moisture supply advantage conferred by CWD for the survival and growth of seedlings in the coastal climate of British Columbia? We compared survival and growth of seedlings planted in decaying wood compared to non-woody humus forms and mineral soil under heavy shade and full light conditions. Low light environments are of particular interest since reports of the strong association between CWD and regeneration has primarily referred to understory seedlings and saplings in old-growth forests.

The forest floor represents the uppermost organic and organicenriched mineral soil horizons. They have been formed by the deposition of organic material and the subsequent biologically mediated decomposition. The forest floor influences rooting-zone temperature, aeration, moisture, and nutrient conditions, and hence, forest productivity. Considering the importance of the forest floor, and the fact that it is exposed to disturbance (being the surface layer), we need to assess the potential impacts our logging practices may have. Clearcutting, one of the contentious silvicultural practices used in British Columbia, is imputed to most adversely affect ecosystems and sustainability. We assessed the long-term impact of clearcutting on the forest floor by documenting changes in the thickness, chemical and biotic properties of the humus form across a chronosequence of forest stands. The study was located in the largest and most representative portion of the coastal rainforest the Very Wet Maritime Coastal Western Hemlock (CWHvm) subzone.

In the wetter climates associated with the coastal forests of the Pacific Northwest, coarse woody debris (CWD) accumulations in the form of snags, downed boles, and large branches can be large in natural forest ecosystems. Although maintaining organic matter for sustainable site productivity is not in dispute, the importance of CWD as a source of soil organic matter is questionable. Forest managers attempting to optimize timber production need to know how CWD affects short-term forest tree growth and productivity. This study addresses the question of the immediate value of CWD for growth of mature (90 year old) western hemlock (Hw). Because of practical difficulty with mature trees growing in different substrates, we utilized fine root distribution or proliferation, as an indicator of important substrates.

Knowledge of ecological characteristics of trees, sites and tree growth on different sites is fundamental for silvicultural decision-making and planning. With the biogeoclimatic ecosystem classification in place, silvicultural management in British Columbia has been given an ecological foundation; however, relationships between growth and site have not yet been fully investigated. The purpose of this study was to determine how height growth of Douglas-fir within the drier portion of the CWH zone varies with site. We adopted site index (m @ 50 yr bh) as a species-specific measure of forest productivity, recognizing that it indicates height growth performance at a selected point in time. If forest productivity is correlated with ecological measures of site quality, what site factors should be used to quantify the relationships? Because of compensating effects, the numerous site factors can be reduced to four primary (synoptic) factors that directly affect plant establishment and growth: climate (light and temperature), soil moisture, soil nutrients, and soil aeration (not used in this study).

The three major components in the site classification of the biogeoclimatic ecosystem classification system are: climatic regimes, soil moisture regimes (SMRs) and soil nutrient regimes (SNRs). Both SMRs and SNRs can be identified in the field using soil characteristics and indicator plants. In the case of SMRs a quantitative classification was also developed that allow comparison of SMRs in different subzones. However, similar quantitative classification has not yet been developed for SNRs. This pamphlet summarizes and compares the results of several regional studies conducted in different biogeclimatic zones. Each of theses studies aimes to develop a quantitative SNR classification (Table 1). The comparison will examine: (1) how well the field-based classification matches quantitative classification, and (2) which direct measures distinguish best between field-identified SNRs.

