Dans la plupart des pays du Tiers-monde, et plus encore dans les zones montagneuses,<br />l'Homme puise directement dans son environnement naturel les ressources nécessaires à sa survie.<br />Ainsi, bien que rarement autarciques, les villages de la Haute montagne marocaine n'en sont pas<br />moins tributaires de ce que leur offre le milieu, et plus précisément la forêt, que ce soit pour<br />alimenter ou soigner le bétail, pour bâtir les maisons et les bergeries, ou encore pour se chauffer et<br />faire la cuisine. Mais cette très forte dépendance vis à vis de l'arbre, à laquelle s'ajoute une forte<br />croissance démographique, a un prix, comme en témoigne l'intense dégradation des écosystèmes<br />forestiers.<br />A travers l'étude du peuplement à genévriers thurifères de la vallée de l'Azzaden (Haut Atlas,<br />Maroc), nous nous sommes donc attachés (1) à préciser les potentialités et les ressources de cet<br />écosystème méditerranéen de haute montagne: réserve ligneuse et productivité (développement<br />d'une méthodologie originale non destructive d'estimation de la phytomasse), cycle du carbone et<br />des éléments minéraux; (2) à déterminer le rôle du facteur anthropique dans les processus de<br />dégradation des sols et de la végétation (modélisation de l'évolution régressive du peuplement), et<br />dans les difficultés de régénération naturelle du Genévrier thurifère.<br />Au-delà de la portée locale d'une recherche ciblée sur une espèce menacée de disparition à court<br />terme, et des implications écologiques, économiques et sociales de la déforestation d'une petite<br />vallée du Haut Atlas, cette étude relève d'une problématique plus générale. La vallée de l'Azzaden<br />peut, en effet, être considérée comme un modèle de fonctionnement d'un écosystème méditerranéen<br />perturbé par l'action anthropique, les données obtenues renseignant plus généralement sur les<br />problèmes globaux de la steppisation, du surpâturage, de l'épuisement des ressources énergétiques,<br />de l'érosion des sols et des variations du stock carboné des milieux semi-arides.

Diss. (sammanfattning) Uppsala : Sveriges lantbruksuniv., 2007. / Härtill 4 uppsatser.

Over the past 100–150 years Texas rangelands have dramatically changed from native open savannahs to dense woodlands. On the Edwards plateau, a major management concern is the increasing encroachment of Ashe juniper (Juniperus ashei). Preceding an anticipated brush management program, I investigated the presence, co-occurrence, and habitat relationships of 7 breeding bird species in the Leon River Watershed in central Texas, USA: black-capped vireo (Vireo atricapillus), golden-cheeked warbler (Dendroica chrysoparia), northern bobwhite (Colinus virginianus), white-eyed vireo (Vireo griseus), Bell’s vireo (Vireo bellii), painted bunting (Passerina ciris), and brown-headed cowbird (Molothrus ater). Vegetation characteristics were compared between sites occupied by each species and unoccupied sites using univariate analysis. Models for predicting species site occupancy were developed (using logistic regression) based on habitat characteristics correlated with the presence of each species. Two species of special concern, the endangered black-capped vireo and golden-cheeked warbler occupied 5.6% of sites and 13.8% of sites respectively, while the brood parasite brown-headed cowbird was the most widespread, occupying 86.8% of sites. Species co-occurrence patterns revealed significant associations between the golden-cheeked warbler and each of 5 other species. For most species, variables included in habitat models could be explained by knowledge of species known habitat associations. For example, the black-capped vireo was positively associated with increasing low-growing (<1.5 m) hardwood cover and with Low Stony Hill ecological sites. The golden-cheeked warbler was positively associated with increasing density of larger juniper trees, increasing variability in vertical vegetation structure, and decreasing midstory canopy of deciduous nonoaks (e.g., cedar elm [Ulmus crasifolia]). It also preferred Low Stony Hill and Steep Adobe ecological sites. Site occupancy seemed to be driven by variables that describe overall vegetation structure. In particular, cover of low-growing non-juniper vegetation and juniper tree density appeared to be important in determining site occupancy for several species. Although the models constructed were not very robust, resource managers can still benefit from such models because they provide a preliminary examination of important controlling variables. Managing rangelands to maintain or restore a mosaic of juniper patches and open shrublands are likely to help meet the habitat requirements of these bird communities.

Στην παρούσα διδακτορική διατριβή εκθέτονται η χλωρίδα των Βαρδουσίων ορέων καθώς και τα δεδομένα που προκύπτουν από την διερεύνηση – ανάλυσή της. Στο πεδίο της βλάστησης (φυτοκοινωνιολογία), έχει επιβεβαιωθεί σχεδόν το σύνολο των προηγούμενων αναφορών και έχει συμπληρωθεί με νέα δεδομένα όσο αφορά τις φυτοκοινωνιολογικές ομαδοποιήσεις. Συγχρόνως δικαιολογήθηκε και εξηγήθηκε η ύπαρξη των φυτοκοινωνιολογικών μονάδων σε σχέση με το σύνολο των οικολογικών παραγόντων που ήταν δυνατό να ληφθούν υπόψη (μικροκλίμα, εδαφικοί παράγοντες, γεωλογικό - πετρολογικό υπόστρωμα), συμβάλλοντας έτσι στην συνοικολογική προσέγγιση του θέματος. Επίσης όπου ήταν δυνατό έγινε προσπάθεια να διερευνηθεί η δυναμική εξέλιξης των περισσότερων φυτοκοινωνιολογικών ομάδων, όπως αυτή διαμορφώνεται κάτω από την επίδραση των περιβαλλοντικών μεταβλητών. Πιο κάτω ακολουθεί μια συνοπτική περιγραφή των σημαντικότερων σημείων της διδακτορικής διατριβής, καθώς και τα συμπεράσματα που μπορούν να διατυπωθούν από τα αποτελέσματα της ερευνητικής αυτής προσπάθειας. / In the present doctoral thesis the flora of Mt. Vardousia, as well as the data that have resulted from its analysis are presented. From a vegetation (plant sociology) point of view, almost all the previous reports on the mountain have been confirmed and have been enriched with new data as far as phytosociological groups are concerned. Additionally, the existence of certain phytosociological units was justified and explained based on the ecological factors that were possible to be measured (microclimate, soil, geology, geological substrate), contributing in this way to the synecological approach of the subject. Also, where feasible, an effort was done to study the dynamics of succession of most of the phytosociological groups, as they depend on various environmental variables. A concise description of the most important points of the doctoral thesis, as well as the conclusions that can be extracted from the results of this research follow.