O presente trabalho trata de efeitos de borda, especialmente sobre solo e serapilheira. Foi dividido em dois capítulos independentes, mas complementares: uma revisão bibliográfica e um estudo de caso em cerrado. Revisão bibliográfica. Os estudos sobre borda tiveram, inicialmente, um enfoque exclusivamente voltado aos interesses humanos, ganhando o enfoque ecológico atual há menos de três décadas. Assim, o principal objetivo desta revisão bibliográfica foi gerar um panorama sobre como o assunto \"borda\" vem sendo estudado pelo mundo, de 1990 a 2007, por meio de buscas em periódicos disponíveis online. De forma geral, o número de publicações sobre o tema \"borda\" vem aumentando visivelmente com o passar dos anos. No entanto, a região inter-tropical (latitudes entre 23°26\'22\'\'N e 23°26\'22\'\'S) é a que demanda maior preocupação na atualidade por apresentar menor número de publicações. Estudo de caso. A crescente substituição de áreas de cerrado por cultivos de cana-de-açúcar e eucaliptos tem gerado intensa fragmentação de habitats nativos no estado de São Paulo, deixando-os sujeitos aos efeitos de borda. Neste capítulo, o principal objetivo foi verificar os efeitos de borda causados por monoculturas de cana-de-açúcar e eucaliptos sobre os nutrientes do solo e da serrapilheira em três fragmentos de cerradão, na região nordeste do estado de São Paulo (municípios de Santa Rita do Passa Quatro e Luiz Antônio), visando a conservação das áreas estudadas. Análises físicas e químicas de amostras de solo superficial (0-15 cm) e de serrapilheira, coletadas a distâncias de 0, 10, 20, 40, 80 e 160 m da borda entre os fragmentos nativos e a matriz (monocultura), bem como no interior do fragmento e nos dois tipos de matriz demonstraram a existência de um gradiente na quantidade dos nutrientes, evidenciando efeitos de borda causados por cultivos agrícolas. / This thesis tells about border effects, especially on soil and litter. It contains two independent, but complementary, chapters: a literature review and a case study in cerrado. Literature review. The first studies on edge were exclusively focused on the man\'s point of view, and the focus turned to ecological concern less than three decades ago. Thus, the main objective of this literature review was to get a broad view on how the subject \"border\" is being studied around the world, from 1990 to 2007, by searching in scientific journals available online. In general, the number of publications on the theme is increasing outstandingly with time. But the inter-tropical region (latitudes between 23°26\'22\'\'N and 23°26\'22\'\'S) is the one with fewer publications and requires awareness. Case study. The increasing substitution of cerrado environments by sugar cane and eucalyptus plantations in São Paulo State has caused severe fragmentation of the native habitats, which become exposed to edge effects. The main objective of this chapter was to verify edge effects caused by sugar cane and eucalyptus monocultures on the nutrients of cerrado soil and litter. The research was carried out in three cerradão fragments in the Northeast of São Paulo State (Santa Rita do Passa Quatro and Luiz Antônio municipalities), aiming at the conservation of the studied areas. Chemical and physical analyses of surface soil (0-15 cm) and litter, collected at 0, 10, 20, 40, 80 and 160 m of the edge between the native fragment and the matrix (monoculture), as well as inside the fragment and the monocultures, showed a gradient in the amount of nutrients, supporting the existence of border effects caused by the agriculture.

The Tropical savannas of central Brazil (cerrado) and the Everglades wetland (Florida) ecosystems are ideal systems to study landscape spatial mosaics and their interactions. Both ecosystems show a variety of plant physiognomies distributed within small spatial scales and elevation gradients. Such variety of plant physiognomies provide an opportunity to investigate the roles of climate, topography, nutrient availability and water table dynamics as determinants of plant physiognomic distributions, and their role in shaping regional systems. South Florida Wetlands and the tropical savannas of central Brazil are examples of hydrologically-controlled ecosystems. In hydrologically-controlled ecosystems water sources, the availability of nutrients, and the patterns of water movement play important roles in determining vegetation structure and function. The main objective of this study was to understand ecosystem level processes that shape different physiognomies in two hydrologically-controlled ecosystems. I conducted field work at the IBGE ecological reserve, a field experimental station located in Brasilia, Brazil. I also worked at the Everglades National Park in an area located near the south entrance of the Park in Homestead, Florida. I carried out three interconnected studies investigating water and nutrient dynamics: (1) In a Brazilian savanna I manipulated levels of litter input and measured changes to soil properties, organic matter decomposition and tree growth. I found that changes in litter input affect soil physicochemical properties and soil biochemical processes. I also found that litter dynamics influence tree growth through their effects on soil physicochemical properties. (2) I also studied the effect of water table depth and its temporal variation on spatial patterns of vegetation distribution in the cerrado landscape. I monitored diurnal and seasonal changes in water table depth along two tree-density and topographic gradients. In addition, I measured woody species composition, growth rates of four tree species, litter production, soil nutrients, and nutrient resorption efficiency along those two gradients. I found that water table depth has an important role in determining the spatial distribution of cerrado physiognomies; it also affects tree growth, species composition and nutrient resorption efficiency. (3) In the Everglades I studied patterns of underground water uptake by two vegetation types. I monitored seasonal and diurnal changes in water table depth in a Hammock forest, in a stand dominated by the invasive woody species Schinus terebinthifolius, as well as the water level in an adjacent lake. I estimated stand level transpiration using two different approaches: with sap flow measurements and diurnal oscillations in water table levels. Then, I calculated the total quantity of groundwater withdrawn by evapotranspiration for the wet and dry seasons in the Hammocks and in the exotic invaded site and then compared the results. I found that water uptake by Everglades trees is well coupled to diurnal changes in water table depth and that the amount of water withdrawn from the groundwater was larger during the wet season than during the dry season. Finally, I detected hydrological feedbacks between different vegetation types and nearby bodies of water. Results of this study contributes to the current knowledge of ecosystem level processes in tropical and subtropical ecosystems where water circulation and water availability play a dominant role in shaping vegetation structure and function.

Summary In the Sudanian zone of West Africa like in the Borgou region of Benin, agricultural and livestock systems face more and more difficulties linked to climatic changes, the decrease in soil fertility and the degradation of natural grasslands. This leads to a decrease in agricultural yields and feeding resources for livestock. On another hand, the socio-economical conditions of the small-holders dont allow them to invest in mineral fertilisers to remedy to soil fertility decrease. The introduction of ley pastures in crop rotations may be an affordable solution to increase available feeds for livestock and to improve soil fertility. The first part of this work reviews the potentialities of various local and exotic grasses and legumes which could be cultivated in the Borgou region. The choice of these species take into consideration the pedo-climatic conditions of the region, their aptitude for production, their nutritive value and their contribution to soil fertility. In the second and third parts of the thesis, the results of experimental 3-year ley pastures using 4 grasses (Panicum maximum Jacq cv. C1, Andropogon gayanus Kunth with broad leaves, A. gayanus with narrow limbs and Pennisetum purpureum Schumach.) and 6 légumineuses (Aeschynomene histrix Poir, Stylosanthes fruticosa (Retz.) Alson, Centrosema pubescens Benth., Mucuna pruriens (L.) DC. var. utilis Wall. Ex Wight) Baker ex Burck, Cajanus cajan (L.) Millsp. and Leucaena leucocephala (Lam.) De Wit) and exploited without any supply in mineral fertilisers are described concerning their yields and nutritive values. The grasses productions varied between 2 200 and 4 600 forage units (UF, in French unites fourragères)/ha/year and between 170 and 450 kg digestible crude protein/ha/year according to species and years (P<0.05). The forage legumes produced between 1 400 and 4 800 kg dry matter (DM)/ha/year according to species and years (P<0.05) with crude protein contents ranging from 16 to 27 %. Except Na and Zn, the combination of these two forage groups can satisfy the ruminant needs in macro and micro-minerals. The fourth part of the manuscript presents a soil-fertility balance under the different ley pastures after 3 years of exploitation. At the end of this study, it appears that in the experimental conditions (cut-and-carry system without fertilisers) Andropogon gayanus with broad leaves and the ligneous legume species Cajanus cajan and Leucaena leucocephala ensure the highest DM yields and are the less exhausting for soil fertility due to deeper root systems enabling to bring up to the surface mineral elements from deep layers / Résumé Dans la région soudanienne de lAfrique de lOuest comme dans le Borgou au Bénin, les systèmes de production agricole et délevage connaissent de plus en plus de difficultés liées aux aléas climatiques, à lappauvrissement rapide des sols et à la dégradation des végétations naturelles. Il en résulte une baisse des rendements agricoles et des ressources alimentaires pour le bétail. Par ailleurs, les conditions socio-économiques des producteurs limitent leurs possibilités dachat d'engrais en vue de remédier à la baisse de fertilité des sols. Dans ce contexte lintroduction de jachères fourragères dans les rotations culturales constitue une solution pour augmenter les disponibilités alimentaires destinées au bétail et enrichir le sol. La première partie de ce travail présente un inventaire de graminées et de légumineuses locales ou exotiques qui peuvent être implantées dans le Borgou. Le choix de ces espèces tient compte des conditions pédo-climatiques de la région, de leur aptitude à la production, de leur valeur alimentaire et de leur contribution à la fertilité des sols. Dans les deuxième et troisième parties de la thèse, les résultats de jachères expérimentales de trois ans comprenant 4 graminées (Panicum maximum Jacq var. C1, Andropogon gayanus Kunth à larges feuilles, A. gayanus à fines feuilles et Pennisetum purpureum Schumach.) et 6 légumineuses (Aeschynomene histrix Poir, Stylosanthes fruticosa (Retz.) Alson, Centrosema pubescens Benth., Mucuna pruriens (L.) DC. var. utilis Wall. Ex Wight) Baker ex Burck, Cajanus cajan (L.) Millsp. et Leucaena leucocephala (Lam.) De Wit) et exploitées sans apports de fumure minérale sont décrits en ce qui concerne leurs rendements et leurs valeurs alimentaires. Les productions des graminées ont varié entre 2 200 et 4 600 unités fourragères/ha/an et 170 à 450 kg de protéines brutes digestibles/ha/an selon les espèces et les années (P<0,05). Les légumineuses ont produit entre 1 400 et 4 800 kg de matière sèche/ha/an selon les espèces et les années (P<0,05), avec des teneurs en protéines variant entre 16 et 27 %. A lexception du Na et du Zn, lutilisation combinée des fourrages de ces espèces peut satisfaire les besoins en minéraux et oligo-éléments des ruminants. La quatrième partie du manuscrit présente un bilan de fertilité des sols sous ces différentes cultures après 3 années de jachère. Au terme de cette étude, il apparaît que dans les conditions expérimentales (affouragement en vert sans fumure minérale) la graminée Andropogon gayanus à larges feuilles et les légumineuses arbustives Cajanus cajan et Leucaena leucocephala sont les plus productives et les moins épuisantes pour le sol, eu égard à leur enracinement profond, capable de ramener des éléments minéraux en surface.