Juniper trees (Juniperus osteosperma, J. monosperma, J. deppeana and J. scopulorum) grow under widely varying climatic and edaphic conditions throughout the American southwest. This study is chiefly concerned with a test of the climatic response in the partitioning of the stable isotopes of carbon in such trees. The relationships developed here, for example, might be used to extract paleoclimatic information from ancient juniper samples preserved in cave middens. In order to test for a climatic response in the leaf cellulose δ¹³C values, leaves from a total of 29 trees were sampled in the immediate vicinity of 9 meteorological stations across the state of Arizona. Care was taken to insure that 22 of the trees experienced only the temperature and precipitation values reflected by their site meteorological stations. As a cross-check, 7 trees exposed to temperature and/or precipitation levels clearly deviant from their site averages were also sampled. In general, each tree was sampled at four places, approximately 2 m above the ground. All leaf samples were reduced to cellulose (holocellulose) before combustion and analysis for their δ¹³C value. The δ¹³C value for each site was derived from an average of 2 to 4 trees per site, the value of each tree being the average of its individual samples. The one sigma 13C variation found between trees at any given site is ±0.38‰; within a single tree, ±0.36‰; and for repeat combustions, ±0.20‰. The δ¹³C values of the juniper sites were regressed against the temperature and precipitation of the individual months and running averages of months across the year using polynomial, multiple regression analysis. Temperature and precipitation were entered as separate variables in a general multiple regression model and also as a combined, single variable (T /P) in a more specific approach. The pattern formed by the multiple correlation coefficients, when plotted by months across the year, closely follows the seasonal variations in photosynthetic activity. Cellulose δ¹³C values have minimum correlation with temperature and precipitation (considered jointly) during summer months and maximum correlation during spring months. For an individual month, the temperature and precipitation (jointly) of April correlated at the highest level with a multiple adj. R = 0.994 and an F = 166; for a maximum seasonal response, March-May reached a multiple adj. R = 0.985, F = 66. The results using the combined, single variable (T /P) were nearly equivalent for the same months: April's adj. R = 0.957, F = 45; March-May's adj. R = 0.985 with an F = 132. The ability of T and P as independent predictors is considerably less than their ability in combination; e.g., 13C g(T) for March-May has an adj. R = 0.80 and 6 13C = h(P) has an adj. R = -0.67 compared to their in- concert adj. R value of 0.985. The results of this study, therefore, strongly support a high degree of climatic sensitivity in the partitioning of the stable isotopes of carbon in juniper leaf cellulose: the correlation coefficients and their F statistics are sufficiently high to consider temperature and precipitation (acting jointly) as accurate predictors of cellulose δ¹³C values in the system studied.

Plant macrofossils from packrat middens in two southeastern Utah caves outline development of modern plant zonation from the late Wisconsin. Allen Canyon Cave (2195 m) and Fishmouth Cave (1585 m) are located along a continuous gradient of outcropping Navajo Sandstone that extends from the Abajo Mountains south to the San Juan River. By holding the site constant, changes in the floral composition for a plot of less than one hectare can be observed, even if sporadically, over tens of millennia. At Allen Canyon Cave, engelmann spruce-alpine fir forest was replaced by the present vegetation consisting of pinyon-juniper woodland on exposed ridgetops and cliffside stands of Douglas fir, ponderosa pine, and aspen. Xerophytic woodland plants such as pinyon, Plains prickly pear, and narrowleaf yucca arrived sometime in the middle Holocene between 7200 and 3400 B.P. At Fishmouth Cave, Utah juniper in Holocene middens replaced blue spruce, limber pine, Douglas fir, and dwarf and Rocky Mountain junipers in late Wisconsin samples. Disharmonious associations for the late Wisconsin occur only at the lower site with the xerophytes Mormon tea, Plains prickly pear, and narrowleaf yucca growing alongside subalpine conifers. One possible explanation involves the late Wisconsin absence of ponderosa and pinyon pines from the Colorado Plateaus. Released from competition at their lower limits, subalpine conifers were able to expand into lower elevations and mix with xerophytic plants found today in understories of pinyon-juniper and ponderosa pine woodlands. Quantitative climatic estimates are derived for the late Wisconsin by applying vertical lapse rates for temperature and precipitation to the amount of vegetation depression. The Fishmouth Cave sequence indicates a minimum lowering of 850 m for blue spruce, limber pine, and dwarf juniper. A depression of at least 700 m for engelmann spruce and alpine fir is suggested for the Allen Canyon locality. Use of conservatively low lapse rates for stations below 2080 m yields a 3-4°C cooling from present mean annual temperature and 35 to 60 percent more rainfall than today. Steeper lapse rates associated with more mountainous terrain suggest a 5°C lowering in temperature and up to 120 percent increase over modern precipitation.